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Acrostilicus Hubbard, 1896 and Pachystilicus Casey, 1905 are North American genera known from only one and two species, respectively, and have never been a subject of a modern revision. In fact, Acrostilicus was not even properly described as its author provided only a sketchy diagnosis of the genus and species. Here, we provide a redescription of the genus Acrostilicus and its species and illustrate the habitus and male genital features. For the first time, we also redescribe Pachystilicus and its two species, and provide their differential diagnoses. Additionally, we tested the phylogenetic position of both genera. They were scored into a morphological matrix supplemented with molecular data and the analyses were run using Bayesian inference and maximum likelihood methods. A total of 119 morphological characters and 4859 bp of nuclear (28S, TP, Wg, CADA, CADC, ArgK) and mitochondrial (COI) sequences were analysed for 46 taxa. The results confirmed that both Acrostilicus and Pachystilicus are members of the subtribe Stilicina, but at the same time challenged the monophyly of the subtribe in its current composition. Additionally, we provided further evidence for non-monophyly of the subtribe Medonina and discussed the biology of Acrostilicus and Pachystilicus.
New species of genus Atractides Koch, 1837 (Acari: Hydrachnidiae, Hygrobatidae) from Qinghai, China
(2022)
The paper deals with five new species of the genus Atractides Koch, 1837 (Acari: Hydrachnidiae, Hygrobatidae) collected from Qinghai Province, P.R. China, Atractides (Atractides) biprojectus Zhang, Li & Guo sp. nov., Atractides (Atractides) smiti Zhang, Li & Guo sp. nov., Atractides (Atractides) menyuanensis Zhang, Li & Guo sp. nov., Atractides (Atractides) longiprojectus Zhang, Li & Guo sp. nov. and Atractides (Atractides) xianmiensis Zhang, Li & Guo sp. nov. All the new species are described and illustrated in detail, and all the type specimens are deposited in the Institute of Entomology, Guizhou University, Guiyang, China (GUGC).
Our expanded efforts in genomic sequencing to cover additional skipper butterfly (Lepidoptera: Hesperiidae) species and populations, including primary type specimens, call for taxonomic changes to restore monophyly and correct misidentifications by moving taxa between genera and proposing new names. Reconciliation between phenotypic characters and genomic trees suggests three new tribes, two new subtribes, 23 new genera, 17 new subgenera and 10 new species that are proposed here: Psolosini Grishin, new tribe (type genus Psolos Staudinger, 1889), Ismini Grishin, new tribe (type genus Isma Distant, 1886), Eetionini Grishin, new tribe (type genus Eetion de Nicéville, 1895), Orphina Grishin, new subtribe (type genus Orphe Godman, 1901), Carystoidina Grishin, new subtribe (type genus Carystoides Godman, 1901), Fulvatis Grishin, new genus (type species Telegonus fulvius Plötz, 1882), Adina Grishin, new genus (type species Nascus adrastor Mabille and Boullet, 1912), Ornilius Grishin, new genus (type species Ornilius rotundus Grishin, new species), Tolius Grishin, new genus (type species Antigonus tolimus Plötz, 1884), Lennia Grishin, new genus (type species Leona lena Evans, 1937), Trida Grishin, new genus (type species Cyclopides barberae Trimen, 1873), Noxys Grishin, new genus (type species Oxynthes viricuculla Hayward, 1951), Gracilata Grishin, new genus (type species Enosis quadrinotata Mabille, 1889), Hermio Grishin, new genus (type species Falga ? hermione Schaus, 1913), Eutus Grishin, new genus (type species Cobalus rastaca Schaus, 1902), Gufa Grishin, new genus (type species Phlebodes gulala Schaus, 1902), Godmia Grishin, new genus (type species Euroto chlorocephala Godman, 1900), Rhomba Grishin, new genus (type species Eutychide gertschi Bell, 1937), Rectava Grishin, new genus (type species Megistias ignarus Bell, 1932), Contrastia Grishin, new genus (type species Hesperia distigma Plötz, 1882), Mit Grishin, new genus (type species Mnasitheus badius Bell, 1930), Picova Grishin, new genus (type species Vorates steinbachi Bell, 1930), Lattus Grishin, new genus (type species Eutocus arabupuana Bell, 1932), Gubrus Grishin, new genus (type species Vehilius lugubris Lindsey, 1925), Koria Grishin, new genus (type species Hesperia kora Hewitson, 1877), Corta Grishin, new genus (type species Eutychide lycortas Godman, 1900), Calvetta Grishin, new genus (type species Hesperia calvina Hewitson, 1866), Oz Grishin, new genus (type species Astictopterus ozias Hewitson, 1878), Praxa Grishin, new subgenus (type species Nascus prax Evans, 1952), Bron Grishin, new subgenus (type species Papilio broteas Cramer, 1780), Turis Grishin, new subgenus (type species Pyrgus 1955, and Synale Mabille, 1904 of Carystus Hübner, [1819]. The following 20 genera are treated as junior subjective synonyms: Leucochitonea Wallengren, 1857 of Abantis Hopffer, 1855; Sapaea Plötz, 1879 and Netrobalane Mabille, 1903 of Caprona Wallengren, 1857; Parasovia Devyatkin, 1996 of Sebastonyma Watson, 1893; Pemara Eliot, 1978 of Oerane Elwes and Edwards, 1897; Ankola Evans, 1937 of Pardaleodes Butler, 1870; Arotis Mabille, 1904 of Mnaseas Godman, 1901; Chalcone Evans, 1955, Hansa Evans, 1955, and Propertius Evans, 1955 of Metrocles Godman, 1900; Jongiana O. Mielke and Casagrande, 2002 of Cobaloides Hayward, 1939; Pamba Evans, 1955 of Psoralis Mabille, 1904; Brownus Grishin, 2019 of Styriodes Schaus, 1913; Mnasilus Godman, 1900 of Papias Godman, 1900; Sucova Evans, 1955 of Mnasitheus Godman, 1900; Pyrrhocalles Mabille, 1904 and Asbolis Mabille, 1904 of Choranthus Scudder, 1872; Miltomiges Mabille, 1903 of Methionopsis Godman, 1901; Sacrator Evans, 1955 of Thracides Hübner, [1819]; and Lychnuchoides Godman, 1901 of Perichares Scudder, 1872. Arunena Swinhoe, 1919 is a junior subjective synonym of Stimula de Nicéville, 1898 (not of Koruthaialos Watson, 1893). The following 27 names are species-level taxa (some in new combinations) reinstated from synonymy: Salantoia gildo (Mabille, 1888) (not Salatis cebrenus (Cramer, 1777)), Bungalotis corentinus (Plötz, 1882) (not Bungalotis midas (Cramer, 1775)), Telegonus cretellus (Herrich-Schäffer, 1869) (not Telegonus cassander (Fabricius, 1793)), Santa palica (Mabille, 1888) (not Chiothion asychis (Stoll, 1780)), Camptopleura cincta Mabille and Boullet, 1917 (not Camptopleura auxo (Möschler, 1879)), Camptopleura orsus (Mabille, 1889) (not Nisoniades mimas (Cramer, 1775)), Metron voranus (Mabille, 1891) and Metron fasciata (Möschler, 1877) (not Metron zimra (Hewitson, 1877)), Limochores catahorma (Dyar, 1916) (not Limochores pupillus (Plötz, 1882)), Pares viridiceps (Mabille, 1889) (not Thoon modius (Mabille, 1889)), Tigasis wellingi (Freeman, 1969) (not Tigasis arita (Schaus, 1902)), Rectava sobrinus (Schaus, 1902) (not Papias phainis Godman, 1900), Nastra subsordida (Mabille, 1891) (not Adlerodea asema (Mabille, 1891), previously in Eutychide Godman, 1900), Lerema pattenii Scudder, 1872 (not Lerema accius (J. E. Smith, 1797)), Lerema (Morys) ancus (Möschler, 1879) (not Cymaenes tripunctus theogenis (Capronnier, 1874)), Cobalopsis zetus (Bell, 1942) (not Cobalopsis nero (Herrich-Schäffer, 1869)), Lerema (Geia) etelka (Schaus, 1902) (not Lerema (Geia) geisa (Möschler, 1879), previously in Morys Godman, 1900), Cymaenes isus (Godman, 1900) (not Cymaenes trebius (Mabille, 1891)), Vehilius labdacus (Godman, 1900) (not Vehilius inca (Scudder, 1872)), Papias amyrna (Mabille, 1891) (not Papias allubita (Butler, 1877), previously in Mnasilus Godman, 1900), Papias integra (Mabille, 1891) (not Papias subcostulata (Herrich-Schäffer, 1870)), Metiscus atheas Godman, 1900 (not Hesperia achelous Plötz, 1882), Dion agassus (Mabille, 1891) (not Dion uza (Hewitson, 1877), previously in Enosis Mabille, 1889), Picova incompta (Hayward, 1942) (not Lerema (Morys) micythus (Godman, 1900), previously in Morys Godman, 1900), Lucida melitaea (Draudt, 1923) (not Lucida lucia (Capronnier, 1874)), Methionopsis modestus Godman, 1901 (not Methionopsis ina (Plötz, 1882)), and Thargella (Volus) volasus (Godman, 1901) (not Eutocus facilis (Plötz, 1884)). The following 57 taxa are elevated from subspecies to species, new status (some in new combinations): Dyscophellus doriscus (Hewitson, 1867) (not Dyscophellus porcius (C. Felder and R. Felder, 1862), Phocides vida (A. Butler, 1872) (not Phocides urania (Westwood, 1852)), Tagiades (Daimio) ceylonica Evans, 1932 (not Tagiades litigiosa Möschler, 1878), Tagiades (Daimio) tubulus Fruhstorfer, 1910 (not Tagiades sambavana Elwes and Edwards, 1897), Tagiades (Daimio) kina Evans, 1934, Tagiades (Daimio) sheba Evans, 1934, Tagiades (Daimio) martinus Plötz, 1884, Tagiades (Daimio) sem Mabille, 1883, and Tagiades (Daimio) neira Plötz, 1885 (not Tagiades trebellius (Hopffer, 1874)), Tagiades (Daimio) korela Mabille, 1891 and Tagiades (Daimio) presbyter Butler, 1882 (not