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Widely distributed phoretic blister beetles usually display unstructured geographic patterns of genetic diversity within species, resulting from recurrent long-dispersal events across their range. To investigate the extent of this pattern in the phoretic genus Lampromeloe, and particularly in western Mediterranean and European populations of L. variegatus, we performed: (1) a phylogeographic analysis based on fragments of the mitochondrial genes COI and 16S, and (2) a morphological revision based on qualitative and morphometric traits. Two allopatric lineages were recovered within L. variegatus, one distributed across Europe and the other in North Africa. These lineages are readily distinguishable based on morphological traits. We conclude that these lineages constitute two diagnosable evolutionary units and, consequently, we describe the North African populations as a new species, Lampromeloe pantherinus sp. nov. The new species is closely related to L. variegatus, from which it differs mainly by the elytral macrosculpture and by the shape of male genitalia. The elytral macrosculpture of L. pantherinus is somewhat similar to that of L. cavensis, a species widely distributed in the lowlands of the Mediterranean Basin. However, these two non-sister species can be distinguished by the morphology and macrosculpture of the pronotum and by the shape of the male genitalia.
The “trachystreptoform” species of Spirostreptidae, i.e., species which would formerly have been ascribed to the tribe Trachystreptini, from the Udzungwa Mountains are (re)described, including one new genus and five new species: Attemsostreptus reflexus Akkari & Enghoff, 2019, A. cataractae Enghoff sp. nov., A. leptoptilos Enghoff sp. nov., A. julostriatus Enghoff sp. nov., Lophostreptus tersus (Cook, 1896) (= L. ptilostreptoides Carl, 1909 syn. nov.), L. magombera Enghoff sp. nov., and Udzungwastreptus marianae Enghoff gen. et sp. nov. The type material of Lophostreptus regularis Attems, 1909 (= L. tersus) is discussed. The discussion includes paragraphs on the classification and the Udzungwa fauna of Spirostreptidae, on grouping of the Udzungwa trachystreptoform species in relation to altitude, and on the possibly recent immigration of A. reflexus and L. tersus into the Udzungwa Mts.
Six species of the spider genus Spinirta Jin & Zhang, 2020 from southern China (Araneae: Corinnidae)
(2024)
In the current study, six species of the genus Spinirta from southern China are recognized, four new species are described: S. hongyui Wang, Lu & Z.S. Zhang sp. nov. (♂♀), S. liuae Wang, Lu & Z.S. Zhang sp. nov. (♂♀), S. simianshan Wang, Lu & Z.S. Zhang sp. nov. (♂♀) and S. yintiaoling Wang, Lu & Z.S. Zhang sp. nov. (♂). Females of S. aviforma Jin & Zhang, 2020 (♂♀) and S. quadrata Jin & Zhang, 2020 (♂♀) are described for the first time. Photos of the body and copulatory organs, as well as the locality map are provided.
Two new species and one new genus of Lamiinae (Coleoptera: Cerambycidae) are described in Acanthocinini: Embera flava new species, new genus, from Panama; and Nealcidion lingafelteri new species from Costa Rica. The species-group name of Nealcidion napoensis Nascimento and McClarin, 2018 is corrected and morphological variations in Leptostylus cristulatus Bates, 1872 are reported. Trypanidius mimicavus Carelli, Monné, and Souza, 2013 is redescribed and transferred to Carphina Bates, 1872, forming the new combination Carphina mimicavus. Colombicallia curta Galileo and Martins, 1992 (Calliini) is recorded from Panama for the first time.
ZooBank registration. urn:lsid:zoobank.org:pub:7DD2EF0F-7DC9-4A85-83DA-10F4EE8CA8CB
Identifications of recently collected Eucnemidae (Coleoptera: Elateroidea) borrowed from two collections have resulted in the discovery of two new species: Entomophthalmus abbreviatus Otto (Cuba) and Trigonopleurus cordobaalfaroi Otto (Guatemala and Louisiana, USA). Images of the two newly described species along with two New World Entomophthalmus Bonvouloir species for comparative purposes are included.
ZooBank registration. urn:lsid:zoobank.org:pub:E83B2AC6-33ED-4680-8F2E-4E455A26D4C8
Several adult female monstrilloid copepods, collected in March 2022 from the protected reef area of Xcalak, on the southern part of the Mexican Caribbean coast, proved to belong to two undescribed species of Monstrilla Dana, 1849. They are described here as M. xcalakensis sp. nov. and M. annulata sp. nov., partly by use of scanning electron microscopy. Females of the two species are generally similar but differ in: (1) antennular segmental structure, (2) antennular armature and ornamentation, (3) structure and setation of the fifth leg, (4) number and modifications of the caudal setae, and (5) integumental ornamentation. Comparison with congeneric species revealed distinctive features for both species that support their status as new. These two species are interesting additions to the reef-dwelling monstrilloid copepod fauna of the Mexican Caribbean and confirm Monstrilla as the most diverse genus of monstrilloids in this area, now represented by nine species.
A taxonomic revision of the oil-collecting bees of the subgenus Epicharis (Epicharitides) Moure, 1945 is provided. A total of nine species were recognized: E. cockerelli Friese, 1900; E. duckei Friese, 1901; E. iheringi Friese, 1899; E. luteocincta Moure & Seabra, 1959; E. minima (Friese, 1904); E. obscura Friese, 1899, and E. rufescens Moure & Seabra, 1959, along with E. mesoamericana sp. nov. and E. lia sp. nov., two new species from the Central American and Amazonian provinces, respectively. Redescriptions, diagnoses, and figures of specimens of both sexes, floral records, distribution maps, an identification key, and an updated catalogue of all species of the group are also provided. In addition, the lectotype of E. duckei was also designated to stabilize the application of the name.
The genus Neocranaus Roewer, 1913 is revisited, its composition is expanded from two to five species and a new generic diagnosis is presented. Neocranaus albiconspersus Roewer, 1913, type species of the genus, is redescribed. The genus Tolimaius Roewer, 1915 syn. nov. is considered as a junior subjective synonym of Neocranaus, its sole member being transferred to Neocranaus – N. pectinitibialis (Roewer, 1915) comb. nov. – and redescribed here. The new combination Neocranaus laevifrons (Roewer, 1917) comb. nov. is proposed for Holocranaus laevifrons Roewer, 1917. The new species Neocranaus gladius Villarreal & Kury sp. nov. is described, from P.N.N. Yariguíes, Santander Department, Colombia. For the first time, the genital structure of this genus is illustrated. A key to the identification of the males of Neocranaus and some considerations about the reproductive biology of N. albiconspersus and N. pectinitibialis are presented.
Empty shells (thanatocoenoses) have been reviewed of species in the genus Granulina (Gastropoda: Granulinidae) from the lower shelf and upper bathyal zones off Mauritania and Western Sahara. We encountered nine species of which four were already known from off Mauritania. Four new species are proposed herein: Granulina reginae sp. nov., G. ronaldi sp. nov., G. sandrae sp. nov. and G. sigridae sp. nov. These four sympatric new species lack labial denticles and they probably form a phyletic clade with a common ancestor. Most hitherto known species in Granulina from the NE Atlantic Ocean and the Mediterranean possess labial denticles. One additional new species with denticles was left in open nomenclature because the material available was considered inadequate. Species in Granulina from Mauritania and Western Sahara have not been found off NW Morocco or the Canary Islands, and the species known from NW Morocco and the Canary Islands have not been found off Mauritania and Western Sahara. The southern extents of the distributions of the Mauritanian species are currently uncertain as additional sampling would be required off Senegal or further South. A key to Mauritanian species in Granulina is given.
A comprehensive review of the enigmatic genus Masona van Achterberg is provided. Two new species are described from the USA: Masona neon Dal Pos & Martens sp. nov. from Puerto Rico, and Masona wow Dal Pos & Martens sp. nov. from California. A key to the world species of Masona is presented, together with a discussion of the morphology of the genus and an annotated catalogue of the species. [Masona] timpaynei Quicke, 2019, is excluded from Braconidae and placed as incertae sedis in the Ichneumonidae, subfamily Neorhacodinae.
Five new and four known species of the genus Dorylaimellus Cobb, 1913 are described and illustrated from the Western Ghats of India. Dorylaimellus attenuatus sp. nov. has a 0.73–0.84 mm long body; lip region rounded, offset; odontostyle attenuated with indistinct lumen, odontophore 9–10 µm long; expanded part of pharynx 39–48% of neck length; female genital system amphidelphic and tail long filiform. Dorylaimellus cylindricaudatus sp. nov. has a 1.0–1.15 mm long body; lip region rounded, offset; odontostyle 4 µm long, odontophore 11–12 µm long; expanded part of pharynx 47–56% of neck length; female genital system amphidelphic, and tail elongate-cylindrical with rounded terminus. Dorylaimellus karnatakensis sp. nov. has a 1.2–1.3 mm long body; lip region rounded, offset; odontostyle 5 µm long, odontophore 12–13 µm long; expanded part of pharynx 45–53% of neck length; female genital system amphidelphic; spicules 22 µm long, ventromedian supplements four, and tail elongate-conoid dorsally convex. Dorylaimellus kasplateauensis sp. nov. has a 0.68–0.83 mm long body, lip region with a weak perioral disc; odontostyle 6–7 µm long, odontophore 11–14 µm long; expanded part of pharynx 38–43% of neck length; female genital system amphidelphic, and tail elongate arcuate conoid. Dorylaimellus tropicus sp. nov. has a 0.6–0.7 mm long body; lip region rounded, offset; odontostyle 4–5 µm long, odontophore 8–10 µm long; hemizonid present; expanded part of pharynx 43–52% of neck length; female genital system amphidelphic and tail elongate-filiform, sudden tapering, ending with rounded tip. Dorylaimellus andrassyi, D. discocephalus, D. belondirelloides, D. chakpilus are redescribed based on the specimens collected from several localities of the Western Ghats.
Two new species of aulacid wasps, Pristaulacus iuliae Turrisi & Nobile sp. nov. from South India (Karnataka) and Pristaulacus ninae sp. nov. from Peninsular Malaysia (Pahang), are described, figured and compared with most related species. Based on the present addition, the Indomalayan area currently includes 27 species of Pristaulacus Kieffer out of 61 known from the whole Oriental Region, which is however a largely underestimated number. An identification key to species and an updated checklist of Pristaulacus occurring in the Indomalayan area are provided.
During exploratory surveys of the fungal diversity in Margalla Hills National Park, Islamabad, we collected a new species of the genus Hymenopellis R.H.Petersen. This is the second report of any species of this genus from Pakistan. Hymenopellis areolata F.Razzaq & Khalid sp. nov. is characterized by an areolate pileus, small basidiospores, and transitional pileipellis (hymeniderm and epithelium) with small pileocystidia. Molecular phylogenetic analyses of the nucleotide sequences of nrITS and nrLSU regions, and morphological data support the description of this new species. A comparison with other closely related species confirmed that the newly described species is distinct from others.
This paper reviews little-known species of the dysderid spider genera Dysdera Latreille, 1804, and Dysderella Dunin, 1992 based on specimens collected in the Caucasus, Middle East, and Central Asia. After combining molecular phylogeny of five mitochondrial and three nuclear genes with morphological evidence, Dysderella is proposed as a junior synonym of Dysdera. In addition, three species are described as new to science: D. jaegeri Bellvert & Dimitrov sp. nov., D. naouelae Bellvert & Dimitrov sp. nov., and D. kourosh Bellvert, Zamani & Dimitrov sp. nov. Four combinations are proposed: Dysdera caspica Dunin, 1990 comb. rev., Dysdera transcaspica Dunin & Fet, 1985 comb. rev., Dysdera elburzica (Zamani, Marusik & Szűts, 2023) comb. nov. and Dysdera sancticedri (Brignoli, 1978) comb. nov. (ex. Dasumia Thorell, 1875). Furthermore, we report a first record of D. festai Caporiacco, 1929 in Turkey and its male cheliceral polymorphism. Our results illustrate the deficiencies that undermine the current taxonomy of this genus. For example, many species are described based on only one or few specimens or limited locality data. The advancements in DNA sequencing technologies applied to museum specimens reduce the need for fieldwork collection and export of fresh specimens. This highlights the significance of museum collections for improving research in this field.
The genus Cerabilia Laporte de Castelnau (Carabidae: Abacetini) is revised for New Zealand. Thirteen species are recognized.
Seven species are described as new: Cerabilia (Cerabilia) cordata Larochelle and Larivière new species, Cerabilia (Cerabilia) kaihoka Larochelle and Larivière new species, Cerabilia (Cerabilia) laevis Larochelle and Larivière new species, Cerabilia (Cerabilia) motunau Larochelle and Larivière new species, Cerabilia (Cerabilia) rugosa Larochelle and Larivière new species, Cerabilia (Cerabilia) takaka Larochelle and Larivière new species, Cerabilia (Cerabilia) willi Larochelle and Larivière new species.
A lectotype is designated for Zabronothus striatulus Broun, 1893.
A revision of all species of Cerabilia (Cerabilia) is provided. Descriptions, an identification key, illustrations of male genitalia, habitus photos, distributional data, and maps are given. Information on ecology, biology, dispersal power, and collecting techniques is included for each species.
ZooBank registration. urn:lsid:zoobank.org:pub:7E3F093D-A5EA-4912-8B30-8380A6F2D890
We propose several nomenclatural changes for taxa in the subfamily Cladodinae and the lampyrid tribes Cratomorphini, Lamprocerini, Lampyrini, Photinini, and Pleotomini in the subfamily Lampyrinae (Coleoptera: Lampyridae). We present evidence for the correct year of description of Calotrachelum flavolineatum Pic, 1941, Diaphanes lateapicalis breveapicalis Pic, 1954, Photinus conradti Pic, 1940, and all species described by Charles Émile Blanchard. Spellings are corrected for Diaphanes lateapicalis breveapicalis Pic, 1954, Diaphanes latipennis dartevillei Pic, 1952, Diaphanes longecarinatus Pic, 1955, Diaphanes moultoni latemarginatus Pic, 1938, Lampyris olivieriana von Heyden, 1890, and Photinus reductemarginalis Pic, 1941. We propose Lucidota puertoestrellaensis Keller and Martin nomen novum to replace Lucidota boliviana Pic, 1927, and Photinus laticollis brasiliensis Keller and Martin nomen novum for Photinus laticollis latior Pic, 1941. Seventy taxa described as variations or aberrations in the tribes Cratomorphini, Lamprocerini, Lampyrini, Photinini, and Pleotomini in the subfamily Lampyrinae by McDermott (1966), are evaluated to subspecies with their availability determined based on ICZN (1999: Article 45.6). Photinus motschulskyi Zaragoza-Caballero, Zurita-Garcia, and Ramírez-Ponce, 2023 is synonymized under Photinus intercalatus Gemminger, 1870. Lastly, we correct the type species of the genus Lucidina Gorham, 1833, and address the dates of publication for fireflies described by Blanchard.
