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Numerous footprints areas has been found, since 1962, in Permian, Triassic and Hettango-Sinemurian. All show precised oriented trackways. So that they were undestood like walking ways but their precise meaning is still enigmatic.
Footprints are numerous in France where they found from Carboniferous to Quaternary levels. Those of Middle Triassic (Anisian-Ladinian) are especialy interesting because they are tridactyl footprints made probably by first dinosaurs. First skelettons are known from Upper Ladinian / lower Carnian.
Comparaison des ichnopopulations des Grands Causses (Sud de la France) et du Veillon (Vendée)
(2003)
From descriptive statistic method concerning length (L), width (W) and L/W, it is suggested that Grallator variabilis samples from Vendée (Le Veillon) and Grands Causses belong to the same ichnospecies. Also, by this way, G. maximus of Hettangian of Vendée could be synonym of G. minusculus from USA Hettangian.