Tagiades nestus (C. Felder, 1860)), Tagiades obscurus Mabille, 1876, Tagiades ravi (Moore, [1866]), Tagiades atticus (Fabricius, 1793), Tagiades titus Plötz, 1884, Tagiades janetta Butler, 1870, Tagiades inconspicua Rothschild, 1915, and Tagiades hovia Swinhoe, 1904 (not Tagiades japetus (Stoll, [1781])), Tagiades silvia Evans, 1934 and Tagiades elegans Mabille, 1877 (not Tagiades gana (Moore, [1866])), Tapena bornea Evans, 1941 and Tapena minuscula Elwes and Edwards, 1897 (not Tapena thwaitesi Moore, [1881]), Darpa dealbata (Distant, 1886) (not Darpa pteria (Hewitson, 1868)), Perus manx (Evans, 1953) (not Perus minor (Schaus, 1902)), Canesia pallida (Röber, 1925) (not Carrhenes canescens (R. Felder, 1869)), Carrhenes conia Evans, 1953 (not Carrhenes fuscescens (Mabille, 1891)), Anisochoria extincta Hayward, 1933 and Anisochoria polysticta Mabille, 1876 (not Anisochoria pedaliodina (Butler, 1870)), Anisochoria verda Evans, 1953 (not Anisochoria minorella Mabille, 1898), Bralus alco (Evans, 1953) (not Bralus albida (Mabille, 1888)), Ephyriades jamaicensis (Möschler, 1879) (not Ephyriades brunnea (Herrich-Schäffer, 1865)), Koruthaialos (Stimula) frena Evans, 1949 (not Koruthaialos focula (Plötz, 1882)), Euphyes kiowah (Reakirt, 1866) (not Euphyes vestris (Boisduval, 1852)), Mnaseas inca Bell, 1930 (not Mnaseas bicolor (Mabille, 1889)), Metron hypochlora (Draudt, 1923) (not Metrocles schrottkyi (Giacomelli, 1911), previously in Metron Godman, 1900), Decinea huasteca (H. Freeman, 1969), Decinea denta Evans, 1955, and Decinea antus (Mabille, 1895) (not Decinea decinea (Hewitson, 1876)), Xeniades pteras Godman, 1900 (not Xeniades chalestra (Hewitson, 1866)), Xeniades difficilis Draudt, 1923 (not Xeniades orchamus (Cramer, 1777)), Xeniades hermoda (Hewitson, 1870) (not Tisias quadrata (HerrichSchäffer, 1869)), Hermio vina (Evans, 1955) (not Hermio hermione (Schaus, 1913), previously in Lento Evans, 1955), Cymaenes loxa Evans, 1955, (not Cymaenes laureolus (Schaus, 1913)), Niconiades peri (Evans, 1955) (not Rhinthon bajula (Schaus, 1902), previously in Neoxeniades Hayward, 1938), Gallio danius (Bell, 1941) (not Vehilius seriatus (Mabille, 1891)), Gallio massarus (E. Bell, 1940) (not Gallio garima (Schaus, 1902) previously in Tigasis Godman, 1900), Cymaenes edata (Plötz, 1882), Cymaenes miqua (Dyar, 1913) and Cymaenes aequatoria (Hayward, 1940) (not Cymaenes odilia (Burmeister, 1878)), Lychnuchus (Enosis) demon (Evans, 1955) (not Lychnuchus (Enosis) immaculata (Hewitson, 1868), previously in Enosis Mabille, 1889), Naevolus naevus Evans, 1955 (not Naevolus orius (Mabille, 1883)), Lucida scopas (Mabille, 1891), Lucida oebasus (Godman, 1900), and Lucida leopardus (Weeks, 1901) (not Lucida lucia (Capronnier, 1874)), Corticea schwarzi (E. Bell, 1941) and Corticea sylva (Hayward, 1942) (not Corticea mendica (Mabille, 1898)), and Choranthus orientis (Skinner, 1920) (not Choranthus antiqua (Herrich-Schäffer, 1863), previously in Pyrrhocalles Mabille, 1904). Borbo impar bipunctata (Elwes and J. Edwards, 1897) is a valid subspecies, not a synonym of Borbo impar tetragraphus (Mabille, 1891), here placed in synonymy with Lotongus calathus (Hewitson, 1876), new synonym. We confirm the species status of Telegonus cassius (Evans, 1952) and Lerema (Morys) valda Evans, 1955. Euphyes chamuli Freeman, 1969 is placed as a subspecies of Euphyes kiowah (Reakirt, 1866), new status. The following 41 taxa are junior subjective synonyms, either newly proposed or transferred from synonymy with other species or subspecies: Telegonus mutius Plötz, 1882 of Euriphellus phraxanor (Hewitson, 1876), Telegonus erythras Mabille, 1888 of Dyscophellus damias (Plötz, 1882), Aethilla jaira Butler, 1870 of Telegonus cretellus (Herrich-Schäffer, 1869), Paches era Evans, 1953 of Santa palica (Mabille, 1888), Antigonus alburnea Plötz, 1884 of Tolius tolimus robigus (Plötz, 1884) (not of Echelatus sempiternus simplicior (Möschler, 1877)), Echelatus depenicillus Strand, 1921 of E. sempiternus simplicior (not of T. tolimus robigus), Antigonus aura Plötz, 1884 of Theagenes dichrous (Mabille, 1878) (not of Helias phalaenoides palpalis (Latreille, [1824])), Achlyodes impressus Mabille, 1889 of Camptopleura orsus (Mabille, 1889), Augiades tania Schaus, 1902 of Metron voranus (Mabille, 1891), Pamphila verdanta Weeks, 1906 of Metron fasciata (Möschler, 1877), Niconiades viridis vista Evans, 1955 of Niconiades derisor (Mabille, 1891), Pamphila binaria Mabille, 1891 of Conga chydaea (A. Butler, 1877) (not of Cynea cynea (Hewitson, 1876)), Psoralis concolor Nicolay, 1980 of Ralis immaculatus (Hayward, 1940), Hesperia dido Plötz, 1882 of Cynea (Quinta) cannae (Herrich-Schäffer, 1869) (not of Lerema lochius (Plötz, 1882)), Proteides osembo Möschler, 1883 of Cynea (Cynea) diluta (Herrich-Schäffer, 1869) (not of Cynea (Quinta) cannae (Herrich-Schäffer, 1869)), Cobalopsis brema E. Bell, 1959 of Eutus rastaca (Schaus, 1902), Psoralis panamensis Anderson and Nakamura, 2019 of Rhomba gertschi (Bell, 1937), Cobalus asella Herrich-Schäffer, 1869 of Amblyscirtes alternata (Grote and Robinson, 1867) (not of Amblyscirtes vialis (W. H. Edwards, 1862)), Papias trimacula Nicolay, 1973 of Nastra subsordida (Mabille, 1891), Pamphila bipunctata Mabille, 1889 and Sarega staurus Mabille, 1904 of Lerema pattenii Scudder, 1872 (not of Cymaenes lumina (Herrich-Schäffer, 1869), previously in Lerema Scudder, 1872), Hesperia aethra Plötz, 1886 of Lerema lineosa (Herrich-Schäffer, 1865) (not of Lerema (Morys) compta Butler, 1877), Megistias miaba Schaus, 1902 of Cobalopsis valerius (Möschler, 1879), Phanis sylvia Kaye, 1914 of Lerema etelka (Schaus, 1902) (not of Lerema (Geia) geisa (Möschler, 1879), previously in Morys Godman, 1900), Carystus odilia Burmeister, 1878, Pamphila trebius Mabille, 1891 and Megistias corescene Schaus, 1902 of Cymaenes lumina (Herrich-Schäffer, 1869), Hesperia phocylides Plötz, 1882 of Cymaenes edata (Plötz, 1882) (not of Lerema accius (J. E. Smith, 1797)), Pamphila xenos Mabille, 1898 of Vehilius inca (Scudder, 1872), Mnasilus guianae Lindsey, 1925 of Papias amyrna (Mabille, 1891), Pamphila nubila Mabille, 1891 of Papias integra (Mabille, 1891) (not of Cynea corisana (Plötz, 1882)), Enosis matheri H. Freeman, 1969 of Metiscus atheas Godman, 1900 (previously in Enosis Mabille, 1889), Hesperia infuscata Plötz, 1882 of Mnaseas derasa derasa (Herrich-Schäffer, 1870) (previously Arotis Mabille, 1904), (not of Papias subcostulata (Herrich-Schäffer, 1870)), Pamphila astur Mabille, 1891 of Metiscus angularis (Möschler, 1877) (not of Cymaenes tripunctus theogenis (Capronnier, 1874)), Anthoptus macalpinei H. Freeman, 1969 of Anthoptus inculta (Dyar, 1918), Methionopsis typhon Godman, 1901 of Methionopsis ina (Plötz, 1882), Methionopsis dolor Evans, 1955 of Thargella volasus (Godman, 1901), Hesperia cinica Plötz, 1882 of Dubiella dubius (Stoll, 1781), Cobalus disjuncta Herrich-Schäffer, 1869 of Dubiella dubius (Stoll, 1781) (not of Vettius lafrenaye (Latreille, [1824])), and Saliana vixen Evans, 1955 of Neoxeniades parna (Evans, 1955). The following are new and revised genusspecies combinations: Euriphellus cebrenus (Cramer, 1777) (not Salatis Evans, 1952), Gorgopas extensa (Mabille, 1891) (not Polyctor Evans, 1953), Clytius shola (Evans, 1953) (not Staphylus Godman and Salvin, 1896), Perus narycus (Mabille, 1889) (not Ouleus Lindsey, 1925), Perus parvus (Steinhauser and Austin, 1993) (not Staphylus Godman and Salvin, 1896), Pholisora litus (Dyar, 1912) (not Bolla Mabille, 1903), Carrhenes decens (A. Butler, 1874) (not Antigonus Hübner, [1819]), Santa palica (Mabille, 1888) (not Chiothion Grishin, 2019), Bralus nadia (Nicolay, 1980) (not Anisochoria Mabille, 1876), Acerbas sarala (de Nicéville, 1889) (not Lotongus Distant, 1886), Caenides sophia (Evans, 1937) (not Hypoleucis Mabille, 1891), Hypoleucis dacena (Hewitson, 1876) (not Caenides Holland, 1896), Dotta tura (Evans, 1951) (not Astictopterus C. Felder and R. Felder, 1860), Nervia wallengrenii (Trimen, 1883) (not Kedestes Watson, 1893), Testia mammaea (Hewitson, 1876) (not Decinea Evans, 1955), Oxynthes trinka (Evans, 1955) (not Orthos Evans, 1955), Metrocles argentea (Weeks, 1901) (not Paratrytone Godman, 1900), Metrocles scitula (Hayward, 1951) (not Mucia Godman, 1900), Metrocles schrottkyi (Giacomelli, 1911) (not Metron Godman, 1900), Niconiades derisor (Mabille, 1891) (not Decinea Evans, 1955), Paratrytone samenta (Dyar, 1914) (not Ochlodes Scudder, 1872), Oligoria (Cobaloides) locutia (Hewitson, 1876) (not Quinta Evans, 1955), Psoralis (Saniba) laska (Evans, 1955) (not Vidius Evans, 1955), Psoralis (Saniba) arva (Evans, 1955) and Psoralis (Saniba) umbrata (Erschoff, 1876) (not Vettius Godman, 1901), Psoralis (Saniba) calcarea (Schaus, 1902) and Psoralis (Saniba) visendus (E. Bell, 1942) (not Molo Godman, 1900), Alychna gota (Evans, 1955) (not Psoralis Mabille, 1904), Adlerodea asema (Mabille, 1891) and Adlerodea subpunctata (Hayward, 1940) (not Eutychide Godman, 1900), Ralis immaculatus (Hayward, 1940) (not Mucia Godman, 1900), Rhinthon braesia (Hewitson, 1867) and Rhinthon bajula (Schaus, 1902) (not Neoxeniades Hayward, 1938), Cymaenes lochius Plötz, 1882 (not Lerema Scudder, 1872), Paracarystus ranka (Evans, 1955) (not Thoon Godman, 1900), Tricrista aethus (Hayward, 1951), Tricrista canta (Evans, 1955), Tricrista slopa (Evans, 1955), Tricrista circellata (Plötz, 1882), and Tricrista taxes (Godman, 1900) (not Thoon Godman, 1900), Gallio madius (E. Bell, 1941) and Gallio seriatus (Mabille, 1891) (not Vehilius Godman, 1900), Gallio garima (Schaus, 1902) (not Tigasis Godman, 1900), Tigasis corope (HerrichSchäffer, 1869) (not Cynea Evans, 1955), Tigasis perloides (Plötz, 1882) (not Cymaenes Scudder, 1872), Amblyscirtes (Flor) florus (Godman, 1900) (not Repens Evans, 1955), Vidius fraus (Godman, 1900) (not Cymaenes Scudder, 1872), Nastra celeus (Mabille, 1891) (not Vehilius Godman, 1900), Nastra nappa (Evans, 1955) (not Vidius Evans, 1955), Vehilius warreni (Weeks, 1901) and Vehilius limae (Lindsey, 1925) (not Cymaenes Scudder, 1872), Cymaenes lumina (Herrich-Schäffer, 1869) (not Lerema Scudder, 1872), Cobalopsis valerius (Möschler, 1879) (not Cobalopsis Godman, 1900), Cobalopsis dictys (Godman, 1900) (not Papias Godman, 1900), Lerema (Morys) venias (Bell, 1942) (not Cobalopsis Godman, 1900), Papias latonia (Schaus, 1913) (not Cobalopsis Godman, 1900), Dion iccius (Evans, 1955) and Dion uza (Hewitson, 1877) (not Enosis Mabille, 1889), Vistigma (Vistigma) opus (Steinhauser, 2008) (not Thoon Godman, 1900), Saturnus fartuga (Schaus, 1902) (not Parphorus Godman, 1900), Phlebodes fuldai (E. Bell, 1930) (not Vettius Godman, 1901), Mnasitheus padus (Evans, 1955) (not Moeris Godman, 1900), Naevolus brunnescens (Hayward, 1939) (not Psoralis Mabille, 1904), Lamponia ploetzii (Capronnier, 1874) (not Vettius Godman, 1901), Mnestheus silvaticus Hayward, 1940 (not Ludens Evans, 1955), Rigga spangla (Evans, 1955) (not Sodalia Evans, 1955), Corticea vicinus (Plötz, 1884) (not Lento Evans, 1955), Mnasalcas thymoetes (Hayward, 1942) (not Mnasicles Godman, 1901), Mnasalcas boyaca (Nicolay, 1973) (not Pamba Evans, 1955), Vertica brasta (Evans, 1955) (not Lychnuchus Hübner, [1831]), Carystina discors Plötz, 1882 (not Cobalus Hübner, [1819]), Zetka irena (Evans, 1955) (not Neoxeniades Hayward, 1938), and Neoxeniades parna (Evans, 1955) (not Niconiades Hübner, [1821]). The following are new or revised species-subspecies combinations: Tagiades neira moti Evans, 1934, Tagiades neira canonicus Fruhstorfer, 1910, Tagiades sheba vella Evans, 1934, Tagiades sheba lola Evans, 1945, Tagiades korela biakana Evans, 1934, Tagiades korela mefora Evans, 1934, Tagiades korela suffusus Rothschild, 1915, Tagiades korela brunta Evans, 1949, Tagiades ravi ravina Fruhstorfer, 1910, Tagiades atticus carnica Evans, 1934, Tagiades atticus nankowra Evans, 1934, Tagiades atticus helferi C. Felder, 1862, Tagiades atticus balana Fruhstorfer, 1910, Tagiades inconspicua mathias Evans, 1934, Tagiades hovia kazana Evans, 1934, Tagiades elegans fuscata de Jong and Treadaway, 2007, Tagiades elegans semperi Fruhstorfer, 1910, Metron hypochlora tomba Evans, 1955, Decinea denta pruda Evans, 1955, and Choranthus orientis eleutherae (Bates, 1934) (previously in Pyrrhocalles Mabille, 1904). In addition to the abovementioned changes, the following new combinations involve newly proposed genus group names: Fulvatis fulvius (Plötz, 1882) and Fulvatis scyrus (E. Bell, 1934) (not Salatis Evans, 1952); Adina adrastor (Mabille and Boullet, 1912) (not Bungalotis Watson, 1893); Nascus (Praxa) prax Evans, 1952, Nascus (Bron) broteas (Cramer, 1780), and Nascus (Bron) solon (Plötz, 1882) (not Pseudonascus Austin, 2008); Chirgus (Turis) veturius (Plötz, 1884); Paches (Tiges) liborius (Plötz, 1884), and Paches (Tiges) mutilatus (Hopffer, 1874) (not Antigonus Hübner, [1819]); Paches (Tiges) exosa (A. Butler, 1877); Tolius tolimus (Plötz, 1884) and Tolius luctuosus (Godman & Salvin, 1894) (not Echelatus Godman and Salvin, 1894); Ancistroides (Ocrypta) caerulea (Evans, 1928), Ancistroides (Ocrypta) renardi (Oberthür, 1878), Ancistroides (Ocrypta) waigensis (Plötz, 1882), Ancistroides (Ocrypta) aluensis (Swinhoe, 1907), Ancistroides (Ocrypta) flavipes (Janson, 1886), and Ancistroides (Ocrypta) maria (Evans, 1949) (not Notocrypta de Nicéville, 1889); Lennia lena (Evans, 1937), Lennia binoevatus (Mabille, 1891), Lennia maracanda (Hewitson, 1876), and Lennia lota (Evans, 1937) (not Leona Evans, 1937); Trida barberae (Trimen, 1873) and Trida sarahae (Henning and Henning, 1998) (not Kedestes Watson, 1893); Noxys viricuculla (Hayward, 1951) (not Oxynthes Godman, 1900); Xeniades (Tixe) quadrata (Herrich-Schäffer, 1869), Xeniades (Tixe) rinda (Evans, 1955), Xeniades (Tixe) putumayo (Constantino and Salazar, 2013) (not Tisias Godman, 1901); Gracilata quadrinotata (Mabille, 1889) (not Styriodes Schaus, 1913); Hermio hermione (Schaus, 1913) (not Lento Evans, 1955); Cynea (Nycea) hycsos (Mabille, 1891), Cynea (Nycea) corisana (Plötz, 1882), Cynea (Nycea) popla Evans, 1955, Cynea (Nycea) iquita (E. Bell, 1941), Cynea (Nycea) robba Evans, 1955, Cynea (Nycea) melius (Geyer, 1832), and Cynea (Nycea) irma (Möschler, 1879); Eutus rastaca (Schaus, 1902) (not Eutychide Godman, 1900); Eutus yesta (Evans, 1955) (not Thoon Godman, 1900); Eutus mubevensis (E. Bell, 1932) (not Tigasis Godman, 1900); Gufa gulala (Schaus, 1902) (not Mucia Godman, 1900); Gufa fusca (Hayward, 1940) (not Tigasis Godman, 1900); Godmia chlorocephala (Godman, 1900) (not Onophas Godman, 1900); Rhomba gertschi (E. Bell, 1937) (not Justinia Evans, 1955); Mnasicles (Nausia) nausiphanes (Schaus, 1913) (not Tigasis Godman, 1900); Amblyscirtes (Flor) florus (Godman, 1900) (not Repens Evans, 1955); Rectava ignarus (E. Bell, 1932) (not Papias Godman, 1900); Rectava vorgia (Schaus, 1902) (not Cobalopsis Godman, 1900); Rectava nostra (Evans, 1955) (not not Vidius Evans, 1955); Lerema (Geia) geisa (Möschler, 1879) and Lerema (Geia) lyde (Godman, 1900) (not Morys Godman, 1900); Contrastia distigma (Plötz, 1882) (not Cymaenes Scudder, 1872); Mit (Mit) badius (E. Bell, 1930) (not Styriodes Schaus, 1913); Mit (Mit) gemignanii (Hayward, 1940), (not Mnasitheus Godman, 1900); Mit (Rotundia) schausi (Mielke and Casagrande, 2002), (not Enosis Mabille, 1889); Picova steinbachi (E. Bell, 1930) (not Saturnus Evans, 1955); Lattus arabupuana (E. Bell, 1932) (not Eutocus Godman, 1901); Gubrus lugubris (Lindsey, 1925) (not Vehilius Godman, 1900); Thargella (Pseudopapias) tristissimus (Schaus, 1902) (not Papias Godman, 1900); Koria kora (Hewitson, 1877) (not Justinia Evans, 1955); Justinia (Septia) septa Evans, 1955; Corta lycortas (Godman, 1900) (not Orthos Evans, 1955); Vertica (Brasta) brasta (Evans, 1955) (not Lychnuchus Hübner, [1831]); Calvetta calvina (Hewitson, 1866) (not Cobalus Hübner, [1819]); Neoxeniades (Bina) gabina (Godman, 1900) (not Orthos Evans, 1955); Oz ozias (Hewitson, 1878) and Oz sebastiani Salazar and Constantino, 2013 (not Lychnuchoides Godman, 1901); and Carystoides (Balma) balza Evans, 1955 and Carystoides (Balma) maroma (Möschler, 1877). Finally, unless stated otherwise, all subgenera, species, subspecies and synonyms of mentioned genera and species are transferred together with their parent taxa, and taxa not mentioned in this work remain as previously classified.
Japanese species of the genus Georissus are revised. Seven species, including two new, G. (Neogeorissus) takahashii sp. nov. and G. (Ne.) satoi sp. nov. are recognized. Georissus (Neogeorissus) sakaii Satô, 1972 and G. (Ne.) katsuoi Nakane, 1995 are synonymized with G. (Ne.) japonicus Satô, 1972. Based on the type series and additional specimens, three subgenera and seven species are (re)described with SEM micrographs of the external morphology and figures of male and female genitalia. The terminology of some structures of the head and the pronotum are redefined for convenience of taxonomy. A key to the Japanese species is also given.
Five new species of the genus Andes Stål, 1866 from China (Hemiptera, Fulgoromorpha, Cixiidae)
(2022)
Five new species of the genus Andes Stål, 1866, A. balteiformis Wang, Zhi & Chen sp. nov., A. bifidus Wang, Zhi & Chen sp. nov., A. furcutus Wang, Zhang & Chen sp. nov., A. latanalus Wang & Chen sp. nov. and A. pallidus Wang & Chen sp. nov. from China, are described and illustrated. A key to the species of Andes in China is provided.
Seven new species of the genus Meta C.L. Koch, 1836 from Southwest China are described here: M. bowo sp. nov. (♂♀), M. cona sp. nov. (♂♀), M. gyirong sp. nov. (♂), M. hongyuan sp. nov. (♂♀), M. tibet sp. nov. (♂♀), M. wanglang sp. nov. (♂♀) and M. weining sp. nov. (♂♀). Detailed descriptions, photos of somatic features and copulatory organs as well as line drawings, comparisons with closely related species, and a distribution map are provided.