ZooBank registration. urn:lsid:zoobank.org:pub:DCD31B02-DF17-4115-A9A8-A6CBB02D8684
The genus Lichnanthe Burmeister (Coleoptera: Scarabaeoidea: Glaphyridae) has been comprised of eight Nearctic species and contains the only New World members of their family. Here, two new species of arenicolous bumblebee scarabs are described from the central United States. The recent discovery of an undescribed Wyoming species led to the reevaluation of a dubious 134-year-old Nebraska Lichnanthe specimen, indicating that it also was an undescribed species. These two species, Lichnanthe brusti new species from central Wyoming and L. bruneri new species from central Nebraska are immediately distinguishable from all existing species of the genus by their square mandibles.
ZooBank registration. urn:lsid:zoobank.org:pub:401927DA-66D9-47A1-AE68-965210824713
Anthrenus (Anthrenus) muehlei, a new species (Coleoptera: Dermestidae: Megatominae) from Iran
(2024)
A new species, Anthrenus muehlei Holloway and Herrmann (Coleoptera: Dermestidae: Megatominae), from Iran is described. Images of internal and external features are presented. Only female specimens were found and described, but the bursa copulatrix contains obvious sclerites enabling easy differentiation from all other known species from the Palaearctic A. pimpinellae complex. The possible function and taxonomic implication of the sclerites is mentioned.
ZooBank registration. urn:lsid:zoobank.org:pub:97661C20-803E-4785-A4B0-58B16C94D9A2
We describe a new species of karst-dwelling pitviper from Chumphon Province of Peninsular Thailand, in the Isthmus of Kra, based on morphological and molecular data (2427 bp from cyt b, ND4 and 16S rRNA mitochondrial DNA genes). Morphologically, Trimeresurus kraensis sp. nov. is distinguished from other congeners by the following combination of morphological characters: a dark/bottle-green dorsum with reddish-brown or purple crossbands; pale green venter lacking dark dots; stripes present on the lateral sides of the ventrals; internasals generally in contact; one large supraocular scale on each side of the head; iris pale copper; tail brown with dark purplish-brown crossbars; dorsal scales in 21–21–15 rows; ventral scales 167 in a single male, 169–171 in females; subcaudal scales 62 in a single male, 52–54 in females, all paired. White vertebral spots present in males, located on approximately every two or four dorsal scales; dark brown spots forming discontinuous pattern present on 1–3 lateral dorsal scale rows; males with reddish-brown postocular stripe with jagged edges. The new species differs from the morphologically similar species Trimeresurus venustus s. str. by a notable divergence in cytochrome b mitochondrial DNA gene sequences (p = 5.9%).
A re-analysis of the morphological phylogeny of the Nopinae is made, based on an update in the description of Aamunops Galán-Sánchez & Álvarez-Padilla, 2022 and the addition of the recently described genera Nopsma Sánchez-Ruiz, Brescovit & Bonaldo, 2020 and Roddemberryus Sánchez-Ruiz & Bonaldo, 2023. Two new species, Aamunops hoof sp. nov. (male) and Aamunops yiselae sp. nov. (male and female), are also described, which allows a better understanding of the genus morphology and resulted in an emended diagnosis. The description of Aamunops has been updated to include several characteristics of the ultrastructural morphology, legs, chelicerae, palps and female genitalia. The inclusion of these new characters of Aamunops along with those of Nopsma and Roddemberryus in the previous data matrix resulted in a new, completely different hypothesis of the relationships of the nopine genera: Nopsma is part of a group formed by Cubanops and Nyetnops, while Aamunops and Roddemberryus are grouped with representatives of Tarsonops. The four-eyed Nopsides ceralbonus Chamberlin, 1924 was recovered as the most basal species of Nopinae. The relationships among genera of Nopinae and the phylogenetic position of three species, whose taxonomic position is doubtful (Cubanops luquillo Sánchez-Ruiz, Brescovit & Alayón, 2015, Orthonops confuso Galán-Sánchez & Álvarez-Padilla, 2022 and Tarsonops irataylori Bond & Taylor, 2013), is also discussed.
The subgenus Indonthophagus Kabakov, 2006 of Onthophagus Latreille, 1802 (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini) is reviewed. A morphological definition of it is provided. A diagnosis, based both on external and internal characters, of the seven species deemed to belong to the group according to literature, is supplied. As a result, Onthophagus spinifex (Fabricius, 1781), previously placed with some uncertainty in the subgenus, is definitively excluded. A key to Indonthophagus species is also provided, as well as images of male, aedeagus and endophallus for each species. New country records of Onthophagus turbatus Walker, 1858 for Pakistan, Nepal and Bhutan are given.
ZooBank registration. urn:lsid:zoobank.org:pub:9D0CE9F2-35CF-449D-8984-1B4C722762F7
U.S. port of entry interception data revealed that Frankliniella Karny (Thysanoptera: Thripidae) was the most frequently intercepted genus of Thripidae. No published identification resource for intercepted Frankliniella is available to USDA port of entry entomology identifiers. A morphological review of intercepted species was conducted. A dichotomous key for the identification of intercepted slide mounted adult females was created. Morphological diagnosis, description of each species with their geographic region(s) of origin and frequency of interception is provided.
ZooBank registration. urn:lsid:zoobank.org:pub:403B362E-9A7F-4385-A0F1-9DB87FE09AD2
Four new species of the jumping spider genus Stenaelurillus Simon, 1886 are described from India: Stenaelurillus feral Tripathi, Kuni & Kadam sp. nov. (♂, from the Eastern Coastline), S. naldurg Kuni, Kadam & Tripathi sp. nov. (♂♀, from the Deccan Plateau), S. judithbleisterae Kadam, Tripathi & Kuni sp. nov. (♂♀, from the Western Ghats) and S. solapur Kuni, Tripathi & Kadam sp. nov. (♂♀, from the Deccan Plateau). Detailed diagnoses, descriptions, illustrations, natural history and distribution map are provided. Additionally, new locality records for the other five species within the states are presented and mapped.
Two new species of Strandesia Stuhlmann, 1888 are described from the northeastern part of Thailand: S. karanovicae sp. nov. and S. amnatcharoenensis sp. nov. Strandesia karanovicae is characterized by a tumid carapace in dorsal view, a small anterior overlap, the absence of a dorso-subapical seta on the first segment of the antennule (A1), a large aesthetasc Y on the antenna, an α seta shape with a needle-like tip and a large β seta on the mandibular palp (Md-palp), serrated bristles on the maxilla, a long h1 seta on the second thoracopod (T2) and a slender caudal ramus (CR). Strandesia amnatcharoenensis has a small compressed posterior part of the right valves (RV) which makes it closely related to S. pholpunthini Savatenalinton, 2015. The new species can be distinguished primarily by a considerably small anterior overlap of the left valve over the right valve (RV), a postero-ventral flange of the RV, a remarkably large claw Ga of the CR and the chaetotaxy of the limbs, especially A1, Md-palp and T2. In addition, in the present study, the sexual population of S. martensi Savatenalinton, 2015 is recorded for the first time, and thus the first description of the male is provided here. Moreover, the morphological examination of both males and females revealed differences between asexual and sexual females and also points to the fact that S. martensi is a mixed reproduction species. This is the first record of the mixed reproductive mode in the genus Strandesia or even in the Cypricercinae.
Two new species of Palpimanus Dufour, 1820 are described from India: P. godawan Tripathi & Sankaran sp. nov. (♂♀), collected from the Thar Desert in Rajasthan, and P. maldhok Kuni, Tripathi & Sankaran sp. nov. (♂♀), collected from Maharashtra. Images of the endogyne and male palp of the holotype and paratype of P. narsinhmehtai Parajapati, Hun & Raval, 2021 are presented to facilitate its identification. A key to Indian species of Palpimanus and a catalogue of Indian palpimanid spiders are provided. The current distribution of all the known Indian palpimanid spiders is mapped.
A comprehensive checklist of Habenaria from Chapada dos Veadeiros, State of Goiás, was performed alongside morphologic and molecular phylogenetic studies, revealing three new taxa endemic to this region. A total of 61 taxa (59 species and two varieties) of Habenaria are recorded for Chapada dos Veadeiros, representing a two-fold increase compared to previous lists and comprising one of the greatest diversities of the genus in Brazil. Of this total, four taxa are locally endemic. Habenaria cultellifolia, until recently known only from the type collection, was rediscovered in the region after 127 years without records and represents this species’ only known extant population. Three proposed new taxa of Habenaria (H. minuticalcar J.A.N. Bat. & Bianch. sp. nov., H. proiteana J.A.N. Bat., A.A. Vale & Bianch. sp. nov., and H. lavrensis var. xanthodactyla J.A.N. Bat. & Bianch. var. nov.) are corroborated by molecular phylogenetic analyses based on nuclear and plastid markers. They are described, illustrated, tentatively assessed as threatened, and compared to phylogenetically and morphologically related species. Since some areas of this mountain range have not yet been floristically sampled, additional taxonomic novelties and new records are still expected in the future.
A new planthopper genus, Aodingus Chen & Li gen. nov. and three new species (A. hainanensis Chen & Li gen. et sp. nov., A. obscurus Chen & Li gen. et sp. nov. and A. cuongi Chen & Li gen. et sp. nov.) are described from China and Vietnam. The new genus is superficially similar to Procidelphax Bartlett, 2009 in general appearance in that the body is strongly dorsoventrally flattened. Distinctive features of the new taxon include broadly compressed body with vertex broad, apical margin broadly rounded, middle part concave, median and submedian carinae absent, frons broad and large, wider at base than at apex, forewing broad and long, aedeagus tubular, curved ventrally. A diagnosis for all species, illustrations and an identification key of new genus are provided. A key to the Chinese genera of Tropidocephalini is also provided.
Strange new spiders: on Roddenberryus, a new and unusual caponiid genus (Araneae, Caponiidae)
(2023)
The new nopine genus Roddenberryus gen. nov. is erected to include in total five species: three new species, R. kirk gen. et sp. nov. (male and female) from Costa Rica, R. spock gen. et sp. nov. (female) from Campeche, Mexico and R. mccoy gen. et sp. nov. (male) from Baja California Sur, Mexico, as well as two species previously misplaced in Caponina Simon, i.e., R. sargi (Pickard-Cambridge, 1899) gen. et comb. nov. from Guatemala and R. pelegrina (Bryant, 1940) gen. et comb. nov. from Cuba. A male specimen reported as C. sargi Pickard-Cambridge from Costa Rica by E. Kritscher (1957) is assigned to Roddenberryus kirk together with one female collected at the same locality. The new genus is characterized by the distally projected endites and a triangular, projected labium, a conformation unique among nopines; also by a triangular, very short, scaly gladius, serrula with interspersed multiple tooth rows, and a tarsal organ with strongly projected margins. Roddenberryus shares with Tarsonops Chamberlin the cracked tarsi and metatarsi, with multiple adesmatic joints intertwined on the cuticle and an unusual internal respiratory system with both posterior tracheae fused in a single trunk.
The planthopper family Caliscelidae (Hemiptera: Auchenorrhyncha: Fulgoromorpha) and the tribe Adenissini are recorded here for the first time from the Socotra Archipelago (Yemen). A new genus, Oceatra gen. nov., is described for two new species from Socotra Island – Oceatra litoralis gen. et sp. nov. (type species) from coastal sand dunes and salt marshes, and O. scandens gen. et sp. nov. from evergreen montane woodland and shrub communities of the Hagher mountains. Oceatra gen. nov. is similar to Perissana Metcalf, 1952 but differs in the structure of the male style and phallobase. A key to distinguish the genera of the subtribe Adenissina is provided.
Panaeolus punjabensis M. Asif, Q. Firdous, A. Izhar, Niazi & Khalid sp. nov. was collected from three different localities (Bahawalnagar, Kasur, and Lahore) in Punjab, Pakistan. Morphological observations and phylogenetic analyses based on nuclear encoded internal transcribed spacers (ITS1-5.8S-ITS2 = ITS) and D1/D2 domain of large subunit (28S) rDNA confirmed the taxonomic distinctness of this species. The new species is potentially hallucinogenic and characterized by a parabolic pileus with a light brown center, broadly fusiform basidiospores, presence of cheilocystidia, pileocystidia, and caulocystidia, and absence of pleurocystidia and clamp connections. The DNA sequences of the species clustered together in a well-supported distinct clade. We present a detailed description, photographs, and line drawings, and elucidate and discuss the phylogenetic position of the new species. Morphological comparisons with phylogenetically and morphologically allied species are discussed.
The genus Parandes Muir, 1925 (Cixiinae, Andini) is recorded from China for the first time with two new species, Parandes circinatus Wang & Chen sp. nov. and Parandes fuscus Wang & Chen sp. nov. Color images for the adults of the two new species and line drawings for the genitalia are provided. A key is presented to separate all species within the genus.
This work provides the first revision and illustration of the type material of the ammonite species Parahoplites abichi Anthula [Type species of Protacanthoplites Tovbina] and Parahoplites aschiltaensis Anthula [Type species of Acanthohoplites Sinzow] from the upper Aptian (Lower Cretaceous) of Dagestan, Russia. The close affinities and synonymy between these two species are confirmed, and Acanthohoplites aschiltaensis is here retained as the senior valid name by its long quoting history and its historical use as a zonal index of the upper Aptian. The genus Protacanthoplites should be thus synonymised with Acanthohoplites by priority in the date of publication. Comparison with, and distinction from, closely allied Acanthohoplitidae is provided.
We simultaneously considered morphology and molecular phylogeny to modify the generic classification of the ‘pyropterine clade’ (Lycidae, Erotinae, Dictyopterini). To place species previously included in Benibotarus Kôno, 1932 in reciprocally monophyletic genera, we propose Gomezzuritus gen. nov. with the type-species Dictyopterus alternatus Fairmaire, 1856. Further, we transfer Gomezzuritus alternatus (Fairmaire, 1856) comb. nov., G. longicornis (Reiche, 1878) comb. nov., and G. rubripes (Pic, 1897) comb. nov. from Benibotarus to Gomezzuritus gen. nov. The pyropterine clade contains five genera in the Palaearctic region: Pyropterus Mulsant, 1838, Gomezzuritus gen. nov., Helcophorus Fairmaire, 1891, Greenarus Kazantsev, 1995, and Benibotarus Kôno, 1932. The arrangement of longitudinal elytral costae proved misleading for consideration of relationships. Two genera in distant positions share only four primary costae (Pyropterus and Helcophorus), and three similarly distant genera share the shortened primary costa 3, resulting in three primary and four secondary longitudinal costae (Gomezzuritus, Greenarus, and Benibotarus). The larva of Gomezzuritus alternatus is described in detail, and it is compared with the larvae of other Dictyopterini, including the presumed larva of G. longicornis.
