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Dr. Nonfin (1931) in his book on the "Biology of the Amphibia", while discussing the inter-relationships of Pelobatidae, divides the family into Megophrynae, Pelobatinae and Sooglossinao und points out that among these three "the most primitive genus in the sub-family is the wide spread Megopluys or Megalophrys (including Leptobrachium)". ...