An annotated list, including information on type species, distribution, and number of species, is provided for all of the non-flea-beetle galerucine genera known to occur in the New World (tribes Galerucini, Metacyclini, and Luperini). A diagnostic key to the genera is provided. Habitus illustrations are provided for most genera. The following new genera are proposed: Amplioluperus gen. nov., Cornuventer gen. nov., Geethaluperus gen. nov., Megarhabda gen. nov., Mexiluperus gen. nov., Monoaster gen. nov., Pyesexora gen. nov., Texiluperus gen. nov., Trachyelytron gen. nov. and Yingabruxia gen. nov. The following new taxonomic placements are proposed: Microbrotica Jacoby, 1887 is transferred from the tribe Metacyclini to the section Diabroticites Chapuis, 1875 (tribe Luperini, subtribe Diabroticina Chapuis, 1875); Pteleon Jacoby, 1888 is transferred from the section Exosomites Wilcox, 1973 (tribe Luperini, subtribe Luperina Gistel, 1848) to the section Scelidites Chapuis, 1875 (subtribe Luperina). The following new combinations are proposed: Luperodes histrio Horn, 1895, Luperus maculicollis LeConte, 1884, and Scelolyperus cyanellus Horn, 1895 are transferred from Pseudoluperus Beller & Hatch, 1932 to Amplioluperus; Luperodes tuberculatus Blake, 1942 is transferred from Pseudoluperus to Cornuventer; Luperus flavofemoratus Jacoby, 1888 is transferred from Pseudoluperus to Geethaluperus; Trirhabda obscurovittata Jacoby, 1886 is transferred from Trirhabda LeConte, 1865 to Megarhabda; Cneorane nigripes Allard, 1889 is transferred from Scelida Chapuis, 1875 to Metacycla Baly, 1861; Luperodes wickhami Horn, 1893 and Luperus dissimilis Jacoby, 1888 are transferred from Pseudoluperus to Mexiluperus; Scelolyperus tenuimarginatus Bowditch, 1925, is transferred from Scelida to Mimastra Baly, 1865 and is synonymized with Mimastra semimarginata Jacoby, 1886 syn. nov.; Pseudoluperus fulgidus Wilcox, 1965 and Pseudoluperus linus Wilcox, 1965 are transferred from Pseudoluperus to Monoaster; Crioceris detrita detrita Fabricius, 1801, Malacosoma detrita laevicollis Jacoby, 1887, Pyesia detrita meridionalis Bechyné, 1958, Pyesia elytropleuralis elytropleuralis Bechyné, 1958, and Pyesia elytropleuralis subalutacea Bechyné, 1958 are transferred from Pyesia Clark, 1865 to Pyesexora; Luperodes spretus Horn, 1893 and Luperodes texanus Horn, 1893 are transferred from Pseudoluperus to Texiluperus; Chthoneis smaragdipennis Jacoby, 1888 is transferred from Platymorpha Jacoby, 1888 to Trachyelytron; Luperus albomarginatus Jacoby, 1888 is transferred from Pseudoluperus to Trichobrotica Bechyné, 1956; and Galleruca sordida LeConte, 1858, Monoxia apicalis Blake, 1939, Monoxia batisia Blatchley, 1917, and Monoxia brisleyi Blake, 1939 are transferred from Monoxia LeConte, 1865 to Yingabruxia; all comb. nov. Pseudoluperus decipiens (Horn, 1893), originally described in Scelolyperus Crotch, 1874, is reduced to a junior synonym of Pseudoluperus longulus (LeConte, 1857), syn. nov. Trachyscelida dichroma Viswajyothi & Clark is proposed as a nom. nov. for Racenisa bicolor Bechyné, 1958 (not Agelastica bicolor LeConte, 1884), as both species are currently placed in the genus Trachyscelida Horn, 1893.
The monotypic ant genus Igaponera gen. nov. is proposed to include its type species I. curiosa (Mackay & Mackay, 2010). Igaponera gen. nov. is described and phylogenetically compared with other ponerine genera based on external morphology. The type species is known from a single gyne originally described in the genus Pachycondyla Smith, 1858. Igaponera curiosa is easily diagnosed by: costate sculpture on head, mesosoma, and petiole; short, robust, triangular mandibles with blunt apex; relatively large eyes set at mid-length on sides of head; lack of stridulitrum; and presence of distinct but relatively small arolia. Putative apomorphies of the new genus are: cuticular flange concealing metapleural gland opening; vertically standing hypostomal tooth with recessed base; stout mandibular shape with blunt apex; absence of stout spine-like setae on meso- and metatibial apices. Our phylogenetic results based on morphology suggest that Neoponera Emery, 1901 and Pachycondyla are the closest lineages to Igaponera, which shows intermediate characteristics as compared to those genera. The genus is apparently arboreal, known only from a seasonally flooded Igapó forest near Manaus, Brazil. Despite the collection site being frequented by researchers, no other specimens of this genus have been collected in over 40 years prior to this study.
Descriptions of the following 23 species of Macrotomoderus Pic, 1901 new to science, from continental China, are provided as an addition to the recently published review of the genus from China and Taiwan (Telnov 2018): M. angelinii, M. belousovi, M. bicrispus, M. boops, M. bordonii, M. dali, M. daxiangling, M. femoridens, M. hajeki, M. hartmanni, M. hengduan, M. imitator, M. kabaki, M. korolevi, M. lapidarius, M. muli, M. palaung, M. similis, M. tenuis, M. transitans, M. truncatulus, M. usitatus, and M. wudu spp. nov. Additional records are provided for some poorly known species. The identification key to the species of Macrotomoderus from China, the Japanese Archipelago, and Taiwan is herewith significantly supplemented and updated. Biogeographical peculiarities and altitudinal gradient of Macrotomoderus distribution in continental China are briefly discussed.
Three new six-eyed species of the spider genus Caponina Simon, 1891 are described, photographed, diagnosed and illustrated: C. alejandroi sp. nov. (male and female) from the Boyacá Department, C. bochalema sp. nov. (male) from the Santander Department and C. huila sp. nov. (male) from the Huila Department. Also, a emended diagnosis for the genus, a distribution map and an identification key for all species of Caponina are provided.
Examination of Coccinellidae material recently collected from Napo Province in Ecuador revealed 38 species of ladybird beetles, of which six are new: Chnoodes yanayacu sp. nov., Cyrea mcclarini sp. nov., Eupalea borowieci sp. nov., Exoplectra misahualli sp. nov., Hyperaspis rutai sp. nov., Toxotoma gonzalezi sp. nov. Seven species are recorded for the first time from Ecuador: Cyrea noticollis (Mulsant, 1850), Eupalea formosa Mulsant, 1850, Epilachna transverselineata (Mader, 1858), Psyllobora marshalli Crotch, 1874, Toxotoma fuscopilosa (Weise, 1902), Toxotoma taeniola (Gordon, 1975), Zenoria linteolata Mulsant, 1850. For three species: Epilachna obtusiforma Gordon, 1975, Hinda ecuadorica Gordon & Canepari, 2013 and Siola atra González, 2015, supplementary notes on the original descriptions are provided. New name combinations are proposed for six species, which are transferred to the genus Toxotoma Weise, 1900: Toxotoma aequatorialis (Gordon, 1975), Toxotoma chigata (Gordon, 1975), Toxotoma flavocirculus (González, 2015), Toxotoma fuscopilosa, Toxotoma hybridula (Gordon, 1975), Toxotoma taeniola. The occurrence of Scymnus interruptus (Goeze, 1777) in Ecuador is also confirmed.
The diversity of Palpigradi is not evenly distributed among its different branches. The widespread genus Eukoenenia includes 80% of the species, while the other genera are poorly known. Allokoenenia certainly is the most understudied genus because it is represented only by the African species Allokoenenia afra Silvestri, 1913. Its description is short and does not include many features depicted in modern taxonomy of Palpigradi. In this paper, we describe two troglobitic species of Allokoenenia, report the occurrence of a third species represented by an immature specimen from Brazilian caves, and provide brief notes on the morphology of A. afra. Allokoenenia canhembora sp. nov., A. stygia sp. nov., and Allokoenenia sp. differ from A. afra by several morphological features, including more elongated appendages and a greater number of blades on lateral organs. Thus, they are considered troglomorphic. These new species are vulnerable to extinction because they are endemic to a single or few caves directly impacted by mining activities and groundwater exploitation. This study represents the first step for the conservation of these species and their habitats, since Brazilian caves with rare troglobites cannot be irreversibly impacted. Also, it brings important contributions on the distribution and morphology of this enigmatic genus.
During a taxonomic study of the species of Croton sect. Adenophylli occurring in Brazil, approximately 140 collections usually identified as Croton echioides, C. rhamnifolius or C. rhamnifolioides caught our attention due to the distinct morphology of vegetative and reproductive organs. After the analysis of these collections, we concluded that they represent a new species, C. sertanejus Sodré & M.J.Silva sp. nov., which was also corroborated by an anatomical study of their leaves. This new species can be differentiated by pseudomonopodial branching, leaves with petioles up to 1.7 cm long, 4–6 subsessile nectary glands and indumentum of sessile trichomes on both surfaces. Detailed descriptions, including details of leaf anatomy, are provided for C. sertanejus sp. nov. and C. echioides, its closely related species. We also formalize the synonymization of C. kalkmannii under C. echioides and re-evaluate the typification of these names, in order to better clarify their taxonomic status.
Two new species of Psyllocarpus sect. Psyllocarpus, P. itakangapyra Sobrado, J.A.M.Carmo & R.M.Salas sp. nov. and P. vianae Sobrado, J.A.M.Carmo & R.M.Salas sp. nov., from the “campo rupestre” of the state of Minas Gerais, southeastern Brazil, are here described and illustrated. We provide comments on their distribution, habitat, and preliminary conservation status, and discuss their taxonomy. In addition, we analyse floral, fruit, and seed micromorphology, as well as pollen grains of the new species. We also provide an updated identification key to the species of P. sect. Psyllocarpus.
The nineteen species of Cybocephalidae (Coleoptera) occurring in North America and the West Indies (including Trinidad) are listed and keyed. Cybocephalus skelleyi new species and Cybocephalus edmondsoni new species are described, and Cybocephalus carrabeus T. R. Smith is found to be a new synonym of Cybocephalus geoffereysmithi T. R. Smith. Illustrations of morphological features, including detailed drawings of male genitalia, host records and distribution data, are provided.
ZooBank registration. urn:lsid:zoobank.org:pub:2425983D-0398-45D4-A728-3BF5991D07BE
Benthic samples collected along the Brazilian central continental margin from Espírito Santo State to Rio de Janeiro State (19° S to 24° S) during the last 20 years and at depths of 50–2200 m yielded the description of six new species of Typhlotanais in the deep-sea (> 200 m): Typhlotanais andradeorum sp. nov., T. bolarticulus sp. nov., T. ischnochela sp. nov., T. longiseta sp. nov., T. priscilae sp. nov. and T. spinibasis sp. nov. Within these species, we found members of four Typhlotanais morpho-groups as ‘cornutus’ group, ‘greenwichensis’ group, ‘spinicauda’ and ‘trispinosus’ group. Typhlotanais ischnochela sp. nov. exhibited the greatest bathymetric range, occurring from the continental shelf until the lower slope (46–1898.7 m depth). The bathymetric distribution of the other five species of Typhlotanais herein described were restricted to the slope. Tables of diagnostic characters of the Typhlotanais morpho-groups and an identification key to the species of Typhlotanaidae found in Brazil are provided. This work brings the total number of Tanaidacea known for Brazilian coast to 64 species, and the family Typhlotanaidae from Brazil is now represented by 11 species in five known genera.