Costanana DeLong & Freytag, 1972 is a Neotropical genus of leafhoppers currently comprising 13 species with a wide distribution from Mexico to Argentina. In this study, eight new species of Costanana are described, C. xenomorpha sp. nov. from Department of Cuzco, Peru, and seven Brazilian species: C. alata sp. nov. from Minas Gerais State; C. bifida sp. nov. from Paraná and Mato Grosso states; C. cifi sp. nov., C. obtusa sp. nov. and C. piraquarensis from Paraná State; C. luzi sp. nov. from Distrito Federal; and C. rubromarginata sp. nov. from Mato Grosso and Pará states. We propose that Costanana cella DeLong & Freytag, 1972 is a junior synonym of C. praecellens (Stål, 1862) and C. asymmetrica DeLong & Freytag, 1972 is a junior synonym of Gypona viridans DeLong & Martinson, 1972. Costanana costata DeLong & Freytag, 1972 and C. flavicosta (Stål, 1862) are transferred to Gypona Germar, 1821, C. minuta (Spångberg, 1878) is transferred to Acuponana DeLong & Freytag, 1970 and Gypona nupera Van Duzee, 1907 is transferred to Costanana. Gypona costata comb. nov. and Costanana santana DeLong & Wolda, 1983 are newly recorded from Pernambuco State, and Gypona flavicosta comb. nov. is recorded from Espírito Santo and Paraná states, Brazil. The female genitalia of Costanana flavina DeLong & Freytag, 1972, C. praecellens and Gypona flavicosta comb. nov. are described and illustrated for the first time and a brief note on feeding behavior is given for C. flavina and C. praecellens. A key to males and a new diagnosis are provided for Costanana. We also describe and illustrate Metacostana cornuta gen. et sp. nov., a new Brazilian genus for Gyponini sharing characteristics with Costanana, Acuthana Domahovski & Cavichioli, 2018, Domahovana Silva et al., 2022 and Dumorpha DeLong & Freytag, 1975.
Scelolabes, historically is a monotypic genus of Ocydromiinae (Hybotidae) that for a long time was not studied due to the absence of the type of the type species (Scelolabes bivittatus Philippi), which is probably lost or destroyed. In this context, we propose an updated diagnosis and redescription of the genus, based on specimens identified by experts and in agreement with the original description of Philippi (1865), as well as illustrating male and female terminalia for the first time. This serves as an aid to discuss and corroborate the status of Scelolabes in the Neotropical region as a genus distinct from Hoplopeza. In addition, two new species are described, and an identification key and a distribution map are provided to all the Neotropical species of the genus.
Three new species of the genus Pelodera Schneider, 1866 viz., P. indica sp. nov., P. adeeli sp. nov. and P. paratretzeli sp. nov. collected from dung beetles and P. cylindrica (Cobb, 1898) collected from soil samples, are described and illustrated. Pelodera indica sp. nov. is characterised by sexual dimorphism in anterior region, cupola-shaped tail with a spike; males having punctated, striated and lobed bursa with no genital papillae originating anterior to cloaca. Pelodera adeeli sp. nov. is characterised by coarsely annulated cuticle; relatively narrow stoma; tail conoid without spike; males with punctated, lobed bursa and nine pairs of genital papillae arranged in a 2/1+2+P+3+1 configuration. Pelodera paratretzeli sp. nov. is characterised by sexual dimorphism in anterior region, stoma wide with three well-developed metastegostomal denticles; tail cupola-shaped with a long spike; males having spicules fused distally up to 12–14% of spicule length; bursa peloderan, anteriorly closed and punctated with nine pairs of genital papillae arranged in a 3/2+P+3+1 configuration. Pelodera cylindrica is described with additional details. The comparative analysis as well as phylogenetic relationship of the species belonging to the coarctata group have been elaborated by incorporating scanning electron microscopic observations. Information on the biogeographical distribution has also been provided.
Two species of Xorides Latreille, 1809 are reported parasitizing wood-boring insects in trunks and relatively larger twigs of Juglans mandshurica Maxim. in Kuandian Manzu Autonomous County, Liaoning, in the Palaearctic part of China. Two new species are described: X. juglanse Sheng, Broad & Sun sp. nov. and X. kuandianense Sheng, Broad & Sun sp. nov. One species, X. sapporensis (Uchida, 1928), was associated with wood-borers in J. mandshurica Maxim. for the first time. A key to the 46 species of Xorides Latreille known from China is provided.
Seven new species of Schismatothele Karsch, 1879 (Araneae, Theraphosidae) are described, almost doubling the diversity of the genus: S. caeri sp. nov.; S. caiquetia sp. nov.; S. merida sp. nov.; S. moonenorum sp. nov.; S. quimbaya sp. nov.; S. timotocuica sp. nov. and S. wayana sp. nov. An identification key for all species of Schismatothele (except S. kastoni) is presented, as well as a complementary diagnosis for the genus. Also, a standardized nomenclature is proposed to describe the prolateral keels of male palpal bulbs of species of Schismatothele.
Three new species of pseudoscorpion, Allochthonius lini sp. nov. (Xiaoguoquan Cave) and Selachochthonius yinae sp. nov. (Xiao Cave) from Yunnan Province, Allochthonius xuae sp. nov. (Yelaoda Cave) from Guizhou Province, are described and illustrated. An identification key is provided for all known Chinese representatives of the family Pseudotyrannochthoniidae.
We describe and illustrate Croton restingae Sodré & Riina sp. nov., a new species endemic to a restricted area in the state of Rio de Janeiro, Brazil, characterized by a particular type of vegetation of the Atlantic Forest known as restinga. The new species belongs to Croton sect. Adenophylli subsect. Laceratoglandulosus, which is supported by morphology and a phylogenetic analysis based on nrDNA ITS sequence data. Croton restingae is most closely related to C. echioides and C. laceratoglandulosus, also in subsect. Laceratoglandulosus, with which it shares the fruit columella with three non-inflated, flat or slightly ascending apical appendages. Croton restingae differs from them by its sessile or shortly stipitate leaf nectaries, revolute pistillate sepals, styles distally 2-fid and basally united forming a column, and larger capsules (7.5–8 × 8.5–9.3 mm). Croton restingaeʼs habitat, conservation status, phenology, morphology and phylogenetic relationships are discussed, and a map with its geographic distribution is also included.
Here I describe a new genus, Iviephengus gen. nov., based on a single species, Iviephengus ferreirai gen. et sp. nov., from Peru. This new genus is characterized by the following combination of characters: interantennal distance close to 3 × the antennal socket length; antenna 12-segmented, IV to XI each with two long symmetrical compressed and apically slightly enlarged branches; labrum fused to frontoclypeus; mandibles short, obliquely crossed, each with a notch on the external margin to fit the other mandible and without extra teeth; maxillary palpi 4-segmented, last segment digitiform; labial palpi 2-segmented; posterior tentorial pit consisting of a single small fossa; wing with radial cell closed and transverse, vein r4 interrupted, r3 absent; first tarsomere of pro- and mesotarsus with a ventral comb covering the posterior half of the tarsomere; claws simple, without any teeth; aedeagus with paramere symmetrical, apex unevenly round, toothed inward, with short and sparse bristles. I provide a key to Mastinocerinae genera with 12-segmented antennae and the first pro- and mesotarsomere with ventral combs. I also provide illustrations for the diagnostic features for this new genus. Finally, I discuss the presence and function of some modifications in the mandible and the sternite VIII in Phengodidae and other Coleopteran families.
Inocybe hopeae sp. nov. and first record of Pseudosperma keralense (Inocybaceae) from Thailand
(2023)
Based on genetic studies, supported further by morphological and ecological differences, we present a taxonomic novelty (Inocybe hopeae Raghoonundon & Raspé sp. nov.) and a new geographical record (Pseudosperma keralense) from forests of Northern Thailand. Inocybe hopeae is characterized by medium-sized basidiomes, brownish orange to brown pileus that is darker towards the margin, off-white to pale brown context, light brown to dark brown stipe with off-white basal mycelium and pale brown to grayish brown lamellae. A three-gene phylogeny (LSU, tef1, rpb2) coupled with macroscopic / microscopic descriptions and illustrations are provided confirming the species’ positions in their respective generic clades. Inocybe hopeae was sister to I. thailandica with strong support (BS = 100%, PP = 1.0). Our Thai collections of OR1629 had similar morphological characters and 100% identical sequences with the holotype of Pseudosperma keralense from India.
During mycological explorations, a new smooth spored species, Inocybe subhimalayanensis Razzaq, Naseer & Khalid sp. nov. was collected from moist temperate sub-Himalayan region, Pakistan. Phylogeny of ITS and LSU regions of nrDNA, and morphoanatomical data make it distinct from other known species of the genus. The taxon is characterized by: a yellowish orange to brown pileus with prominent fibrillose, prominent umbo; ellipsoid to amygdaliform smooth larger basidiospores (8.4‒)8.6‒12.2(‒12.6) × (4.9‒)5.1‒7.1(‒7.3) µm; and lack of velipellis. Molecular phylogenetic analyses further support the recognition of the new species.
From 2004 to 2012, ten multidisciplinary oceanographic surveys were conducted along the coast of Northwest Africa, between the Strait of Gibraltar and the northern border of Sierra Leone. A total of five species of Euryalida Lamarck, 1816 belonging to three families were captured at 29 of the 1298 stations sampled in the area. Among them, Astrodendrum juancarlosi sp. nov. is described and figured in this paper. Ophiocreas oedipus Lyman, 1879 is recorded for the first time on West African continental margin and Gorgonocephalus pustulatum (H.L. Clark, 1916), an Indo-Pacific species only known from South African coast in the Atlantic, is reported off Guinea-Bissau, greatly extending to the North its Atlantic distribution. In addition, Asteroschema inornatum Koehler, 1906, a northeast Atlantic species, is recorded for the first time in African waters, off Western Sahara, extending its range of distribution to the south. Our data also extend the bathymetric distribution of A. inornatum to shallower waters and of G. pustulatum to deeper waters. The association of some euryalids with certain species of pennatulaceans and gorgonians is also described.
Description of three new Acanthocinini (Coleoptera: Cerambycidae: Lamiinae) species from Ecuador
(2023)
Three new species of Acanthocinini (Coleoptera: Cerambycidae: Lamiinae) are described from Napo province, Ecuador: Anisopodus micromaculatus new species; Parabaryssinus katerinae new species; and Paracleodoxus minutus new species. A key to species of Paracleodoxus Monné and Monné (2010) is provided.
ZooBank registration. urn:lsid:zoobank.org:pub:E7C66DA1-6F5F-4F94-922E-43E0B83331DD
The Egyptian fauna of mealybugs (Hemiptera: Coccoidea: Pseudococcidae) is reviewed and an illustrated key to the 30 genera and 54 species is provided. Phenacoccus madeirensis Green is reported for the first time in Egypt. A new genus, Ezzatacoccus Evans and Abd-Rabou, is described and illustrated with Amonostherium arabicum Ezzat, 1960 designated as its type species. Octococcus salicicola Priesner and Hosny, 1935 is reinstated as a valid taxon and transferred to Misericoccus Ferris, new combination. Ripersia cressae Hall is transferred to Maconellicoccus Ezzat, new combination and Planococcus lindingeri (Bodenheimer) is transferred back to Formicococcus Takahashi, revised status.
ZooBank registration. urn:lsid:zoobank.org:pub:8CA7B000-E8D4-463D-95B0-431BA0A7BA57
A catalog of the species of Trichodesma LeConte (Coleoptera: Ptinidae) for the world is provided, including synonyms and misspellings. Six additional genera, Anobiopsis Fall, Belemia Español, Nanodesma Zahradník, Nicobium LeConte, Trichobiopsis White, and Trichodesmina Español, are also cataloged for completeness. The species name, author, year, and page number of description are given for each species, as well as references for combinations by different authors. The type depository is given when known, and followed by a “?” when unknown but suspected, and a general distribution is given by region of the world followed by country. Authorship of the genus Trichodesma in Lepidoptera is transferred to Schaus.
ZooBank registration. urn:lsid:zoobank.org:pub:99BDA976-4B58-4D42-8CF5-4D3034B95C7B
We provide a floristic account for the Begoniaceae of Peru. The family is represented in Peru by 76 species, which are all members of the genus Begonia and represent eighteen sections. Twelve new species are described: B. condorensis Jara & Moonlight sp. nov., B. deltoides Moonlight sp. nov., B. huancabambae Moonlight sp. nov., B. imbrexiformis Moonlight sp. nov., B. longinqua Moonlight sp. nov., B. longitepala Moonlight sp. nov., B. nunezii Moonlight sp. nov., B. occultata J.P.Allen & Moonlight sp. nov., B. pedemontana Moonlight sp. nov., B. serratistipula Moonlight sp. nov., B. vargasii Moonlight sp. nov. and B. yuracyacuensis Moonlight sp. nov. We also provide four new records for the country: B. andina Rusby, B. brandbygeana L.B.Sm. & Wassh., B. neoharlingii L.B.Sm. & Wassh. and B. unilateralis Rusby. We provide an identification key to all species. The ecology, distribution, and conservation status of all Peruvian Begonia species are discussed, including provisional IUCN threat assessments. Most species are illustrated by either historical illustrations, contemporary photographic plates, or line drawings. Twenty-six names are newly synonymised including fifteen previously accepted species, and we designate eighty-three lectotypes, four neotypes, and four epitypes.
Metafruticicola is a diverse land snail genus inhabiting the north-eastern Mediterranean region from S Albania to Israel. In this study, we describe Metafruticicola kavafis sp. nov., a new species for science from Kasos Island. We also focus on the reproductive system of steno-endemic species and subspecies of the genus from small islands of the Aegean Archipelago, whose anatomy was previously completely unknown. These species are M. crassicosta, M. pieperi, M. coartata gemina and M. nicosiana conciliatrix. For the latter subspecies, only the outlines of the genitalia were previously given. Additionally, we investigated in detail for the first time the genital anatomy of M. nicosiana nicosiana from Cyprus. Hitherto, information on the genital apparatus of this subspecies was based on sparse data derived from sexually immature specimens. These new anatomical data contribute to a plethora of controversies concerning the subgeneric division of Metafruticicola, which was based solely on conchological features and especially the microsculpture of the teleoconch. We suggest that the current subgeneric division of the genus should be abandoned.
The early Tournaisian (Early Carboniferous; Mississippian) ammonoids from the classical abandoned limestone quarry of Gattendorf (Upper Franconia) are revised, using the historical collections as well as so far undescribed material. The ammonoid assemblage is composed of prionoceratid ammonoids of the six genera Mimimitoceras, Paragattendorfia, Stockumites, Acutimitoceras, Gattendorfia and Gattenpleura, which indicate a stratigraphic position near the Devonian–Carboniferous boundary in the earliest Carboniferous. The new species Stockumites hofensis sp. nov. and S. nonaginta sp. nov. are described.
The taxonomy of Loboscelidiinae in Vietnam is revised, with 16 new species being described: Loboscelidia bachmaensis sp. nov., L. barbata sp. nov., L. cilia sp. nov., L. convexa sp. nov., L. cucphuongensis sp. nov., L. cuneata sp. nov., L. do sp. nov., L. flavipes sp. nov., L. glabra sp. nov., L. komedai sp. nov., L. mediata sp. nov., L. parallela sp. nov., L. piriformis sp. nov., L. squamosa sp. nov., L. vang sp. nov. and L. vietnamensis sp. nov. In total, 24 species of Loboscelidia are recognized in the fauna of Vietnam. Keys to Indo-Chinese male and world female of Loboscelidia are provided. A brief observation of the foraging behavior of L. squamosa sp. nov. is also reported. Host-carriage and subsequent host egg burying are considered primary nesting behaviors of solitary wasps.
Eight new species of Itauara Müller, 1888 are described based on specimens collected in southeastern Brazil: I. bispinata sp. nov., I. caparao sp. nov., I. cipoensis sp. nov., I. cristata sp. nov., I. holzenthali sp. nov., I. mangaratiba sp. nov., I. robertsonae sp. nov., and I. rupicola sp. nov. In addition, we provided an updated distributional list of species of Itauara, with new records of I. jamesii Robertson & Holzenthal, 2011 and I. lucinda Robertson & Holzenthal, 2011 for Espírito Santo State (Brazil), I. plaumanni (Flint, 1974) for Paraná State (Brazil), and I. tusci Robertson & Holzenthal, 2011 for Minas Gerais State (Brazil). We performed an equal weighted parsimony analysis adding the new species to the dataset provided by Robertson & Holzenthal (2013), with modification of the interpretation of some morphological characters. The genus was recovered as monophyletic, but overall statistic support for clades was weak. Itauara is restricted to South America and has a disjunct distribution, with some species occurring in northern South America and others in southeastern South America. Probably, the diversification of Itauara in South America is related to the connections between the Amazon Forest and the Atlantic Forest through historical events at different times.
A new species of Strepsiptera of the genus Paraxenos Saunders, 1872 (Xenidae) from the United Arab Emirates is described. It was recorded from the host species Bembix kohli Morice, 1897 and represents the first occurrence of Paraxenos from Bembix Fabricius, 1775 in the Afrotropical region. A detailed redescription of the female cephalothorax of Paraxenos hungaricus (Székessy, 1955) is provided, together with the first description of the male cephalotheca. The holotype of Paraxenos krombeini Kifune & Hirashima, 1987 was redescribed. Additionally, a key for parasites of Bembix among Paraxenos species is provided based on characters of the female cephalothorax and male cephalotheca. The distribution and conservation status of Paraxenos spp. on Bembix are also discussed.
The number of currently described species of Afrotropical parasitoid wasps does not reflect the true species diversity. One of the most severely understudied parasitoid wasp groups is Ceraphronoidea. In this first study on Afrotropical mainland Ceraphronoidea in more than 20 years, which is also the first ever taxonomic monograph focusing on Ceraphronidae, we describe 88 new species of Ceraphronidae (85 new species) and Megaspilidae (3 new species) from Kakamega Forest (Kenya), Mt. Kilimanjaro (Tanzania) and Ivindo NP (Gabon): Aphanogmus abaluhya sp. nov., A. ashitakai sp. nov., A. idakho sp. nov., A. ikhongamurwi sp. nov., A. isiukhu sp. nov., A. kakamegaensis sp. nov., A. lateritorum sp. nov., A. mangimelii sp. nov., A. mariae sp. nov., A. mashariki sp. nov., A. nehbergi sp. nov., A. njia sp. nov., A. vestrii sp. nov., A. yala sp. nov. (all clavicornis species group), A. dimidiatus sp. nov., A. fraterculus sp. nov., A. guenteri sp. nov., A. kakakili sp. nov., A. kisiwa sp. nov., A. maua sp. nov., A. morriconei sp. nov., A. ndefu sp. nov., A. ngai sp. nov., A. nikii sp. nov., A. pilosicoxa sp. nov., A. rafikii sp. nov., A. robustus sp. nov., A. simbai sp. nov., A. taji sp. nov., A. ukanda sp. nov. (all fumipennis species group), A. campanula sp. nov., A. kikuyu sp. nov., A. pagoda sp. nov. (all tenuicornis species group), Ceraphron banda sp. nov., C. brashi sp. nov., C. breviharpis sp. nov., C. breviscapus sp. nov., C. buyangu sp. nov., C. chemositi sp. nov., C. cingulum sp. nov., C. clavatumeris sp. nov., C. digiti sp. nov., C. eaerendili sp. nov., C. ekero sp. nov., C. ellae sp. nov., C. eulbergi sp. nov., C. herreni sp. nov., C. hitagarciai sp. nov., C. insolitus sp. nov., C. isecheno sp. nov., C. isukha sp. nov., C. ivindoensis sp. nov., C. kaharabu sp. nov., C. kaimosiensis sp. nov., C. kakamegaensis sp. nov., C. kidole sp. nov., C. kimathii sp. nov., C. lirhanda sp. nov., C. longiharpis sp. nov., C. longisetae sp. nov., C. longumerunus sp. nov., C. maathaiae sp. nov., C. malava sp. nov., C. mamamutere sp. nov., C. metapleuralis sp. nov., C. mikoi sp. nov., C. mwekaensis sp. nov., C. nandi sp. nov., C. nzoia sp. nov., C. onesimusi sp. nov., C. pilosiharpis sp. nov., C. pleurosulcus sp. nov., C. reinholdi sp. nov., C. salazar sp. nov., C. sataoi sp. nov., C. semira sp. nov., C. sungura sp. nov., C. tenuimeris sp. nov., C. tiriki sp. nov., C. trietschae sp. nov., Cyoceraphron dhahabudorsalis sp. nov., C. harpe sp. nov., C. invisibilis sp. nov., C. kahawia sp. nov., C. njano sp. nov. (all Ceraphronidae), Conostigmus kijiko sp. nov., C. koleo sp. nov., and Dendrocerus wachagga sp. nov. (all Megaspilidae). In addition, we describe four species of Aphanogmus and five species of Ceraphron without formal naming. A neotype is designated for Dendrocerus anneckei Dessart, 1985 (Megaspilidae). With these new species we more than double the number described from the Afrotropical mainland (65 vs 153). The species numbers found allow us to estimate the real worldwide species number of Ceraphronoidea as being roughly 12 000–21 000, i.e., 16–29 times the number of the currently described species (~730, including the species described herein). This study is meant to highlight that it is necessary and also possible to study the parasitoid wasps of tropical regions and provide momentum for exploring the diversity of small and diverse insect groups in the Afrotropics and elsewhere while also providing the basic knowledge that is much needed for protecting biodiversity and understanding evolution and the networks of life on earth. All described species are diagnosed and illustrated, with focus on the male genitalia. Furthermore, we provide an identification key to males of Afrotropical Ceraphronidae.
The genus Lacronia Strand, 1942 is herein revised, and a maximum parsimony phylogenetic analysis of morphological characters (30 taxa, 115 characters, 2962 scorings) is performed to test its monophyly. As a result, Lacronia is herein made monophyletic by means of the inclusion of Discocyrtus boraceae B. Soares, 1942, Discocyrtus niger Mello-Leitão, 1923 and Discocyrtus tenuis Roewer, 1917. Lacronia including those species is the sister group of Discocyrtus s. str. inside the DRMN-clade, and should be removed from Pachylinae Sørensen, 1884, although without a new subfamilial assignment for now. Four new junior subjective synonyms are detected for the first time (Discocyrtus rarus B. Soares, 1944 = Discocyrtus fazi Piza, 1942 = Discocyrtus niger Mello-Leitão, 1923; Discocyrtus infelix Mello-Leitão, 1940 = Discocyrtus nigrolineatus Mello-Leitão, 1923 = Discocyrtus tenuis Roewer, 1917). Accordingly, the following new combinations are proposed: Lacronia boraceae (B. Soares, 1942) comb. nov., Lacronia nigra (Mello-Leitão, 1923) comb. nov. and Lacronia tenuis (B. Soares, 1942) comb. nov. Lacronia is endemic to the Atlantic province of Brazil, with verified records from the states of Rio de Janeiro, Santa Catarina and São Paulo. As a geographic note, the record of ‘D. fazi’ for Chile is here discussed and considered incorrect.
Neotropical Xiphocentronidae may have arrived in South America from Central America in the late Miocene or via the Greater Antilles land bridge during the Oligocene. This would give from 10 to 30 Mya of diversification of the family in South America. However, only 11 species were previously known from the Andean foothills. In this study, five new species are described from Peru, four of Xiphocentron (X. ashaninka sp. nov., X. harakbut sp. nov., X. matsigenka sp. nov., X. yine sp. nov.) and one of Machairocentron (M. amahuaca sp. nov.). The new species are most similar to species described from the Yungas of Argentina, and the Pacific dominion of Colombia and Venezuela. The association with species from the Pacific may suggest a species divergences prior to the major Andean uplift and the Amazon basin formation. The spine-like setae on the basal region of the inferior appendage of Xiphocentron were recognized as topologically homologous to the setal brushes on the ventral projection of Caenocentron. Furthermore, based on the morphology of male and female genitalia of Machairocentron, a mating position different from that described for Psychomyiidae is inferred.
Four new crane fly species of the subgenus Tipula (Vestiplex) Bezzi, 1924, T. (V.) medialobata sp. nov. (Yunnan), T. (V.) singularis sp. nov. (Yunnan), T. (V.) cibagouensis sp. nov. (Tibet) and T. (V.) paramonovi sp. nov. (Tibet) are described and illustrated. Redescription and detailed illustrations of T. (V.) nestor Alexander, 1934 are provided with first distributional record in mainland China.
Two species of the genus Ophelina Örsted, 1843 (Annelida, Opheliidae) are reported from the coast of Kuwait (Arabian Gulf) after specimens collected in the intertidal and shallow subtidal, namely Ophelina arabica sp. nov. and Ophelina grandis (Pillai, 1961). The new species is mainly characterised by features of the anal tube, which is provided with about 25 annulations at each side; the ventral margins are fully fused while dorsal margins are fused at most of their length but are free at the distal end in the shape of a conspicuous incision; the posterior end is opened with free margins; the anal tube also lacks marginal papillae but bears a pair of basal papillae and an unpaired anal cirrus attached to ventral margin at mid-length. Ophelina grandis is reported for the first time in the Arabian Gulf; specimens are fully described and compared with similar species. A key for species of Ophelina in the Indo-Pacific, Southern Asia, Indo-Malay Archipelago and Australia, is also provided.
Two new species, Russula pseudopunicea C.L.Hou, G.Q.Cheng & H.Zhou sp. nov. and R. wulingshanensis C.L.Hou, G.Q.Cheng & H.Zhou sp. nov., from Yanshan mountains in North China are described herein based on morphological and phylogenetic analyses of nrITS, and nrLSU-rpb2-mtSSU gene regions. Morphologically, R. pseudopunicea sp. nov. is characterised by a reddish brown, light brown to brownish orange pileus with a greyish yellow margin, subglobose to broadly ellipsoid basidiospores with warts forming a partial reticulum and pleurocystidia turning grey to purplish red in sulfovanillin. Russula wulingshanensis sp. nov. is characterised by a purple pinkish pileus with a grey-white to grey-purple margin, subglobose to broadly ellipsoid basidiospores with isolated warts, and pileocystidia turning black in sulfovanillin. Phylogenetic and morphological analyses resolved the two species in Russula subg. Heterophyllidia. Russula pseudopunicea sp. nov. and R. wulingshanensis sp. nov. were placed in the lineages of subsect. Virescentinae and subsect. Griseinae, respectively.
The freshwater snail genus Mercuria is widely distributed in lowland waters across Western Europe, Northern Africa and the Mediterranean islands. Approximately two-thirds of the currently recognised species are described based on their shell morphology, which may vary within species due to biotic and abiotic factors. Recent molecular phylogenies that included numerous previously documented populations recovered 14 species clades, nine of which correspond to nominal species and five, to undescribed taxa. Here, we formally describe the five undescribed taxa as new species and provide morphological descriptions of the shell and other anatomical structures for three of the other inferred clades and for the species M. maceana to elucidate their taxonomic status and assess the utility of morphological characters for species delimitation in Mercuria. Taken together, the morphological and molecular evidence suggest new identifications and synonymies, having implications on the known geographic range of the studied species, including the type species M. similis. Anatomical measurements and geometric morphometric analysis of shell shape revealed no clear differentiation among the species analysed, predicting the importance of molecular data in elucidating the species diversity of the genus.
A synthesis of the Phaeogenini occurring in the Afrotropical region is provided. Three species are newly described: Centeterichneumon nambi Dal Pos, Diller & Di Giovanni sp. nov. from Uganda, Chauvinia ganota Claridge sp. nov. from Kenya, and Kibalus nonnaritae Dal Pos & Di Giovanni sp. nov. from Uganda. Heterischnus mfongosi Rousse & van Noort, 2013 is newly recorded for Kenya and Tanzania and the male of the species is diagnosed for the first time. Also, the female of Arearia oxymoron Rousse & van Noort, 2013 is diagnosed for the first time from one of the paratype localities. Lusius tenuissimus (Heinrich, 1938) and Chauvinia nyanga Rousse & van Noort, 2013 are recorded for the first time for Uganda and Kenya, respectively. In addition, new localities are given for Chauvinia nitida (Heinrich, 1938), Heterischnus olsoufieffi (Heinrich, 1938) and Hoplophaeogenes curticornis Heinrich, 1938. A new combination, Nesostenodontus mkomazi (Rousse & van Noort, 2013) comb. nov., is proposed to accommodate Heterischnus mkomazi. An updated key to the Afrotropical genera of Phaeogenini and keys to the Afrotropical species of the genera Arearia Seyrig, Centeterichneumon Heinrich, Chauvinia Heinrich, Heterischnus Heinrich, Hoplophaeogenes Heinrich, Kibalus Rousse, van Noort & Diller, and Lusius Tosquinet are provided. Updated online Lucid keys to genera and species are available from http://www.waspweb.org.