A new genus, Thaicypris gen. nov., in the tribe Herpetocypridini Kaufmann, 1900 of the subfamily Herpetocypridinae Kaufmann, 1900 is established to accommodate a new species from Thailand. The present contribution deals with the description of a new genus and species, Thaicypris panhai gen. et sp. nov., which is mainly characterized by the distinctive and raised, inwardly displaced selvage at the postero-ventral part of the right valve (RV) that is not parallel to the valve margin, the absence of an anterior inner list on the RV, the prominent and elevated double inner list on the posterior part of the left valve, the small and three-segmented Rome organ on the first antenna (A1), the spatulated terminal segment of the maxillular (Mx1) palp, the slender caudal ramus (CR) with long and thin Sp seta, the presence of basal triangle on the CR attachment, and the pointed projection at the terminal segment base of the prehensile palps. The hemipenis of the new genus and species is outstanding, especially the medial lateral shield which has a long, beak-shaped protrusion on the distal part. The discovery of this Thai taxon is the first record of the tribe Herpetocypridini in Thailand and the second species of the tribe in Southeast Asia.
Martensina thailandica gen. et sp. nov., a freshwater ostracod species representing a new subfamily, Martensininae subfam. nov., in the family Cyprididae, is here described from a swamp in Maha Sarakham Province, Thailand. The new genus and species is mainly characterized by the 7-segmented antennula which has a Rome organ and remarkably long aesthetasc ya, the morphology of the sexually dimorphic antenna (A2), the markedly elongated A2 terminal segment, the short and thin α- and β-setae on the mandibular palp, the elongated terminal segment of the maxillula, the obviously 2-segmented male prehensile palp, the presence of d1 and d2 setae on the protopod of the second thoracopod (T2), the sexually dimorphic T2, the distinctive terminal segment of the third thoracopod bearing three long setae, the well-developed caudal ramus, the large hemipenis which has a complex internal structure, and the Zenker organ with funnel-shaped ends and numerous spiny whorls.
New information is presented for Neotropical Cerambycidae (Coleoptera). Dolichestola vittipennis Breuning, 1948 is synonymized with D. annulicornis Breuning, 1942, and the species is newly recorded from the Brazilian state of São Paulo. Dolichestola densepunctata Breuning, 1942 is newly recorded from Venezuela and Brazil, and the difference between it and D. annulicornis is reported. Mecas skillmani Santos-Silva and Androw, new species, is described from Mexico (Jalisco). Pannychella callicera (Bates, 1881) is illustrated, and notes on the genus and species are provided. Pannychis Thomson, 1864 is considered a genus different from Mecas LeConte, 1852, and notes on the genus and P. sericea Thomson, 1864, new combination, are given; therefore, Mecas has no subgenera. Pannychina Gilmour, 1962 is synonymized with Dylobolus Thomson, 1868, and Pannychina atripennis (Bates, 1885) is synonymized with Dylobolus rotundicollis Thomson, 1868.
ZooBank registration. urn:lsid:zoobank.org:pub:DCAB0F19-79E2-462F-B7AB-940BD901237D
Eight new Neotropical species of Dexosarcophaga Townsend, 1917 are described, five from Brazil, Dexosarcophaga phoenix sp. nov., Dexosarcophaga jandainae sp. nov., Dexosarcophaga patiuorum sp. nov., Dexosarcophaga petra sp. nov., and Dexosarcophaga sphaera sp. nov., one from Costa Rica, Dexosarcophaga limon sp. nov., one from Ecuador, Dexosarcophaga napo sp. nov., and one from Colombia, Dexosarcophaga pallida sp. nov. Male and female morphology is documented with photographs and illustrations, including details of the male terminalia for all new species and female terminalia of Dexosarcophaga phoenix sp. nov. and Dexosarcophaga sphaera sp. nov. With the addition of these new species, 58 species of Dexosarcophaga are now known, with records from the American continent spanning from the southern United States to northern Argentina.
A revision of microscope slides deposited between 2014 and 2017 in the collection of Rocha and Doma (National University of La Pampa, Argentina) revealed three new species of the genus Milnesium Doyère, 1840: M. pelufforum sp. nov., M. irenae sp. nov. and M. quiranae sp. nov. Milnesium pelufforum sp. nov. is mostly characterized by ten transverse bands of sculptured cuticle and pseudoplates (the first band, until now, never detected in the genus), six peribuccal lamellae and claw configuration [2-2]-[2-2] in young or [2-3]-[3-2] in senior specimens. Milnesium irenae sp. nov. is mostly characterized by complex cuticular ornamentation including a fine reticulation different from the typical one in the genus; it also has pseudoplates, six peribuccal lamellae, medioventral peribuccal papilla reduced, stylets, their furcae and supports very developed, and claw configuration [2-3]-[2-2]. Milnesium quiranae sp. nov. is mostly characterized by smooth cuticle, six peribuccal lamellae, and claw configuration [3-3]-[3-3]; with growing, the medioventral peribuccal papilla reduces and the buccal tube becomes wider. With the present contribution the genus Milnesium now has 48 valid species, and the number of described limno-terrestrial tardigrade species from South America has risen to 11, including 8 from Argentina with 5 from Salta and La Pampa province.
Two new species of Ageniella Banks, 1912 (Hymenoptera: Pompilidae) from Brazil and updated keys
(2022)
Ageniella Banks, 1912 is a paraphyletic group exclusive to the Nearctic and Neotropical regions. The genus has a remarkable morphological diversity among species and strong sexual dimorphism, hampering taxonomic studies of it. Herein, we add two new species to the Neotropical fauna of Ageniella: A. caerulea sp. nov. belonging to the subgenus Ameragenia and A. ruschi sp. nov. belonging to the subgenus Priophanes; both species are described and illustrated. A brief discussion of subgeneric characters and an updated taxonomic key to the species of Ageniella known from Brazil are provided.
Here, we present the results of our field survey in Sri Lanka and describe ten new species of Sericini: Selaserica fabriziae sp. nov., Sel. sororinitida sp. nov., Neoserica pophami sp. nov., Maladera haniel sp. nov., M. kishi sp. nov., M. windy sp. nov., M. karunaratnae sp. nov., M. hiyarensis sp. nov., M. dambullana sp. nov., and M. deenstana sp. nov. All seven of the newly described species of Maladera belong to the M. fistulosa species group, which is an endemic radiation on the island that is characterized by entirely reduced or fused parameres. An updated key to the Maladera fistulosa group is provided. Further, new locality records for 23 already known species are given. The genitalia and habitus of all new species are illustrated, the distribution of the new species is shown with maps.
A new species of the Neotropical genus Platygonia Melichar, 1925 is described and illustrated from the municipality of Ipixuna, State of Amazonas, Northern Brazil. Platygonia nigra sp. nov. can be distinguished from the other species of the genus by the following combination of features: (1) dark brown to black ground color of dorsum; (2) presence of a white to pale yellow spot at the distal portion of corium; (3) male pygofer with a conspicuous diagonal cleft; (4) connective Y-shaped, keeled, with the stem longer than the arms; and (5) aedeagus with an unpaired basiventral process directed anteriorly. This is the first record of the genus from the Brazilian Amazon Rainforest. Notes on P. undecimmaculata (Fowler, 1899), which is a taxon of uncertain taxonomic position, a key to the species of Platygonia, and a map showing their distribution are added.
La distribución geográfica y las características morfológicas de Ectinogonia chalyboeiventris chalyboeiventris Germain y Kerremans, 1906 y E. chalyboeiventris wagenknechti Cobos, 1954 (Coleoptera: Buprestidae) fueron identificadas a partir del estudio de parte de sus ejemplares tipo. Como resultado de la comparación de estos taxones con ejemplares de Ectinogonia Spinola recolectados en las provincias de Elqui y Choapa (Región de Coquimbo), se reconocen y describen dos nuevas especies del norte de Chile: E. gemmula Pineda y Mondaca, nueva especie y E. interruptissima Pineda y Mondaca, nueva especie. Se presentan caracteres diagnósticos, fotografías de los adultos y de los órganos genitales del macho y de la hembra, junto a observaciones sobre el hábitat, historia natural, y distribución de los nuevos taxa. Adicionalmente, se presenta una lista actualizada que incluye a la totalidad de las especies del subgénero Ectinogonia descritas hasta el momento.
ZooBank registration. urn:lsid:zoobank.org:pub:7DC20EF3-1568-484B-B1C9-3D1A9F12460E
A new species of the gecarcinucid freshwater crab genus, Spiralothelphusa Bott, 1968, is described from the Andhra Pradesh State of South India, with supporting data from mitochondrial cytochrome c oxidase subunit I. This brings the number of known species of Spiralothelphusa to seven, five from India and two from Sri Lanka. The identity of S. wuellerstorfi (Heller, 1862) has remained uncertain because the former lectotype designation was not clear. Bott (1968), while designating a male lectotype of S. wuellerstorfi from the Naturhistorisches Museum Wien, Vienna, Austria, had neither stated the catalogue number nor the size of the specimen. It is now difficult to trace the particular lectotype among several paralectotypes of the depository. A new lectotype is, therefore, designated herein for S. wuellerstorfi, and the species is redescribed. The new species, S. andhra sp. nov., has morphological affinities with S. wuellerstorfi, but can be distinguished from the latter species by the shape of the male telson and outer margin of the non-twisted portion of the male first gonopod. Phylogenetic analyses of the molecular data also corroborate their separation. An illustrated identification key is provided for the species of Spiralothelphusa.
A new genus and new species of potamid crab, Gurumon gurumayum gen. et sp. nov., are described from the Arunachal Pradesh State of northeastern India. Gurumon gen. nov. has affinities with Abormon Mitra, Pati & Ng, 2021, Pararanguna Dai & Chen, 1985, and some species of Potamiscus Alcock, 1909, but the new genus can easily be differentiated from them mainly by its low external orbital angle, the relatively stouter exopod of the third maxilliped, and the relatively broader male pleonal somite 6. Their male first gonopods are also different from each other. Counting Gurumon gurumayum gen. et sp. nov., India is currently known for ten genera and 24 species of potamiscine freshwater crabs, and Arunachal Pradesh for eight genera and 11 species of these crabs. The current nomenclatural problems with Potamiscus are also discussed.
New records for a new species and other exotic Dirrhagofarsus Fleutiaux (Coleoptera: Eucnemidae: Melasinae: Dirhagini) species are reported from throughout much of the northeastern and southern areas of the United States. Dirrhagofarsus brevis, new species, is hereby described in this study. Two other exotic species, Dirrhagofarsus modestus (Fleutiaux) and Dirrhagofarsus unicolor (Hisamatsu), new country records, are redescribed and diagnosed as they were compared with other Dirrhagofarsus species present in the Nearctic region. All Dirrhagofarsus species in this study are imaged, highlighting essential character states to facilitate better diagnosis in conjunction to the new identification key provided in this study.
ZooBank registration. urn:lsid:zoobank.org:pub:24DCE05F-14CE-465B-AB11-A2F34012FAB6
The amerophidian snake radiation is a Late Cretaceous superfamily that encompasses two families: Aniliidae, pipe snakes, and Tropidophiidae, dwarf boas. We describe a new dwarf boa snake species, from the Tropidophiidae family, from the cloud forest in northeastern Ecuador. Tropidophis cacuangoae sp. nov. can be diagnosed from its congeners based on external and osteological morphology. The new species inhabits eastern tropical piedmont and lower evergreen montane forests, in the Amazon Tropical Rainforest biome, and is likely to be an Ecuadorian endemic. We also discuss the relationships of the new species with South American tropidophiids and provide a key to the identification of mainland South American dwarf boas.