Three new deep-sea species of Thyasiridae (Mollusca: Bivalvia) from the northwestern Pacific Ocean
(2023)
The Thyasiridae is one of the species-richest families in the abyssal and hadal zones of the northwestern Pacific Ocean. Many thyasirid species dominate benthic communities in terms of abundance and play an important role in the functioning of deep-sea ecosystems. Most of the thyasirid species in the region are new to science and have not been described. Based on the material collected from 1954 to 2016 by seven deep-sea expeditions, three new species of Thyasiridae (Parathyasira coani sp. nov., P. pauli sp. nov., and Thyasira kharkovensis sp. nov.) are described from the abyssal and hadal zones (3210–7540 m depth) of the Sea of Okhotsk, the Bering Sea, as well as the Kuril-Kamchatka and Japan trenches. The new species are remarkable among their congeners due to the combination of the following characters: an obliquely-rhomboidal shell with a weak and shallow posterior sulcus and a large prodissoconch with sculpture of lamellated folds. Comparisons with related species are provided.
Based on molecular and morphological data of four specimens of Pareas Wagler, 1830 collected from the type locality of P. yunnanensis (Vogt, 1922), along with examination of the type specimens of P. yunnanensis, we revalidate this poorly known, secretive species. Furthermore, based on molecular and morphological lines of evidence we also describe a new species of Pareas from Xishuangbanna Prefecture, Yunnan Province, China. Morphologically, the new species closely resembles its sister species P. nigriceps Guo & Deng, 2009. However, the new species is divergent from the latter in cytochrome b mtDNA gene sequences, and can be distinguished from all congeners by the following combination of morphological characteristics: single preocular, postocular fused with subocular, loreal not bordering orbit, vertebral scales enlarged, 3–5 rows of mid-dorsal scales keeled at the middle of the body, ventral scales 160–171; subcaudals 62–64, dorsal surface of head solid black or reddish-brown, dark nuchal band present, iris brownish-black or reddish-brown.
A new genus of Lebinthina (Orthoptera: Gryllidae: Eneopterinae) is erected based on species from Maluka Islands near northern Sulawesi (Indonesia): Platybinthus gen. nov. This new genus currently consists of three species. Platybinthus punctatus (Brunner von Wattenwyl, 1898) gen. et comb. nov. from Halmahera Island is assigned as the type species. Platybinthus striolatus gen. et comb. nov., also from Halmahera Island, is redescribed. We also describe a new species: Platybinthus sandyi gen. et sp. nov. from Morotai Island.
Analyses of whole genomic shotgun datasets, COI barcodes, morphology, and historical literature suggest that the following 13 butterfly species from the family Hesperiidae (Lepidoptera: Papilionoidea) in Texas, USA are distinct from their closest named relatives and therefore are described as new (type localities are given in parenthesis): Spicauda atelis Grishin, new species (Hidalgo Co., Mission), Urbanus (Urbanus) rickardi Grishin, new species (Hidalgo Co., nr. Madero), Urbanus (Urbanus) oplerorum Grishin, new species (Hidalgo Co., Mission/Madero), Telegonus tsongae Grishin, new species (Starr Co., Roma), Autochton caballo Grishin, new species (Hidalgo Co., 6 mi W of Hidalgo), Epargyreus fractigutta Grishin, new species (Hidalgo Co., McAllen), Aguna mcguirei Grishin, new species (Cameron Co., Brownsville), Polygonus pardus Grishin, new species (Hidalgo Co., McAllen), Arteurotia artistella Grishin, new species (Hidalgo Co., Mission), Heliopetes elonmuski Grishin, new species (Cameron Co., Boca Chica), Hesperia balcones Grishin, new species (Travis Co., Volente), Troyus fabulosus Grishin, new species (Hidalgo Co., Peñitas), and Lerema ochrius Grishin, new species (Hidalgo Co., nr. Relampago). Most of these species are known in the US almost exclusively from the Lower Rio Grande Valley in Texas. Nine of the holotypes were collected in 1971-1975, a banner period for butterfly species newly recorded from the Rio Grande Valley of Texas; five of them collected by William W. McGuire, and one by Nadine M. McGuire. At the time, these new species have been recorded under the names of their close relatives. A Neotype is designated for Papilio fulminator Sepp, [1841] (Suriname). Lectotypes are designated for Goniurus teleus Hübner, 1821 (unknown, likely in South America), Goniloba azul Reakirt, [1867] (Mexico: Veracruz) and Eudamus misitra Plötz, 1881 (Mexico). Several taxonomic changes are proposed. The following taxa are species (not subspecies): Spicauda zalanthus (Plötz, 1880), reinstated status (not Spicauda teleus (Hübner, 1821)), Telegonus fulminator (Sepp, [1841]), reinstated status (not Telegonus fulgerator (Walch, 1775), Telegonus misitra (Plötz, 1881), reinstated status (not Telegonus azul (Reakirt, [1867])), Autochton reducta (Mabille and Boullet, 1919), new status (not Autochton potrillo (Lucas, 1857)), Epargyreus gaumeri Godman and Salvin, 1893, reinstated status (not Epargyreus clavicornis (Herrich-Schäffer, 1869)), and Polygonus punctus E. Bell and W. Comstock, 1948, new status (not Polygonus savigny (Latreille, [1824])). Urbanus ehakernae Burns, 2014 and Epargyreus socus chota Evans, 1952 are junior subjective synonyms of Urbanus alva Evans, 1952 and Epargyreus clavicornis (Herrich-Schäffer, 1869), respectively, and Epargyreus gaumeri tenda Evans, 1955, new combination is not a subspecies of E. clavicornis.
ZooBank registration. https://zoobank.org/D5462F9E-E08D-46C6-898D-76EE7466DD19
Abstract. More than 1300 specimens of Eucnemidae collected from Heredia Province in Costa Rica during the 1990s Arthropods of La Selva (ALAS) survey were studied from 2018 through 2022. One new genus of false click beetle, Absensiugum Otto, Muona and Córdoba-Alfaro, is described. Nematodes teres Horn, from the Nearctic and Caribbean regions, is transferred to this new genus to form Absensiugum teres, new combination. Sixteen new species of false click beetle (Coleoptera: Eucnemidae) are described from Costa Rica. These new species are: Adelothyreus brevis, Adelothyreus costaricensis, Adelothyreus totus, Quirsfeldia stethonoides, Lacus pectinatus, Maelodrus costaricensis, Onichodon confluentus, Onichodon rufus, Isarthrus striatus, Absensiugum brunneum, Dromaeolus americanus, Dromaeolus brunneus, Dromaeolus herediensis, Dromaeolus holdridgei, Deltometopus bicolor and Nematodes apicalis. Three additional records outside of the Heredia Province from the Osa Peninsula and Panama for Lacus pectinatus are included in this study. Identification keys are provided for species of Adelothyreus Chevrolat, Onichodon Newman, Dromaeolus Kiesenwetter, Deltometopus Bonvouloir and Nematodes Berthold in Costa Rica. Diagnostic differences are briefly noted for each species within the Neotropical region. A list of Eucnemidae from Heredia Province is provided.
ZooBank registration. urn:lsid:zoobank.org:pub:C1D5B819-A964-4679-B090-84CDBBC59D6A
Four new species of Coleoptera are described: Pentanodes clavatus new species, from Ecuador; Pentanodes baldwini new species, from Panama (Cerambycidae: Cerambycinae: Tillomorphini); Elytrimitatrix (Grossifemora) dilatata new species, from Mexico (Disteniidae) and Novantinoe falsa new species, from Mexico (Disteniidae). Taxonomical notes on Euderces cribellatus (Bates, 1885) (Tillomorphini) and Oreodera Audinet-Serville, 1835 (Lamiinae: Acrocinini) are provided. Additionally, new records are provided for Oreodera noguerai McCarty, 2001 (new state record), Adetus croton Heffern, Santos-Silva and Botero, 2019 (new country record), Icimauna sarauaia Martins and Galileo, 1991 (new country record), and Phoebe mexicana Bates, 1881 (new state record).
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We propose several nomenclatural changes for taxa in the lampyrid subfamilies Ototretinae, Photurinae, and Psilocladinae (Coleoptera: Lampyridae). Evidence for the correct year of description of Drilaster albicornis lateobscura (Pic, 1921) is presented. Spellings are corrected for Drilaster debilis holzi (Pic, 1914), Drilaster impustulata fukienensis (Pic, 1955), Drilaster moutoni (Pic, 1911), Drilaster pendleburyi (Pic, 1943) and Pyrogaster lunifera (Eschscholtz, 1822). We also explain the validity of the name Photuris flavicollis Fall, 1927. Fifty-eight taxa described as variations or aberrations in the subfamilies Ototretinae, Photurinae, and Psilocladinae by Delkeskamp (1977), McDermott (1966), Wittmer (1944), and Pic (1924c), are evaluated, with their availability determined based on ICZN (1999: Article 45.6).
ZooBank registration. urn:lsid:zoobank.org:pub:65697128-997F-4A52-AC9E-6860B0BF997D
Autarcontes lopezi Fisher, 1925 (Coleoptera: Buprestidae), is transferred to the genus Agrilus Curtis, 1825 (new combination). Colobogaster bella Kirsch, 1873, is transferred to the genus Chrysobothris Eschscholtz, 1829 (new combination). Ectinogonia isamarae Moore, 1994, is resurrected as the valid name for the species previously called E. obscuripennis Cobos, 1954, as the latter is unavailable as infrasubspecific. Conognatha jakobsoni Obenberger, 1928, is resurrected over C. germaini Théry in Hoscheck, 1934, as the former name has priority. Callimicra lucida Waterhouse, 1889, is resurrected as the valid name over C. hoscheki Obenberger, 1922, which has been used due to several historical errors and misinterpretations.
ZooBank registration. urn:lsid:zoobank.org:pub:651D001D-1C91-4A1A-B8BE-335BC7E7DD0F
Several nomenclatural changes for taxa in the firefly subfamily Luciolinae (Coleoptera: Lampyridae) are proposed. Evidence is presented to correct the year of description for Luciola dregei Motschulsky 1853 to Motschulsky 1854 when the name was ratified. We correct the authorities and years of descriptions for type species designations for the genera Delopleurus Motschulsky, 1853, and Delopyrus Motschulsky, 1853 to Motschulsky 1854 for both. All remaining taxa described as variations in the subfamily Luciolinae by McDermott (1966), are evaluated to subspecies with their availability determined based on ICZN (1999) Article 45.6.
ZooBank registration. urn:lsid:zoobank.org:pub:618A5442-2644-4E9A-BE70-07C413810DB9
Lectotypes are designated for Acmaeodera amabilis Horn, 1878 and Acmaeodera disjuncta Fall, 1899 (Coleoptera: Buprestidae). The defining characters of Acmaeodera opacula LeConte, 1858 are compared with those of A. disjuncta Fall, 1899, as well as habitat and host plant. Acmaeodera disjuncta is synonymized with A. opacula. The distinguishing characters of congeners whose general appearance at times can resemble A. opacula are discussed, and a new state record for Mexico is provided for A. opacula.
ZooBank registration. urn:lsid:zoobank.org:pub:AA3C1BA7-9BF8-45F7-84D6-362B52305931
A new species of Acmaeodera Eschscholtz, 1829 (Coleoptera: Buprestidae) from southern Nevada, USA
(2023)
A new species of Acmaeodera Eschscholtz, 1829, A. raschkoi Westcott (Coleoptera: Buprestidae), is described from southern Nevada, USA, and compared to four other species in the genus. Its habitat and means of capture are discussed in detail.
ZooBank registration. urn:lsid:zoobank.org:pub:6EA78E6C-AD58-470B-9E9B-2815EA0B9340
The Lasioglossum (Dialictus) gemmatum species complex, also known as the L. tegulare species group and the L. parvum species complex, is a very common, widespread, diverse, and recognisable lineage of sweat bees, containing 22 previously described species and several known undescribed species. The species were recently revised for the eastern Nearctic region and the Greater Antilles, but remain poorly known in the western Nearctic along with most other L. (Dialictus). These characteristics make it a prime candidate for revision in ongoing taxonomic work on the western Nearctic L. (Dialictus). Here we present the results of this revision, including 10 new species descriptions, one new synonymy, a preliminary phylogeny, and keys to known Nearctic species. Species of the eastern Nearctic and a few primarily Neotropical species which can occur in the Nearctic are also included. We report that the L. (D.) gemmatum species complex is likely a monophyletic group arising from the L. (D.) comulum group, but that the enlarged tegula has arisen independently in at least two other L. (Dialictus) lineages, and it contains multiple cases of allopatric speciation. The following species are described as new: Lasioglossum (Dialictus) angelicum sp. nov., L. (D.) deludens sp. nov., L. (D.) diabolicum sp. nov., L. (D.) eremum sp. nov., L. (D.) gloriosum sp. nov., L. (D.) indagator sp. nov., L. (D.) holzenthali sp. nov., L. (D.) magnitegula sp. nov., L. (D.) profundum sp. nov., and L. (D.) rufodeludens sp. nov. Previously undescribed males of L. (D.) perparvum (Ellis, 1914) and L. (D.) pseudotegulare (Cockerell, 1896) and the female of L. (D.) gaudiale (Sandhouse, 1924) are diagnosed and figured for the first time. Lasioglossum (Dialictus) hunteri (Crawford, 1932) is a new subjective junior synonym of L. (D.) ellisiae (Sandhouse, 1924). Pre-2022 specimen records of L. (D.) hunteri and L. (D.) tegulariforme (Crawford, 1907) are attributable to a heterogeneous mix of species, and records of L. (D.) perparvum are likely attributable to L. (D.) deludens.
Three novel species collected from Dinghushan Biosphere Reserve (DHSBR) in southern China, Russula cylindrica Y.Song sp. nov. and R. lacteocarpa Y.Song sp. nov. in subgenus Archaeae and R. reticulofolia Y.Song sp. nov. in subg. Compactae, are described based on morphological and molecular data. In addition, Russula leucobrunnea Y.Song nom. nov. is proposed in replacement of R. leucocarpa nom. illeg. in subg. Brevipedum, as R. leucocarpa (T.Lebel) T.Lebel had been described earlier. Differences between the three novel species and their closely related taxa were analyzed. Another two known species in subg. Brevipedum, R. callainomarginis J.F.Liang & J.Song and R. japonica Hongo were also identified among specimens from DHSBR and are described and illustrated. Phylogenetic analyses of ITS and a five-locus phylogeny (concatenated LSU, mtSSU, rpb1, rpb2 and tef1) support the recognition of these taxa.
Based on morphological and molecular evidence, two new species of Silene are recognized and described here, S. penduliflora F.Jafari, Keshavarzi & Doostm. sp. nov. and S. thyrsiantha F.Jafari, Mirtadz. & Keshavarzi sp. nov. The newly discovered species are distributed in the central and southeastern parts of Iran, growing in rocky habitats. Relationships among these species and their close relatives are demonstrated using nrDNA ITS and cpDNA rps16 phylogenies. Silene ghahremaninejadii, S. parrowiana, and S. shahrudensis form a clade with these new species. A key to S. penduliflora and S. thyrsiantha and their close relatives is provided.
Enicospilus Stephens, 1835 is the largest genus of subfamily Ophioninae (Ichneumonidae) with more than 700 extant species worldwide that are mostly nocturnal and parasitoids of larvae of Lepidoptera. In this paper, the Vietnamese species of Enicospilus are reviewed for the first time. Of the total 82 recorded species, 10 species are described as new: E. aequiscleritalis sp. nov., E. bulbipennis sp. nov., E. centraliscleritiger sp. nov., E. circuliscleritalis sp. nov., E. gialaiensis sp. nov., E. hiepi sp. nov., E. melanothoracicus sp. nov., E. nigristernalis sp. nov., E. trui sp. nov., and E. tuani sp. nov. Fifty-two species are recorded for the first time from the country: E. abdominalis (Szépligeti, 1906), E. acutus Shimizu, 2020, E. argus Gauld & Mitchell, 1981, E. atoponeus Cushman, 1947, E. bacillaris Wang, 1997, E. bakerielli Gauld & Mitchell, 1981, E. bifasciatus (Uchida, 1928), E. biharensis Townes, Townes & Gupta, 1961, E. concentralis Cushman, 1937, E. corculus (Tosquinet, 1903), E. dasychirae Cameron, 1905, E. eastopi Gauld & Mitchell, 1981, E. enicospilus Nikam, 1972, E. exaggeratus Chiu, 1954, E. fittoni Nikam, 1980, E. flavocephalus (Kirby, 1900), E. formosensis (Uchida, 1928), E. fusiformis Chiu, 1954, E. gasteralis Nikam, 1980, E. grandis (Cameron, 1905), E. hamatus Gauld & Mitchell, 1981, E. hedilis Gauld & Mitchell, 1981, E. iapetus Gauld & Mitchell, 1981, E. insinuator (Smith, 1860), E. ixion Gauld & Mitchell, 1981, E. javanus (Szépligeti, 1910), E. laqueatus (Enderlein, 1921), E. longitarsis Tang, 1990, E. mythrus Gauld & Mitchell, 1981, E. nathani Gauld & Mitchell, 1981, E. nigribasalis (Uchida, 1928), E. nigristigma Cushman, 1937, E. nigriventris Nikam, 1975, E. nigronotatus Cameron, 1903, E. nigropectus Cameron, 1905, E. pallidistigma Cushman, 1937, E. pantanae Tang, 1990, E. pinguivena (Enderlein, 1921), E. pseudoconspersae (Sonan, 1927), E. purifenestratus (Enderlein, 1921), E. rhetus Gauld & Mitchell, 1981, E. riukiuensis (Matsumura & Uchida, 1926), E. sauteri (Enderlein, 1921), E. selmatos Chiu, 1954, E. strigilatus Tang, 1990, E. teleus Gauld & Mitchell, 1981, E. transversus Chiu, 1954, E. tripartitus Chiu, 1954, E. urus Gauld & Mitchell, 1981, E. verticinus (Roman, 1913), E. yonezawanus (Uchida, 1928), and E. zebrus Gauld & Mitchell, 1981. A key to all Vietnamese species of Enicospilus is provided.
The Cyrtodactylus irregularis group, originally considered to consist of only one taxon, has been split into 26 species. We herein present the distribution of all species within the group in Cambodia, Laos and Vietnam and describe two new species based on integrative analyses. Cyrtodactylus chumuensis sp. nov. is discovered from Dak Lak Province and distinguished from the remaining taxa by more than 11.86% genetic divergence and by the following distinct morphological characters: size medium (SVL 67.5 mm); enlarged femoral scales on each thigh 4–5, femoral pores 0–2 in males; precloacal pores 6–7 in males; ventral scale rows 43–45; lamellae under toe IV 17–21. Cyrtodactylus arndti sp. nov. is described from Binh Dinh Province and genetically differentiated from its congeners by a minimum of 11.42% and by the following characters: adult size medium (SVL 73.4–80.8 mm); enlarged femoral scales on each thigh 5–11; femoral pores 0–2 in males; 6 precloacal pores in males, females with 6 pitted precloacal pores; ventral scale rows 26–38; lamellae under toe IV 17–22; subcaudal scales transversely enlarged. Additionally, we highlight the potential cryptic diversity with the taxon currently regarded as C. pseudoquadrivirgatus and understudied areas in Vietnam where new species will likely be discovered.
Three new species of the bamboo-feeding genus Bambusiphaga Huang & Ding, 1979, B. parvula sp. nov., B. angulosa sp. nov., and B. nigrigena sp. nov., are described and illustrated from China. A key to species of the genus is provided. Habitus photos for adults and illustrations of male genitalia are also given.
The species of Aphaniosoma Becker, 1903 described from Egypt by Collin in 1949 are reviewed and lectotypes designated for Aphaniosoma creperum Collin, 1949, A. fissum Collin, 1949, A. lamellatum Collin, 1949 and A. sodalis Collin, 1949. All species known from the eastern Mediterranean and the Middle East are reviewed, with annotations, distributions and a key to species. The following 19 species are described as new and illustrated: A. acitergum sp. nov., A. aegyptium sp. nov., A. angulitergum sp. nov., A. captiosum sp. nov., A. cristatum sp. nov., A. curvistylus sp. nov., A. daedalum sp. nov., A. denticulosum sp. nov., A. flavipyga sp. nov., A. freidbergi sp. nov., A. interispina sp. nov., A. kravchenkoi sp. nov., A. longilingua sp. nov., A. lucidum sp. nov., A. meltseri sp. nov., A. miricercus sp. nov., A. palestinense sp. nov., A. praeditum sp. nov. and A. yaromi sp. nov. Biology and ecology are commented upon.
Four new species and one new genus of Lamiinae (Coleoptera: Cerambycidae) are described: Nyssodrysilla humeralis new species (Acanthocinini), from Paraguay; Atrypanius schmidi new species (Acanthocinini), from Paraguay; Chicanatonus hovorei, new genus, new species (Acanthocinini), from Mexico; Euryestola transversa new species (Calliini), from Ecuador. The following new combinations are established: Leiopus convexus Melzer, 1934 is transferred to Hyperplatys Haldeman, 1847; Leiopus floccidus Erichson, 1847 is transferred to Anisopodus White, 1855; Leiopus marcelamonneae Audureau and Demez, 2015 and L. pleuriticus White, 1855 are transferred to Atrypanius Bates, 1864. Leiopus histrionicus Gistel, 1848 is newly synonymized with Eutrypanus dorsalis (Germar, 1823). The formal transference of Leiopus soricinus Fairmaire and Germain, 1859 to Lepturges (Lepturges) Bates, 1863, forgotten in recent catalogs and checklists, is reinforced. Atrypanius marcelamonneae new combination is excluded from the Paraguayan fauna. Euryestola cribrata (Bates, 1881) is newly recorded from Panama. Keys to species of Nyssodrysilla Gilmour, 1962 and Euryestola Breuning, 1940 are provided. The occurrence of Colobothea naevia Bates, 1865 in Ecuador is confirmed. Colobothea olivencia Bates, 1865 is newly recorded from Ecuador and from the Brazilian states of Pará and Ceará; variation in the pubescent pattern on the pronotum of this species is reported. Four new species and one new genus of Lamiinae (Coleoptera: Cerambycidae) are described: Nyssodrysilla humeralis new species (Acanthocinini), from Paraguay; Atrypanius schmidi new species (Acanthocinini), from Paraguay; Chicanatonus hovorei, new genus, new species (Acanthocinini), from Mexico; Euryestola transversa new species (Calliini), from Ecuador. The following new combinations are established: Leiopus convexus Melzer, 1934 is transferred to Hyperplatys Haldeman, 1847; Leiopus floccidus Erichson, 1847 is transferred to Anisopodus White, 1855; Leiopus marcelamonneae Audureau and Demez, 2015 and L. pleuriticus White, 1855 are transferred to Atrypanius Bates, 1864. Leiopus histrionicus Gistel, 1848 is newly synonymized with Eutrypanus dorsalis (Germar, 1823). The formal transference of Leiopus soricinus Fairmaire and Germain, 1859 to Lepturges (Lepturges) Bates, 1863, forgotten in recent catalogs and checklists, is reinforced. Atrypanius marcelamonneae new combination is excluded from the Paraguayan fauna. Euryestola cribrata (Bates, 1881) is newly recorded from Panama. Keys to species of Nyssodrysilla Gilmour, 1962 and Euryestola Breuning, 1940 are provided. The occurrence of Colobothea naevia Bates, 1865 in Ecuador is confirmed. Colobothea olivencia Bates, 1865 is newly recorded from Ecuador and from the Brazilian states of Pará and Ceará; variation in the pubescent pattern on the pronotum of this species is reported.
ZooBank registration. urn:lsid:zoobank.org:pub:7C223856-CC09-4A47-8A52-E4F98C445241
Two new genera, Skelleyus Opitz and Divulgoatus Opitz (Coleoptera: Cleridae: Enopliinae), show unusual antennal development. Funicular antennomeres become shorter as they approach an extensive capitulum. This study involves three taxa, Skelleyus leavengoodi Opitz, new species, Divulgoatus kelleri Opitz, new species, and Divulgoatus discrepans (Gorham).
ZooBank registration. urn:lsid:zoobank.org:pub:E059A916-DDE2-4193-A090-8CDD46EDC859
Taxonomic history and review of the Förster genera of Platygastridae (Hymenoptera: Platygastroidea)
(2023)
Platygastridae is a ʻdark taxonʼ, with many genera and species in dire need of professional attention. The taxonomic impediment is especially severe in the Palearctic Platygastrinae due to the abundance of names with vague concepts. Historical descriptions and their associated type material must be examined and clarified before further revisionary work can occur. Arnold Förster described 18 genera of Platygastridae, most of which represent distinct and recognizable lineages. The present study reviews their taxonomic history, providing diagnostic remarks, English translations, and illustrations of important specimens from the Förster collection in the Natural History Museum Vienna. The collection also includes original exemplar specimens of European species whose types have been lost. Neotypes and lectotypes are designated from this material to improve nomenclatural stability in the group. Neotypes are designated for Amblyaspis forticornis (Nees, 1834), Isocybus grandis (Nees, 1834), Platygaster striolata Nees, 1834, and Trichacis tristis (Nees, 1834). Lectotypes are designated for Leptacis spinigera (Nees, 1834) comb. nov. and for Platygaster corvina Förster, 1861, with Platygaster henkvlugi Buhl, 1996 treated as a junior synonym. Platygaster mutica Nees, 1834 stat. rev., nomen dubium, is transferred from Synopeas.
The nitidus species group of the bee genus Scrapter Lepeletier & Serville, 1828 is redefi ned, revised and 15 species are described as new for science: S.caeruleus sp. nov. ♀, S.confusus sp. nov. ♀♂, S.convexoides sp. nov. ♂, S. convexus sp. nov. ♀♂, S. crassipunctatus sp. nov. ♀♂, S.felicis sp. nov. ♀♂, S.fl avipunctatus sp. nov. ♀♂, S.imparilis sp. nov. ♀♂, S. littoralis sp. nov. ♀, S.longicornis sp. nov. ♂, S.montanus sp. nov. ♀♂, S.mpumalangensis sp. nov. ♀♂, S.obtusus sp. nov. ♀♂, S. perpunctatulus sp. nov. ♂ and S.variabilis sp. nov. ♀♂. The previously unknown males of S. divergens (Friese, 1925), S. semirufus Cockerell, 1932 and S. perpunctatus Cockerell, 1933 are described for the fi rst time. All currently known 28 species of the S. nitidus species group are redescribed, imaged and included in a key to facilitate their identifi cation.
Detection of morphologically indistinguishable cryptic species implies using an integrative taxonomic approach with a combination of molecular, contemporary morphological, ecological and other relevant analyses. Within a contemporary morphological analysis, two approaches are commonly used in hoverfly taxonomy: a geometric morphometric analysis of the wing and surstyle shape. Here, the importance of the R4+5 vein shape is tested in cryptic species delimitation within four Merodon species groups using linear and semilandmark geometric morphometric analyses. As expected, geometric morphometrics showed a stronger resolution compared to linear morphometrics. Linear morphometrics failed to detect differences related to sexual dimorphism or differences among the species M. pruni and M. obscurus. However, all cryptic species and sexes were separated with high significance based on the R4+5 vein shape. Moreover, obtained results concurred with the landmark-defined wing shape and molecular results published in previous studies. Additionally, combining two characters, the semilandmark R4+5 vein shape and the landmark-defined wing shape, provided more detailed and precise insights into the shape differences. Our results showed that the R4+5 vein shape stands out as an important character in species delimitation of hoverflies where the sinuation of this vein is present. Therefore, it can be beneficial as a single character or in combination with a landmark-based wing shape analysis.
The Australian genus Pleuroseta Richards, 1973 is revised, including P. wentworthi (Richards, 1973) and three new species: P. ingens sp. nov. (Queensland), P. monteithi sp. nov. (Queensland) and P. occidentalis sp. nov. (Western Australia). Leptocera (Biroina) dorrigonis Richards, 1973 is treated as a new junior synonym of Pleuroseta wentworthi.
The present paper is devoted to the description of three new species of the genus Acanthophorella Antić & Makarov, 2016 from Georgia, western Caucasus: A. devi Antić sp. nov. and A. valerii Antić sp. nov., both presumed troglobionts and each from a single cave, and the epigean A. aurita Antić sp. nov. The troglobiotic Acanthophorella barjadzei Antić & Makarov, 2016 is reported from two additional caves, with further descriptive notes given. Notes on the ecology and localities, and a distribution map for all six species of the genus are presented. A key is given to all members of the Flagellophorella complex. The relationships within the complex and the distribution and troglomorphism in the genus Acanthophorella are briefly discussed.
A new fossil ceratopogonid genus and species from Lower Cretaceous Lebanese amber, Baskintoconops maaloufi Pielowska-Ceranowska gen. et sp. nov., is described and illustrated. The studied material originates from a newly discovered amber site in the Lebanese village Baskinta at a locality dubbed Qanat Bakish. The described genus is typified by its wing venation pattern combining characters of genera Fossileptoconops and Jordanoconops belonging to the subfamily Leptoconopinae.