The West Indian genus Onychotillus Chapin (Coleoptera: Cleridae: Tillinae) is revised and includes O. androwi Opitz, new species, O. apiculus Opitz, new species, O. cinctipennis (Chevrolat,1874), O. cubana de Zayas,1988, O. dimidiatus de Zayas,1988, O. lineatus Opitz, new species, O. minutus de Zayas,1988, O. trinitatis de Zayas,1988, O. woodruffi Opitz, new species, and O. vittatus Chapin,1945.
A new polynoid, Webbnesia maculata gen. et sp. nov., was discovered during benthic surveys conducted around the Canary Islands. Its generic characters (absence of cephalic peaks, ventrally inserted lateral antennae, reduced notopodium and chaetae all stout) place it close to Antinoe Kinberg, 1856, Hermadion Kinberg, 1856 and Malmgrenia McIntosh, 1874, but the combination is unique and justifies the erection of a new genus. The new genus and species are described, figured and discussed in detail. An updated list of taxa and an identification key to all genera of Polynoinae Kinberg, 1856 sensu lato currently reported from the extended Northeast Atlantic are given.
The discovery of a new species of the genus Canthocamptus, C. waldemarschneideri sp. nov., in northern Siberia prompted a taxonomic analysis of this genus. In this work, on the basis of cladistic analysis, we show that the genus is not monophyletic. Based on differences in the structure of the endopods on the second pair of male swimming legs, fifth legs of males and females, and caudal rami, we conclude that the Canthocamptus mirabilis species group is a separate genus, Kikuchicamptus gen. nov. Additionally, two species are transferred to the genus Attheyella, and one species, Canthocamptus gibba, is synonymized. The subgenera Canthocamptus (Baikalocamptus) and Canthocamptus (Canthocamptus) are also synonymized. The new species, Canthocamptus waldemarschneideri sp. nov., is most closely related to the American Canthocamptus assimilis Kiefer, 1931 and differs from it in the ornamentation of the abdominal somites and the shape of the caudal setae.
We describe a new species of Theloderma from northeastern Vietnam based on morphological differences and molecular divergence. Theloderma khoii sp. nov. is distinguishable from its congeners on the basis of a combination of the following characters: large size, SVL 52.1 mm in male, 59.4 mm in female; head length and width equal; vomerine teeth present; snout pointed and truncated, eye large, ED 4.7 mm in male, 5.6 mm in female, spinules on upper eyelid; tibiotarsal articulation reaches to the posterior border of the eye or the tip of the snout; dorsal skin very rough with large irregular gland ridges and warts, ventral surface of body granular; tips of all digits dilated but all considerably smaller than tympanum; dorsal surface mossy green or olive mottled with dark magenta. The distribution of the new species is unknown but probably extends into adjacent high elevation forested areas in Ha Giang Province, Vietnam and in Yunnan Province, China with an extent of occurrence of only < 1000 km2 and continuing decline in the quality of its habitat due to deforestation. Thus, we suggest the species should be considered Endangered following IUCN’s Red List categories.
Lodevoisadia coheni gen. et sp. nov. is described as the ninth species of ‘Grylloblattodea’ from the middle Permian of the Salagou Formation, near Lodève town (France). It is currently not reasonable to place this species into a specific family, even though it seems to share most characters with the small family Tunguskapteridae. The lack of phylogenetic analysis and the current poor delineation of the majority of the grylloblattodean families (lacking synapomorphies) render any attribution of new taxa to a particular family often uncertain.
Iran is a huge but understudied Middle Eastern country with a rich but chronically understudied bee fauna, including for the highly-speciose bee genus Andrena. Examination of unidentified museum material combined with recent field collections and a critical review of the literature has revealed a total of 197 species of Andrena in the Iranian fauna, of which 65 are newly reported for the country, with an additional 16 species new for science. Andrena (Aciandrena) deminuta Wood sp. nov., Andrena (Euandrena) boustaniae Wood sp. nov., Andrena (Euandrena) oblata sp. nov., Andrena (Euandrena) sani sp. nov., Andrena (Micrandrena) elam Wood sp. nov., Andrena (Micrandrena) subviridula Wood sp. nov., Andrena (Notandrena) idigna Wood sp. nov., Andrena (Planiandrena) flagrans Wood sp. nov., Andrena (Planiandrena) sella Wood sp. nov., Andrena (Ulandrena) bulbosa Wood sp. nov., Andrena (incertae sedis) hosseiniiae Wood & Monfared sp. nov., and Andrena (incertae sedis) rostamiae sp. nov. are described from Iran, Andrena (Micrandrena) extenuata sp. nov. is described from Iran and Syria, Andrena (Micrandrena) tabula Wood sp. nov. and Andrena (Micrandrena) obsidiana Wood sp. nov. are described from Iran and Turkey, and Andrena (Planiandrena) huma sp. nov. is described from Iran, Syria, and the Golan Heights. Eight taxa are synonymised (valid name first): Andrena (Melandrena) assimilis Radoszkowski, 1876 = Andrena (Melandrena) gallica Schmiedeknecht, 1883 syn. nov.; Andrena (Notandrena) emesiana Pérez, 1911 stat. resurr. = Andrena (Notandrena) recurvirostra Warncke, 1975 syn. nov.; Andrena (Plastandrena) eversmanni Radoszkowski, 1867 = Andrena (Plastandrena) peshinica Nurse, 1904 syn. nov.; Andrena (incertae sedis) hieroglyphica Morawitz, 1876 = Andrena (Carandrena) cara Nurse, 1904 syn. nov. and Andrena (Carandrena) halictoides Nurse, 1904 syn. nov.; Andrena (Melandrena) induta Morawitz, 1894 = Andrena (Melandrena) patella Nurse, 1903 syn. nov.; Andrena (incertae sedis) minor Warncke, 1975 stat. nov. = Andrena (Carandrena) splendula Osytshnjuk, 1984 syn. nov.; Andrena (Notandrena) zostera Warncke, 1975 = Andrena (Carandrena) subsmaragdina Osytshnjuk, 1984 syn. nov. Overall, these results considerably improve our understanding of the Iranian Andrena fauna, and suggest that overall bee diversity in this country is substantially more than 1000 species.
Stromatium chilensis Cerda, 1968, an endemic species from Chile, is placed in Malcho Mondaca and Beéche, new genus (Coleoptera: Cerambycidae: Cerambycinae: Hesperophanini). The new genus is compared with Stromatium Audinet-Serville, included in a previous key to Hesperophanini, and a diagnosis and illustrations of the species are provided.
ZooBank registration. urn:lsid:zoobank.org:pub:215682BF-4BBD-4C7F-BD30-FCF5285C8DB2
The previously endemic Australian genus Habritella Girault & Dodd, 1915 is reported for the first time in the Afrotropical region with four new species: H. africana sp. nov., H. mandibulata sp. nov., H. noyesi sp. nov., and H. viridifrons sp. nov. The males of Habritella are described for the first time. A new generic synonymy and combination are also proposed: Ezgia Koçak & Kemal, 2008 syn. nov. and H. stylifera (Bouček, 1988) comb. nov. Habritella is redescribed to accommodate the newly discovered and transferred species. An illustrated key to the Afrotropical species is provided.
The present study examines whether the chaetotaxy of the costal vein in the calyptrate families Fanniidae and Muscidae deserves more attention in phylogenetic and taxonomic contexts. An overview of the macrotrichia and their arrangement on wing vein C is given. Special attention is given to the presence/absence of ventral and dorsal setulae on the costal sectors CS1‒3. This is described as one variable character (A) with nine states (A0‒A8). Specimens of both sexes (when possible) of each species belonging to 4 of a total of 5 fanniid genera and 115 of a total of 179 muscid genus-group taxa were examined and scored for character A. It was found that the presumed ancestral state of character A differs between the two families. It is further shown that the main transformational trend in character A in Muscidae has been bi-directional, leading either to the loss of ventral setulae or the gain of dorsal setulae. The utility of character A in the Fanniidae and Muscidae is many-sided and involves taxa ranging from species to family. It is concluded that character A and other aspects of costal chaetotaxy deserve more attention in morphology-based studies of calyptrate flies.
Revision of the Eurasian species of Aegilips Haliday, 1835 (Hymenoptera: Figitidae: Anacharitinae)
(2022)
The knowledge of Aegilips in Eurasia is updated. One new species is described: Aegilips insularis Mata-Casanova and Pujade-Villar sp. nov., from Portugal (Madeira). The genus is also cited for the first time in the Indomalayan region with specimens of A. atricornis found in Northern India and Pakistan. Aegilips curvipes Giraud, 1860, Aegilips gemellus Belizin, 1961, Aegilips laevis (Hedicke, 1914), Aegilips montanus Belizin, 1951 and Aegilips punctatus Belizin, 1951 are synonymized with A. nitidulus (Dalman, 1823). Aegilips vena Fergusson, 1985 is synonymized with Aegilips zaitzevi Kovalev, 1974. Aegilips notatus Belizin, 1951, and Aegilips punctulatus Hedicke, 1928 are designated as incertae sedis. The known distributions of A. atricornis, A. kozlovi, A. nitidulus and A. romseyensis are expanded. Morphological differences are discussed and a key for Eurasian Aegilips species is also given.
A global synonymical checklist of the species and higher taxa of the insect order Megaloptera is provided. The checklist includes both extant and extinct taxa, and recognizes 2 families, 4 subfamilies, 48 genera, 425 species, and 6 subspecies. Both families (Corydalidae and Sialidae), and three of the four subfamilies (Corydalinae, Chauliodinae, and Sialinae) are known from both extant and extinct species; the Sharasialinae (Sialidae) is entirely extinct. Country-level geographic distribution data are provided for all species and subspecies. Synoptic type data are provided for taxa in the family and genus groups. Summary data are given for the numbers of megalopteran species currently known to occur in each of the major biogeographical regions of the world, and for the world fauna. Increase of knowledge about the diversity of the world Megaloptera fauna is summarized in counts of valid species described per decade and in a global taxonomic description curve. An updated set of keys to the world families, subfamilies, and genera of the Megaloptera is also provided.
The genus Pareucamptonyx Olmi, 1991 is endemic to the New World and comprises three described species, two from the Neotropical region and one from the Nearctic. Here is presented the first revision of species of Pareucamptonyx from the Neotropical region, including four new species described from Brazil: P. albopictus Martins sp. nov., P. kumagaiae Martins sp. nov., P. niger Martins sp. nov. and P. paranaensis Martins sp. nov. Detailed illustrations, distribution map, and key to females are provided, as well as a revised diagnosis of the genus and notes about the method of collection. Additionally, new distribution records for P. townesi (Olmi, 1984) are reported.