A review of the Ptocasius Simon, 1885 spiders of Gaoligong Mountains, China (Araneae: Salticidae)
(2023)
Sixteen new species of the genus Ptocasius are described from Mt. Gaoligong, Yunnan: P. angulatus sp. nov. (♀); P. circulus sp. nov. (♀); P. danzhu sp. nov. (♂♀); P. davidi sp. nov. (♀); P. filiformus sp. nov. (♂♀); P. foliolatus sp. nov. (♀); P. geminus sp. nov. (♂♀); P. jietouensis sp. nov. (♂♀); P. longapophysis sp. nov. (♂♀); P. longlingensis sp. nov. (♂♀); P. rectangulus sp. nov. (♀); P. robustus sp. nov. (♀); P. tengchongensis sp. nov. (♂♀); P. umbellulatus sp. nov. (♀); P. zabkai sp. nov. (♂♀) and P. zonatus sp. nov. (♀). Including P. montanus (Żabka, 1981) and P. pseudoflexus (Liu, Yang & Peng, 2016), a total of eighteen species of Ptocasius have been reported from Mt. Gaoligong. For each new species, a morphological description, photos of the body and copulatory organs, line drawings of copulatory organs, and locality maps are provided.
Taxonomic revision of the African assassin bug genus Fusius (Heteroptera: Reduviidae: Peiratinae)
(2023)
The African assassin bug genus Fusius Stål, 1862 is revised after the examination of type specimens with redescriptions of four species. Lectotypes of Pirates (Fusius) H-flavum Reuter, 1881 and Pirates rubricosus Stål, 1855 are designated. The status of P. (Fusius) H-flavum Reuter, 1881 is revalidated with its current name as F. hflavus (Reuter, 1881) stat. rev. et comb. nov. Seven new synonyms are proposed: F. dilutus Miller, 1957 = F. gowdeyi Miller, 1957 syn. nov. = F. liberiensis Miller, 1957 syn. nov. = F. dilutus anonymus Dispons, 1969 syn. nov. = F. dilutus vicinus Dispons, 1969 syn. nov.; F. distinctus Miller, 1957 = F. sylvestris Miller, 1957 syn. nov.; F. hflavus (Reuter, 1881) = F. hargreavesi Miller, 1957 syn. nov.; F. rubricosus (Stål, 1855) = F. ugandensis Miller, 1957 syn. nov. A key is provided to separate the four species of this genus. Diagnosis and distribution of Fusius are briefly discussed.
Four new species of Alloxysta (Hym.: Cynipoidea: Figitidae: Charipinae) from Petr Starý’s collection
(2023)
After the revision of Petr Starý’s Charipinae collection, four new species of Alloxysta Förster, 1869 have been found. This collection represents a good overview of the Charipinae fauna worldwide, although the best represented area is Central Europe. Here, we describe four new species of Alloxysta: A. llumae Ferrer-Suay sp. nov., A. onae Ferrer-Suay sp. nov., A. poli Ferrer-Suay sp. nov. and A. staryi Ferrer-Suay & Pujade-Villar sp. nov. Three of these new species are based on reared specimens. Diagnosis, distribution, and description of the new species are given and illustrated.
A review of the genus Stratiomys from India is presented. The new species Stratiomys brunettii sp. nov. is described based on male and female specimens collected from the Kashmir Himalayas. The only other congener previously recorded in India, Stratiomys approximata, is redescribed. A key to the species is presented.
Serratichneumon Riedel & Sheng gen. nov. and Serratichneumon maculatus Sheng & Riedel gen. et sp. nov. belonging to the tribe Ichneumonini of subfamily Ichneumoninae (Hymenoptera: Ichneumonidae), collected in the Oriental Region (China, Vietnam, and Indonesia), are described and illustrated. The new genus is placed in Tereshkin’s key to the Palaearctic genera of the subtribe Amblytelina, and compared with similar genera, Hepiopelmus Wesmael, 1845, and Tricholabus Thomson, 1894.
The Andean genus Priscula Simon, 1893 includes the largest Neotropical pholcid spiders, but due to their mostly cryptic lifestyle they remain poorly collected and poorly studied. Many species available in collections remain undescribed and nothing has been published about the phylogeny and the biology of the genus. Here, we deal with a recent collection of Priscula spiders from Ecuador, the country of origin of the type species, P. gularis Simon, 1893. We describe eight new species, collected at 17 localities at altitudes from 640–3160 m, all based on males and females: P. azuay sp. nov., P. llaviucu sp. nov., P. espejoi sp. nov., P. esmeraldas sp. nov., P. chapintza sp. nov., P. pastaza sp. nov., P. bonita sp. nov., and P. lumbaqui sp. nov. We use a sample of approximately 26 species-level taxa, mostly from Ecuador and Venezuela, to propose a first hypothesis about relationships within the genus. Our data (mainly CO1) suggest the existence of five species groups, three of which are represented in Ecuador. The cave-dwelling P. pastaza sp. nov. is only slightly troglomorphic (paler than usual; anterior median eyes strongly reduced or lost) but differs dramatically from forest-dwelling congeners in its biology: it hangs fully exposed in its web during the day; it produces egg sacs with only 6–7 eggs (average in 15 other species: 42 eggs); and it produces the largest eggs relative to body size of all studied species.
A faunal study for the order Phasmatodea of the Chicaque Natural Park is presented, including a list of species found, descriptions, redescriptions, and biological notes. A total of nine species were found and studied; two new genera: Ramandeun new genus, Nubilophasma new genus, and four new species: Atratomorpha jorgei new species, Isagoras franciscoverai new species, Nubilophasma chicaquensis new genus and new species, and Ramandeum coronatum new genus and new species are described. The description of the eggs of the new taxa, of the previously unknown eggs of Paraceroys quadrispinosus (Redtenbacher, 1906), and the redescription of the eggs of Libethra rabdota Stål, 1875, and Libethra inchoata Brunner von Wattenwyl, 1907 are provided. Additional comments on the ecology and morphological variation of the reviewed taxa are included. Finally, further studies on the stick insect fauna of the Colombian Andes are discussed and recommended to provide more information to broaden the understanding of the species that inhabit this complex mountain system.
ZooBank registration. urn:lsid:zoobank.org:pub:7D34FF9A-CDEE-4DD4-A643-E0F467E00A5B
The Neotropical eucnemid genus, Dyscolotaxia Horn, is revised following the examination of specimens from five collections, examination of the published original description of Plesiofornax tetratoma Chassain, and from images provided by Fernanda Salazar-Buenaño (QCAZI). Dyscolotaxia championi Horn is redescribed with included descriptions of the male genitalia and a female specimen. Three new species of false click beetles (Coleoptera: Eucnemidae) are described from the Neotropical region. These new species are: Dyscolotaxia chiriquiensis (Panama), Dyscolotaxia hispaniolensis (Dominican Republic) and Dyscolotaxia picea (Ecuador). Amazotaxia new genus is described from South America for Plesiofornax tetratoma Chassain, creating Amazotaxia tetratoma (Chassain) new combination. Morphology of both Dyscolotaxia and Amazotaxia in relation to Pleisofornax Cocquerel are reviewed along with biogeographical hypotheses for these groups. An identification key is provided for species of Dyscolotaxia in the Neotropical region. Images for all species of Dyscolotaxia and Amazotaxia are provided.
Plagiosarus transversus Vlasak and Santos-Silva, new species (Cerambycidae: Lamiinae: Acanthoderini), is described from Costa Rica. The rank of Plagiosarus melampus congestus Bates, 1885 is discussed.
ZooBank registration. urn:lsid:zoobank.org:pub:71496BE9-C69F-43B4-AF33-E3376DA6E083
A century and a half since the time of Hewitson, we are experiencing a renaissance in species discovery fueled by whole genome sequencing. A large-scale genomic analysis of Hesperiidae Latreille, 1809 (Lepidoptera), including primary type specimens, reveals a deluge of species new to science. One hundred of them (one in a new genus) are described here from the New World (type localities are given in parenthesis): Drephalys (Drephalys) diovalis Grishin, new species (Ecuador: Napo), Euriphellus panador Grishin, new species (Ecuador: Esmeraldas), Euriphellus panamicus Grishin, new species (Panama: Panama), Cecropterus (Thorybes) viridissimus Grishin, new species (Ecuador: Zamora-Chinchipe), Cecropterus (Murgaria) dariensis Grishin, new species (Panama: Darien), Urbanus (Urbanus) mericuti Grishin, new species (Ecuador: Napo), Telegonus (Telegonus) pastus Grishin, new species (Panama: Panama), Autochton (Autochton) dora Grishin, new species (Ecuador: Pastaza), Astraptes centralis Grishin, new species (Panama: Colón), Aguna claxonica Grishin, new species (Ecuador: Napo), Aguna esmeralda Grishin, new species (Ecuador: Esmeraldas), Aguna lata Grishin, new species (Guyana), Ridens angulinea Grishin, new species (Peru: Cuzco), Pythonides lera Grishin, new species (Peru: Cuzco), Pythonides latemarginatus Grishin, new species (Panama: Panama), Gindanes variegatus Grishin, new species (Brazil: Mato Grosso), Milanion (Milanion) virga Grishin, new species (Brazil: Rondônia), Milanion (Milanion) furvus Grishin, new species (Panama: Panama), Milanion (Milanion) laricus Grishin, new species (Ecuador: Napo), Charidia ronda Grishin, new species (Brazil: Rondônia), Pseudodrephalys tinas Grishin, new species (Peru: Loreto), Pseudodrephalys argus Grishin, new species (Suriname: Para), Achlyodes calvus Grishin, new species (Brazil: Santa Catarina), Spioniades artemis Grishin, new species (Panama: Panama), Spioniades artemidoides Grishin, new species (Brazil: Santa Catarina), Myrinia orieca Grishin, new species (Ecuador: Orellana), Myrinia aragua Grishin, new species (Venezuela: Aragua), Myrinia maculosa Grishin, new species (Guatemala), Myrinia manchada Grishin, new species (Guyana), Polyctor (Fenops) lamperus Grishin, new species (Panama: Darien), Nisoniades (Nisoniades) lutum Grishin, new species (Mexico: Guerrero. ), Bolla (Stolla) vena Grishin, new species (Venezuela: Aragua), Staphylus (Vulga) vula Grishin, new species (Mexico: Veracruz), Staphylus (Vulga) vulga Grishin, new species (Panama: Darien), Staphylus (Staphylus) rotundalus Grishin, new species (Ecuador: Napo), Staphylus (Staphylus) yucatanus Grishin, new species (Mexico: Quintana Roo/Yucatan), Heliopetes (Heliopetes) lana Grishin, new species (Guatemala), Canesia ella Grishin, new species (Venezuela: Barinas), Paches (Paches) loxeca Grishin, new species (Ecuador: Morona-Santiago), Clito congruens Grishin, new species (Panama: Colón), Cycloglypha corax Grishin, new species (Brazil: Rio de Janeiro), Festivia peruvia Grishin, new species (Peru: Huánuco), Decinea notata Grishin, new species (Ecuador: Napo), Pompeius fuscus Grishin, new species (Brazil: Minas Gerais), Vernia clara Grishin, new species (Panama: Chiriquí), Oligoria (Oligoria) obtena Grishin, new species (Ecuador: Napo), Thespieus mandal Grishin, new species (Brazil: Rio de Janeiro), Psoralis (Saniba) magnamacus Grishin, new species (Panama: Darien), Alychna ayonis Grishin, new species (Ecuador: Napo), Wahydra banios Grishin, new species (Ecuador: Tungurahua), Wahydra cuzcona Grishin, new species (Peru: Cuzco), Cynea (Cynea) aureofimbra Grishin, new species (Ecuador), Cynea (Nycea) quada Grishin, new species (Ecuador: Napo), Cynea (Quinta) achirae Grishin, new species (Mexico: Tamaulipas), Eutus amazonicus Grishin, new species (Peru: Madre de Dios), Eutus incus Grishin, new species (Peru: Cuzco), Eutus septemaculatus Grishin, new species (Brazil: Mato Grosso), Godmia viridicapita Grishin, new species (Ecuador: Napo), Rhomba pulla Grishin, new species (Peru: Cuzco), Niconiades victoria Grishin, new species (Mexico: Tamaulipas), Lancephallus purpurus Grishin, new genus and new species (Guyana), Mnasicles (Remella) ecua Grishin, new species (Ecuador: Pichincha), Amblyscirtes (Amblyscirtes) aeratus Grishin, new species (Mexico: Oaxaca), Amblyscirtes (Mastor) chrysoplea Grishin, new species (Mexico: Oaxaca), Amblyscirtes (Mastor) chrysomisa Grishin, new species (Mexico: Chiapas), Amblyscirtes (Flor) meridus Grishin, new species (Mexico: Veracruz), Rectava chiriquensis Grishin, new species (Panama: Chiriquí), Cobalopsis adictys Grishin, new species (Panama: Veraguas), Cymaenes melaporphyrus Grishin, new species (Mexico: San Luis Potosí), Lerema (Morys) ecuadorica Grishin, new species (Ecuador: Pichincha), Saturnus obscurior Grishin, new species (Panama: Darien), Cantha zoirodicta Grishin, new species (Peru: Madre de Dios), Cantha meiodicta Grishin, new species (Peru: Madre de Dios), Phlebodes duplex Grishin, new species (Guatemala: Cayuga), Lychnuchus (Enosis) valle Grishin, new species (Colombia: Valle), Eutychide ochoides Grishin, new species (Peru: Cuzco), Dion bora Grishin, new species (Panama: Darien), Dion occida Grishin, new species (Peru: Madre de Dios), Eprius (Eprius) veledinus Grishin, new species (Ecuador: Pichincha), Radiatus panamensis Grishin, new species (Panama: Panama), Pheraeus pulcher Grishin, new species (Peru: Madre de Dios), Callimormus rades Grishin, new species (Panama: Panama), Gubrus lubens Grishin, new