Labahitha spiders (Arachnida: Araneae: Filistatidae) from islands in the Indian and Pacific Ocean
(2022)
The genus Labahitha has hitherto comprised two species from peninsular Malaysia and Christmas Island (Australia). We here demonstrate that the genus is widespread in islands and territories across the Indian and Pacific Oceans, including the following species that have been previously assigned to other filistatid genera: Labahitha marginata (Kishida, 1936) comb. nov. (= Filistata bakeri Berland, 1938 syn. nov.), Labahitha garciai (Simon, 1892) comb. nov. (= Pritha heikkii Saaristo, 1978 syn. nov., = Pritha sechellana Benoit, 1978 syn. nov.), Labahitha nicobarensis (Tikader, 1977) comb. nov., Labahitha littoralis (Roewer, 1938) comb. nov., Labahitha insularis (Thorell, 1891) comb. nov., Labahitha sundaica (Kulczyński, 1908) comb. nov. (all transferred from Pritha, the latter three provisionally, pending re-examination of the type material); Labahitha fuscata (Nakatsudi, 1943) comb. nov. and Labahitha ryukyuensis (Ono, 2013) comb. nov. (both transferred from Tricalamus). Many of these species have been collected in synanthropic settings and from disparate islands thousands of kilometers apart. This suggests either high dispersal capabilities or, more likely, human-mediated introductions. At least L. marginata has been introduced to continental America. Two new species of Labahitha are described: Labahitha platnicki sp. nov. from New Caledonia and the Bismarck Islands and Labahitha incerta sp. nov. from Queensland, Australia. The male of Labahitha gibsonhilli (Savory, 1943) is reported for the first time. Wandella loloata sp. nov. is described from Papua New Guinea, representing the first record of this genus outside Australia. Pritha hasselti (Simon, 1906) from Indonesia is shown to be a Filistatinae, and thus the species is provisionally transferred back to Filistata.
A revision of the wild species in the genus Malus Mill. (Rosaceae) is presented based on numerical analyses and specimens from herbaria around the world, while cultivated species such as Malus domestica (Suckow) Borkh. are not included because of their complicated domestication history. Infra- and interspecific morphological variation and species delimitation are clarified based on Principal Component Analyses (PCA) and Cluster Analyses (UPGMA). We found that several morphological characters traditionally used to distinguish species have limited taxonomic value because of high phenotypic variation or plasticity. There is a substantial conflict between traditional morphological and genetic taxonomic concepts, and as a result species lineages are often morphologically indistinguishable. None of the analyses supports the recognition of infraspecific categories in Malus transitoria (Batalin) C.K.Schneid. and interspecific categories between Malus doumeri (Bois) A.Chev. and Malus leiocalyca S.Z.Huang. Based on our analyses, we recognize 26 wild species in the genus, and propose seven new synonymies.
A key, an annotated checklist with detailed distribution, biological and host information, and color photographic plates are provided for the 91 species of dacine fruit flies (Diptera: Tephritidae: Dacini) known to occur in Oceania. Dacus virgatus Coquillett, previously a synonym of Bactrocera psidii (Froggatt), is instead considered a junior synonym of B. facialis (Coquillett). The species originally described in 1971 as Dacus (Asiadacus) perpusillus Drew, later reassigned as Bactrocera (Sinodacus) perpusilla (Drew) and in recent years as Zeugodacus (Sinodacus) perpusillus (Drew) actually belongs to genus Dacus, and is transferred back to Dacus, but to the subgenus Neodacus, restored combination. The presence of B. redunca (Drew) is recorded for the first time in New Caledonia. New male lure records include isoeugenol and dihydroeugenol for both B. neoxanthodes Drew and Romig and B. quadrisetosa (Bezzi) and zingerone for Dacus taui (Drew and Romig), all in Vanuatu.
ZooBank registration. urn:lsid:zoobank.org:pub:A105F057-F2A4-4C14-B82E-14912B319D57
A key, an annotated checklist with detailed distribution, biological and host information, and color photographic plates are provided for the 91 species of dacine fruit flies (Diptera: Tephritidae: Dacini) known to occur in Oceania. Dacus virgatus Coquillett, previously a synonym of Bactrocera psidii (Froggatt), is instead considered a junior synonym of B. facialis (Coquillett). The species originally described in 1971 as Dacus (Asiadacus) perpusillus Drew, later reassigned as Bactrocera (Sinodacus) perpusilla (Drew) and in recent years as Zeugodacus (Sinodacus) perpusillus (Drew) actually belongs to genus Dacus, and is transferred back to Dacus, but to the subgenus Neodacus, restored combination. The presence of B. redunca (Drew) is recorded for the first time in New Caledonia. New male lure records include isoeugenol and dihydroeugenol for both B. neoxanthodes Drew and Romig and B. quadrisetosa (Bezzi) and zingerone for Dacus taui (Drew and Romig), all in Vanuatu.
ZooBank registration. urn:lsid:zoobank.org:pub:A105F057-F2A4-4C14-B82E-14912B319D57
The tribe Amarotypini (Coleoptera: Carabidae: Migadopinae) is revised for New Zealand. Three genera and fourteen species are recognized.
Two genera and thirteen species are described as new: Amarophilus Larochelle and Larivière new genus, Amarophilus lomondensis Larochelle and Larivière new species, Amarophilus otagoensis Larochelle and Larivière new species, Amarophilus rotundicollis Larochelle and Larivière new species, Amarophilus wanakensis Larochelle and Larivière new species, Amarotypus fiordlandensis Larochelle and Larivière new species, Amarotypus glasgowensis Larochelle and Larivière new species, Amarotypus murchisonorum Larochelle and Larivière new species, Amarotypus simoninensis Larochelle and Larivière new species, Amarotypus takaheensis Larochelle and Larivière new species, Amaroxenus Larochelle and Larivière new genus, Amaroxenus arnaudensis Larochelle and Larivière new species, Amaroxenus glacialis Larochelle and Larivière new species, Amaroxenus huttensis Larochelle and Larivière new species, Amaroxenus kahurangiensis Larochelle and Larivière new species.
A revision of all taxa is provided. Descriptions, identification keys, illustrations of male genitalia, habitus photos, distributional data and maps are given. Information on ecology, biology, dispersal power, and collecting techniques is included for each species.
ZooBank registration. urn:lsid:zoobank.org:pub:6BBC7A99-0736-44D1-BAD1-3C719F9A69C2
The subfamily Carventinae (Heteroptera: Aradidae) is revised for New Zealand. Eight genera and fifteen species are recognized. One genus and six species are described as new: Carventaptera hallae Larivière and Larochelle new species, Lissaptera heissi Larivière and Larochelle new species, Modicarventus kirmani Larivière and Larochelle new species, Neocarventus montanus Larivière and Larochelle new species, Neocarventus northlandicus Larivière and Larochelle new species, Neocarventus potterae Larivière and Larochelle new species, Tuataraptera Larivière and Larochelle new genus. One new combination is established: Neocarventus uncus Kirman, 1989 = Tuataraptera unca (Kirman, 1989). One new synonymy is made: Leuraptera yakasi Heiss, 1990 = Leuraptera zealandica Usinger and Matsuda, 1959. A revision of al taxa is provided. Descriptions, identification keys, illustrations of male genitalia, habitus photos, distributional data and maps are given. Extensive information on biology is included for each species.
ZooBank registration. urn:lsid:zoobank.org:pub:CAF794A0-89C7-498F-84D0-940FDDB648F3
Early Carboniferous coiled nautiloids from North Africa are virtually unknown. An assemblage of nine species, all from the family Trigonoceratidae, from the Dalle à Merocanites (Tournaisian-Viséan boundary interval) of Timimoun in western Algeria is described, being the most diverse Carboniferous nautiloid assemblage known from North Africa but much less diverse than the time-equivalent assemblages from Belgium and Ireland. The assemblage consists of the species Maccoyoceras pentagonum sp. nov., Lispoceras orbis sp. nov., Thrincoceras devolvere sp. nov., Rineceras multituberculatum sp. nov., Rineceras rectangulatum sp. nov., Vestinautilus padus sp. nov., Vestinautilus concinnus sp. nov., Planetoceras destrictum sp. nov. and Planetoceras transforme sp. nov. A morphometric analysis of Maccoyoceras pentagonum sp. nov. and Lispoceras orbis sp. nov. shows that the intraspecific variation in these species is within rather narrow limits.
The ammonoids of the suborder Gephuroceratina from the Roteisenstein (Red Ironstone) Formation of the area around Dillenburg (eastern Rhenish Mountains) are revised, mainly based on historical collections stored in the Museum für Naturkunde, Berlin. The new species Ponticeras materni sp. nov. is described and the species Pseudoprobeloceras pernai (Wedekind, 1918), Pseudoprobeloceras applanatum (Wedekind, 1918), Ponticeras aequabile (Beyrich, 1837), Darkaoceras galeatum (Matern, 1931), Taouzites acutus (Matern, 1931), Koenenites lamellosus (Sandberger & Sandberger, 1851), Acanthoclymenia forcipifera (Sandberger & Sandberger, 1851) and Acanthoclymenia planorbis (Sandberger & Sandberger, 1851) are revised. The stratigraphic distribution of the genera is discussed; they are assigned to three assemblages: (1) Pseudoprobeloceras pernai Zone (latest Givetian; genera Pseudoprobeloceras, Ponticeras, Darkaoceras and Taouzites), (2) Koenenites lamellosus Zone (early Frasnian, containing Koenenites lamellosus and Acanthoclymenia forcipifera) and (3) Mesobeloceras kayseri Zone (middle Frasnian, containing Acanthoclymenia planorbis).
The tornoceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea)
(2022)
The ammonoids of the suborder Tornoceratina from the Middle to Late Devonian Red Ironstone Formation of the area around Dillenburg (eastern Rhenish Mountains) are revised, mainly based on historical collections stored in the Museum für Naturkunde, Berlin. The species Tornoceras typus (Sandberger & Sandberger, 1851), Tornoceras frechi Wedekind, 1918 and Epitornoceras mithracoides (Frech, 1888) are re-described based on the original material from the Red Ironstone. The new genus Lentitornoceras gen. nov. is proposed for the new species L. materni gen. et sp. nov.; the new genus Paucitornoceras gen. nov. is proposed with the type species Goniatites paucistriatus. Epitornoceras transmediterraneum sp. nov. is described from the Anti-Atlas of Morocco and compared with E. mithracoides.
This review considers a fascinating, from a zoogeographical viewpoint, group of closely related species: Melitaea lutko Evans, 1932, M. timandra Coutsis & van Oorschot, 2014, M. mimetica Higgins, 1940 stat. rev. and M. shahvarica sp. nov. It is a taxonomical and geographical review of these species, and data on the biology of M. shahvarica sp. nov. and nominate subspecies of M. timandra are discussed. A new species, M. shahvarica sp. nov. from Shahvar Mt. (Iran), and a new subspecies, M. timandra binaludica subsp. nov. from Kuh-e-Binalud Mts (Iran), are described. The specific structure of the group given in previous publications is critically evaluated. Hypotheses about a possible phylogenesis of the study group are provided.