species (Ecuador: Loja), Ludens labens Grishin, new species (Panama: Darien), Rigga isa Grishin, new species (Ecuador: Napo), Flaccilla lactea Grishin, new species (Peru: Cuzco), Falga athena Grishin, new species (Panama: Darien), Panoquina jay Grishin, new species (Peru: Loreto), Calpodes salianus Grishin, new species (Peru: Madre de Dios), Calpodes stingo Grishin, new species (Ecuador: Sucumbíos), Aides nobra Grishin, new species (Panama: Colón), Thracides pavo Grishin, new species (Mexico: Tabasco), Talides eluta Grishin, new species (Peru: Cuzco), Talides laeta Grishin, new species (Peru: Cuzco), Neoxeniades angustior Grishin, new species (Brazil: Rio de Janeiro), Damas zea Grishin, new species (Guyana), Tromba xantha Grishin, new species (Mexico: Veracruz), Perichares fura Grishin, new species (Ecuador: Pichincha), Carystoides (Balma) goliath Grishin, new species (Colombia: Valle), and Agathymus galeana Grishin, new species (Mexico: Nuevo Leon). Additionally, we present evidence to support 22 taxa as species (not subspecies or synonyms) and synonymize one genus and four species. Namely, the following taxa are species: Milanion pilta Evans, 1953 (not Milanion pilumnus Mabille and Boullet, 1917), Milanion latior Mabille and Boullet, 1917 (not a synonym of Milanion marciana Godman and Salvin, 1895), Charidia pilea Evans, 1953, and Charidia pocus Evans, 1953 (not Charidia lucaria (Hewitson, 1868)), Paches (Paches) gloriosus Röber, 1925 and Paches (Paches) loxana Evans, 1953 (not Paches (Paches) loxus (Westwood, 1852)), Spioniades anta Evans, 1953 (not Spioniades abbreviata (Mabille, 1888)), Decinea onasima (Hewitson, 1877) and Decinea formosus (Hayward, 1940) (not Decinea dama (Herrich-Schäffer, 1869)), Thespieus guerreronis (Dyar, 1913) (not Thespieus dalman (Latreille, [1824])), Cynea (Nycea) erebina (Möschler, 1879) and Cynea (Nycea) cleochares (Mabille, 1891) (not Cynea (Cynea) diluta (Herrich-Schäffer, 1869)), Amblyscirtes (Mastor) repta Evans, 1955 (not Amblyscirtes (Flor) florus (Godman, 1900)), Saturnus tiberius (Möschler, 1883), Saturnus conspicuus (E. Bell, 1941), Saturnus meton (Mabille, 1891), and Saturnus obscurus (E. Bell, 1941) (not Saturnus reticulata (Plötz, 1883)), Phlebodes sifax Evans, 1955 (not Phlebodes campo (E. Bell, 1947)), Eutychide ochus Godman, 1900 and Eutychide rogersi (Kaye, 1914) (not a subspecies and a synonym, respectively, of Eutychide subcordata (Herrich-Schäffer, 1869)), Falga mirabilis Evans, 1955, Falga jacta Evans, 1955, and Falga ombra Evans, 1955 (not Falga jeconia (A. Butler, 1870)); and the following taxa are junior subjective synonyms: Libra Evans, 1955 (of Phemiades Hübner, [1819]), Papilio clito Fabricius, 1787 of Milanion hemes hemes (Cramer, 1777), Pamphila hycsos Mabille, 1891 of Cynea (Nycea) erebina (Möschler, 1879), Hesperia olympia Plötz, 1882 of Eutychide subcordata (Herrich-Schäffer, 1869), and Hesperia ocrinus Plötz, 1882 of Aides aegita (Hewitson, 1866). Furthermore, we propose new combinations for genus-species: Lychnuchus (Enosis) ponka (Evans, 1955) (not Thoon Godman, 1900), and species-subspecies: Charidia pocus mayo Evans, 1953 (not Charidia lucaria (Hewitson, 1868)), Decinea onasima boliviensis (E. Bell, 1930) (not Decinea dama (Herrich-Schäffer, 1869)), Cynea (Nycea) erebina somba Evans, 1955 (not Pamphila hycsos Mabille, 1891), Saturnus tiberius suffuscus (Hayward, 1940) (not Saturnus reticulata (Plötz, 1883)), and Falga mirabilis odol Evans, 1955 (not Falga jeconia (A. Butler, 1870)). Then, Milanion pilumnus var. hemestinus Mabille and Boullet, 1917 is a junior subjective synonym of Milanion pilumnus pilumnus Mabille and Boullet, 1917, not of Milanion leucaspis (Mabille, 1878). Lectotypes are designated for nine taxa (names in original combinations below): Pellicia bromias Godman and Salvin, 1894 (Mexico: Veracruz, Atoyac), Nisoniades perforata Möschler, 1879 (Colombia), Helias ascalaphus Staudinger, 1876 (central Panama), Pamphila hycsos Mabille, 1891 (Colombia), Amblyscirtes fluonia Godman, 1900 (Mexico: Guerrero, Xocomanatlan), Mastor anubis Godman, 1900 (Mexico: Guerrero, Omiltemi), Eutychide ochus Godman, 1900 (Mexico: Veracruz, Atoyac), Cobalus subcordata Herrich-Schäffer, 1869 (Southeast Brazil), and Thracides xanthura Godman, 1901 (Panama: Chiriquí Province, Bugaba). A neotype is designated for Eudamus briccius Plötz, 1881 (Guyana: Iwokrama Forest).
ZooBank registration. urn:lsid:zoobank.org:pub:ACDF923B-906D-460E-9707-259E0ECDBCA8
Genomic analysis of Pyrginae Burmeister, 1878 (Lepidoptera: Hesperiidae Latreille, 1809) with an emphasis on the tribes Achlyodini Burmeister, 1878 and Carcharodini Verity, 1940 reveals many inconsistencies between the resulting phylogeny and the current classification. These problems are corrected by proposing new taxa, changing the ranks of others, or synonymizing them, and transferring species between genera. As a result, five subtribes, one genus, 20 subgenera, and one species are proposed as new: Cyclosemiina Grishin, new subtribe (type genus Cyclosemia Mabille, 1878), Ilianina Grishin, new subtribe (type genus Iliana E. Bell, 1937), Nisoniadina Grishin, new subtribe (type genus Nisoniades Hübner, [1819]), Burcina Grishin, new subtribe (type genus Burca E. Bell and W. Comstock, 1948), and Pholisorina Grishin, new subtribe (type genus Pholisora Scudder, 1872), all in Carcharodini; Lirra Grishin, new genus (type species Leucochitonea limaea Hewitson, 1868) in Pythonidina Grishin, 2019; Trifa Grishin, new subgenus (type species Tagiades jacobus Plötz, 1884), Tuberna Grishin, new subgenus (type species Pythonides contubernalis Mabille, 1883), Ebona Grishin, new subgenus (type species Quadrus eboneus E. Bell, 1947), Noctis Grishin, new subgenus (type species Achlyodes accedens Mabille, 1895), and Cyrna Grishin, new subgenus (type species Achlyodes cyrna Mabille, 1895) of Quadrus Lindsey, 1925; Liddia Grishin, new subgenus (type species Helias pallida R. Felder, 1869), Minna Grishin, new subgenus (type species Achlyodes minna Evans, 1953), and Thilla Grishin, new subgenus (type species Eurypterus later Mabille, 1891) of Eantis Boisduval, 1836; Torgus Grishin, new subgenus (type species Ouleus gorgus E. Bell, 1937) of Iliana E. Bell, 1937; Fenops Grishin, new subgenus (type species Cabares enops Godman and Salvin, 1894) of Polyctor Evans, 1953; Bezus Grishin, new subgenus (type species Pellicia bessus Möschler, 1877) and Macarius Grishin, new subgenus (type species Pellicia macarius Herrich-Schäffer, 1870) of Nisoniades Hübner, [1819]; Quadralis Grishin, new subgenus (type species Pterygospidea extensa Mabille, 1891) of Gorgopas Godman and Salvin, 1894; Menuda Grishin, new subgenus (type species Nisoniades menuda Weeks, 1902) and Narycus Grishin, new subgenus (type species Pythonides narycus Mabille, 1889) of Perus Grishin, 2019; Bovaria Grishin, new subgenus (type species Achlyodes cyclops Mabille, 1876), Sebia Grishin, new subgenus (type species Nisoniades eusebius Plötz, 1884), and Stolla Grishin, new subgenus (type species Pholisora balsa E. Bell, 1937) of Bolla Mabille, 1903; Vulga Grishin, new subgenus (type species Achlyodes vulgata Möschler, 1879) and Capilla Grishin, new subgenus (type species Helias aurocapilla Staudinger, 1876, currently a junior subjective synonym of Hesperia musculus Burmeister, 1875) of Staphylus Godman and Salvin, 1896; and Quadrus (Zera) vivax Grishin, new species (type locality in Brazil: Rio de Janeiro). The following 10 are subgenera, not genera or synonyms: Ouleus Lindsey, 1925 and Zera Evans, 1953 of Quadrus Lindsey, 1925; Atarnes Godman and Salvin, 1897 and Eburuncus Grishin, 2012 of Milanion Godman and Salvin, 1895; Pachyneuria Mabille, 1888 and Austinus O. Mielke and Casagrande, 2016 of Sophista Plötz, 1879; Hemipteris Mabille, 1889 and Mictris Evans, 1955 of Pellicia Herrich-Schäffer, 1870; and Hesperopsis Dyar, 1905 and Scantilla Godman and Salvin, 1896 of Staphylus Godman and Salvin, 1896. The following 7 are species, not subspecies: Quadrus (Ebona) cristatus (Steinhauser, 1989) (not Quadrus (Ebona) negrus (Nicolay, 1980)), Quadrus (Quadrus) ophia (A. Butler, 1870) (not Quadrus (Quadrus) lugubris (R. Felder, 1869)), Quadrus (Zera) gellius (Mabille, 1903) and Quadrus (Zera) servius (Plötz, 1884) (not Quadrus (Zera) hyacinthinus (Mabille, 1877)), Mimia pazana Evans,1953 (not Mimia phidyle (Godman and Salvin, 1894)), Polyctor (Polyctor) dagua Evans, 1953 (not Polyctor (Polyctor) polyctor (Prittwitz, 1868)), and Staphylus (Vulga) satrap Evans, 1953 (not Staphylus (Vulga) saxos Evans, 1953); and these 8 are species, not synonyms: Quadrus (Zera) menedemus (Godman and Salvin, 1894) (not Quadrus (Zera) tetrastigma (Sepp, [1847])), Pellicia (Pellicia) bilinea Mabille, 1889 (not Pellicia (Pellicia) dimidiata Herrich-Schäffer, 1870), Pellicia (Hemipteris) nema Williams and Bell, 1939 (not Pellicia (Pellicia) theon Plötz, 1882), Bolla (Bovaria) sodalis Schaus, 1913 (not Bolla (Bolla) imbras (Godman and Salvin, 1896)), Bolla (Bovaria) aplica (E. Bell, 1937) (not Bolla (Sebia) eusebius (Plötz, 1884)), Bolla (Sebia) chilpancingo (E. Bell, 1937) (not Bolla (Bolla) subapicatus (Schaus, 1902)), and Bolla (Stolla) madrea (R. Williams and E. Bell, 1940) and Bolla (Stolla) hazelae (Hayward, 1940) (not Bolla (Stolla) zorilla (Plötz, 1886)). The following 2 are junior subjective synonyms: Achlyodes erisichthon Plötz, 1884 of Quadrus (Zera) servius (Plötz, 1884) (not a subspecies of Quadrus (Zera) tetrastigma (Sepp, [1847]) and Staphylus subapicatus Schaus, 1902 of Bolla (Bolla) imbras (Godman and Salvin, 1896). Furthermore, we propose the following additional new genus-species combination: Gindanes homer (Evans, 1953), Gindanes nides (O. Mielke and Casagrande, 2002), Gindanes maraca (O. Mielke and Casagrande, 1992), Gindanes jenmorrisae (Shuey and Ramírez. 2022), Gindanes tullia (Evans, 1953), Gindanes herennius (Geyer, [1838]), Gindanes proxenus (Godman and Salvin, 1895), Gindanes parallelus (Mabille, 1898), Gindanes braga (Evans, 1953), Gindanes hampa (Evans, 1953), Gindanes rosa (Steinhauser, 1989), Gindanes neivai (Hayward, 1940), Gindanes mundo (H. Freeman, 1979), Gindanes eminus (E. Bell, 1934), Quadrus (Trifa) francesius Freeman, 1969, Quadrus (Trifa) ineptus (Draudt, 1922), Quadrus (Trifa) jacobus (Plötz, 1884), Quadrus (Tuberna) lancea (Hewitson, 1868), Quadrus (Ebona) pescada (E. Bell, 1956), Lirra pteras (Godman and Salvin, 1895), and Lirra limaea (Hewitson, 1868) (not Pythonides Hübner, 1819); Quadrus (Cyrna) zora (Evans, 1953) (not Bolla Mabille, 1903); Eantis later (Mabille, 1891) and Eantis haber (Mabille, 1891) (not Aethilla Hewitson, 1868); Iliana (Torgus) gorgus (E. Bell, 1937) and Iliana (Torgus) taurus (Evans, 1953) (not Eantis Boisduval, 1836); Bolla (Stolla) evemerus (Godman and Salvin, 1896), Bolla (Stolla) chlora (Evans, 1953), Bolla (Stolla) astra (R. Williams and E. Bell, 1940), Bolla (Stolla) balsa (E. Bell, 1937), Bolla (Stolla) tridentis (Steinhauser, 1989), Bolla (Stolla) esmeraldus (L. Miller, 1966), Bolla (Stolla) chlorocephala (Latreille, [1824]), and Bolla (Stolla) incanus (E. Bell, 1932) (not Staphylus Godman and Salvin, 1896). Finally, lectotypes are designated for Achlyodes servius Plötz, 1884 (type locality in Brazil: Rio de Janeiro), Pellicia theon Plötz, 1882 (type locality in South America), and Nisoniades eusebius Plötz, 1884 (type locality in Central America). Unless stated otherwise, all subgenera, species, subspecies, and synonyms of mentioned genera and species are transferred with their parent taxa, and others remain as previously classified.
ZooBank registration. http://zoobank.org/B9AFA1A9-8664-4F31-B4D9-ACF7780C7CC6
Acmaeodera natlovei new species (Coleoptera: Buprestidae) is described from the southwestern United States. Details of phenology, geographic range, larval, flower and adult host plants, and similar species are discussed. Acmaeodera yuccavora Knull, 1962 is newly synonymized with Acmaeodera conoidea Fall, 1899. Acmaeodera thoracata Knull, 1974 and A. bryanti Van Dyke, 1953 are newly synonymized with Acmaeodera neoneglecta Fisher, 1949. New state and host records are reported for United States. A key to the 46 species of Acmaeodera occurring east of the Rocky Mountain states is provided.
ZooBank registration. urn:lsid:zoobank.org:pub:E03CF02E-D212-4AF1-8E3B-5AE3463D2A71
Aksakidion odontokeras, new genus and new species (Coleoptera: Chrysomelidae: Eumolpinae: Eumolpini), is described from four specimens in the Bowditch Collection at Harvard University. It is distinguished from all other genera in the Eumolpinae by an elongate, seta-bearing, tusk-like projection on the dorsal side of each mandible. Although locality data only mention Paraguay as the country of origin, information about the collector indicates it was collected in the late 19th century in the vicinity of Asunción.
ZooBank registration. urn:lsid:zoobank.org:pub:65DABC23-2BF6-42D3-B445-AC2B9EDA66E8