Two new opilionid species from suborder Cyphophthalmi, family Sironidae, Siro franzi Karaman & Raspotnig sp. nov. and Siro ozimeci Karaman sp. nov., from Austria and Croatia respectively, are described and illustrated. Both species show a close relation to two other relict sironid species from the southern and eastern parts of the Alps, Siro valleorum and Siro crassus. All four species are treated here as a monophyletic, alpine group of genus Siro, opposed to the remaining two European sironids, S. rubens and S. carpaticus (palaeoeuropean Siro group). The history of the alpine Siro group parallels the history of a part of the dynamic European archipelago in the Mediterranean Tethys area, which became a part of the Alpine orogeny. Diversification of the alpine Siro group is the result of the orogenic evolution of the Alps, linked to the Austroalpine and South Alpine tectonic units.
Bureschia serbica sp. nov., is described from a cave in the Stara Planina Mt in Serbia as a second species in the genus. It is a troglobite amphibious species, predominantly aquatic like the nominal species of the genus, Bureschia bulgarica. The new species is adapted to amphibious life in running groundwater, unlike the nominal one that lives in subterranean lakes and ponds. Body structure changes, most noticeable the significant changes in the structure of the mouthparts, as a result of preadaptation in aquatic and amphibious Synocheta, are discussed.
Lissomus quisqueya new species and L. woodruffi new species are described. Both species are reported from the Dominican Republic on the island of Hispaniola. These are the first species of Lissomus Dalman reported from the Greater Antilles. Lissomus quisqueya is recorded from montane mesic forests in the Cordillera Central from Dajadon and La Vega provinces. Lissomus woodruffi is recorded from Barahona, Independencia and Pedernales provinces from montane mesic forest in the Sierra de Bahoruco. Morphologically, these species are most similar to members of the L. discedens Bonvouloir species group from Mesoamerica and South America. Known sites of occurrence of L. quisqueya and L. woodruffi are remnants of formerly more extensive mesic forests. Some of the specimens were found in secondary forests and ecotonal areas at sites near to primary forest or remnants, and all these forests are threatened by anthropogenic activities and stochastic climate changes.
The genus Andromma Simon, 1893 is revised. The type species A. aethiopicum Simon, 1893 (♂♀) and the other three known species, A. anochetorum Simon, 1909 (♂), A. bouvieri Fage, 1936 (♀) and A. raffrayi Simon, 1899 (♂♀), are redescribed. The types of the subspecies A. raffrayi inhacorense Lessert, 1936 were not examined. This subspecies is considered a synonym of the nominal species. Nineteen new species are described and illustrated: A. albinovani sp. nov. (♂♀), A. alvoculatum sp. nov. (♀), A. anacardium sp. nov. (♀), A. cyamos sp. nov. (♀), A. cycnotrachelos sp. nov. (♂♀), A. delphiurum sp. nov. (♂♀), A. deogratias sp. nov. (♂♀), A. dicranobelos sp. nov. (♂), A. didrepanum sp. nov. (♂), A. divinagraciae sp. nov. (♂♀), A. elephantactes sp. nov. (♀), A. ghesquierei sp. nov. (♂♀), A. heligmos sp. nov. (♀), A. helix sp. nov. (♀), A. juakalyi sp. nov. (♂), A. katangensis sp. nov. (♀), A. ophiophagum sp. nov. (♀), A. prosopion sp. nov. (♀) and A. velum sp. nov. (♂♀). The distribution of the species and their apparent commensalism with termites are discussed.
Psychodid specimens from one of the world’s biodiversity hotspots – the Caucasus region – were sorted from samples collected during field work in 2019. Pneumia fuehzulii sp. nov. (from two localities in Azerbaijan) and Thornburghiella salihi sp. nov. (from one locality in Azerbaijan and four sites in Georgia) are described from this material. Differential diagnoses are given and some morphological characters illustrated. Sijaricia Krek, 1999, syn. nov. of Thornburghiella Vaillant, 1982 is proposed and T. erinacea (Krek, 1971) comb. nov. from Ulomyia Walker, 1856 is suggested.
Revisions of Holocnemus and Crossopriza: the spotted-leg clade of Smeringopinae (Araneae, Pholcidae)
(2022)
The genera Holocnemus Simon, 1873 and Crossopriza Simon, 1893 are revised. Together with Stygopholcus Kratochvíl, 1932 (revised recently) and the newly described genus Maghreba gen. nov., they constitute the spotted-leg clade within the northern clade of Smeringopinae. Males and females in this group are characterized by dark marks on the leg femora and tibiae. The native area of the spotted-leg clade ranges from northern Africa and the Mediterranean to Central Asia and NW India. A morphological cladistic analysis suggests that Holocnemus is paraphyletic while Crossopriza is monophyletic, but morphology seems only partly adequate to resolve phylogenetic relationships convincingly. The genus Holocnemus includes four species, all of which are redescribed: H. pluchei (Scopoli, 1763); H. reini (C. Koch, 1873) comb. nov. (transferred from Pholcus); H. caudatus (Dufour, 1820); and H. hispanicus Wiehle, 1933. The genus Maghreba gen. nov. includes eight species from NW Africa: M. aurouxi (Barrientos, 2019) gen. et comb. nov. (transferred from Holocnemus; redescribed, female newly described) and seven newly described species. The genus Crossopriza includes six previously described species (of which five are redescribed), and 18 newly described species. The Madagascan C. nigrescens Millot, 1946 is synonymized with C. lyoni (Blackwall, 1867). All new species are described on the basis of both sexes.
Four new species of limnoterrestrial rhabdocoels (‘Typhloplanidae’ Graff, 1905) are described. One of these – Faunulus nielsi Houben, Proesmans & Artois gen. et sp. nov. – could not be unambiguously placed within an existing genus. Faunulus nielsi most closely resembles species of the genus Adenocerca Reisinger, 1924 but can be clearly distinguished by the position of the testes. The three other new species described are Bryoplana belgica Houben, Proesmans & Artois sp. nov., Hoplopera isis Houben, Proesmans & Artois sp. nov., and Protoplanella leiae Houben, Proesmans & Artois sp. nov. All three belong to the subfamily ‘Protoplanellinae’ Reisinger, 1924 and are distinguished based on a detailed description of the reproductive system. Finally, new data are provided for nine other, known typhloplanids: Adenocerca minima Kolasa, 1981; Chorizogynopora italica Kolasa, 1981; Hoplopera opaca Reisinger, 1924; K. subterranea Reisinger, 1933; Krumbachia virginiana (Kepner & Carter, 1931) Ruebush, 1938; Olisthanellinella rotundula Reisinger, 1924; Prorhynchella minuta Ruebush, 1939; Protoplanella simplex Reisinger, 1924; and Ventrociliella romanae Kolasa, 1977. A detailed comparison of our material of V. romanae to what is described for Bockia deses Reisinger, 1924, leads us to consider the latter a nomen dubium.
A list of abbreviations regarding literature, collections and persons as used by early authors (1758–1779) of scarabaeoid beetles (Insecta: Coleoptera) is given together with modern referrals to the Literature Cited. Notes regarding referential errors are included. Hyperlinks to all mentioned and freely online available publications are provided.
ZooBank registration. urn:lsid:zoobank.org:pub:8508A5D0-CA65-4BBB-9FD4-8D14AC261F72
The Bittacidae fauna in Guizhou Province, China is reviewed. Eleven species in the genera Terrobittacus Tan & Hua, 2009 and Bittacus Latreille, 1805 of Bittacidae are documented in Guizhou, including three new species: Bittacus dilobus sp. nov. and Bittacus leigongshanicus sp. nov. from Leigongshan, and Bittacus multisetus sp. nov. from Yushe. A key to species of Bittacidae in Guizhou is provided.
A new lichen species Lecaimmeria pakistanica K.Habib, R.Zulfiqar & Khalid sp. nov. is described and illustrated from rocks in the temperate forests of the Himalaya of Azad Jammu and Kashmir, Pakistan. This species is characterized by its yellow-brown to brown thallus having areoles 0.4 to 1.5 mm across, branched and anastomosing paraphyses, a tall hymenium, large ascospores 20–32 × 10–16 μm, and no substance detected by thin layer chromatography. All other species of the genus have ascospore dimensions in the range of 14–22 × 5–14 μm. A phylogenetic analysis is provided based on ITS nrDNA sequences, and supports the separation of the novel species. Photographs and a comparative analysis with related species of Lecaimmeria are provided to confirm the status of the species.
Taxonomy of fourteen very little known species of Nodosariinae Ehrenberg, 1838 in Icelandic waters is revised. Knowledge of these species in the North Atlantic relies mainly on studies in the late 19th and early 20th centuries, using large volume samplers. Later studies have emphasized quantitative samples of a few cm3 where the Nodosariinae are very rare. This study analysed 879 dredging samples where Nodosariinae occurred in 492 samples, comprising 7598 specimens of about 415 000 of all picked foraminifera. Ordination analysis of species distributions reflects prominent temperature and salinity differences that exist in the sampling area (753 000 km2) north and south of the Greenland-Scotland Ridge (GSR). Eight species are restricted to southern temperate waters (> 2°C): Dentalina mutabilis (Costa, 1855), Dentalina antarctica Parr, 1950, Dentalina antennula d’Orbigny, 1846, Dentalina filiformis (d’Orbigny, 1826), Grigelis pyrula (d’Orbigny, 1826), Grigelis guttifera (d’Orbigny, 1846) comb. nov., Grigelis semirugosus ? (d’Orbigny, 1846) and Nodosaria subsoluta Cushman, 1923. Four species (Nodosaria haliensis Eiland & Guðmundsson, 2004, Nodosaria incerta Neugeboren, 1856, Dentalina elegans d’Orbigny, 1846 and Dentalina frobisherensis Loeblich & Tappan, 1953) occur mainly north of Iceland. Two species, Dentalina obliqua (Linnaeus, 1758) and Pseudonodosaria subannulata (Cushman, 1923), have wide tolerance ranges for physical variables.
Limited data are available for the kinorhynch fauna from the Southern Hemisphere, with little or no data from New Zealand. Here, we provide a first comprehensive overview of the diversity of mud dragons, with an emphasis on species of Echinoderes from the continental slope of New Zealand, from a variety of habitats such as slopes, canyons and seamounts located in the Hikurangi Margin region. The study revealed fifteen species of Echinoderes. Of these, ten are described as new to science: E. aragorni sp. nov., E. blazeji sp. nov., E. dalzottoi sp. nov., E. frodoi sp. nov., E. galadrielae sp. nov., E. gandalfi sp. nov., E. landersi sp. nov., E. leduci sp. nov., E. legolasi sp. nov. and E. samwisei sp. nov. Moreover, Echinoderes juliae Sørensen et al., 2018, Echinoderes sp. aff. E. balerioni, Echinoderes sp. aff. E. galadrielae/beringiensis, Echinoderes sp. aff. E. lupherorum and Echinoderes sp. aff. E. unispinosus are reported in the investigated region. The most abundant among all was E. gandalfi sp. nov., but it was found only in canyons. Interestingly, the second most common species was E. juliae that was found at several stations in canyons, seamount and on the slope. This species is one of the deep-sea species originally found on the abyssal plain off Oregon and along the continental rise off California, Northeast Pacific, recorded in polymetallic nodules in the tropical eastern Pacific, and recently found on the abyssal plains off Chile, east of the Atacama Trench. These findings, together with records of Echinoderes sp. aff. E. lupherorum and Echinoderes sp. aff. E. unispinosus indicate that, despite their low dispersal abilities, kinorhynchs, similar to other meiofaunal species, may exhibit a wider distribution pattern than previously assumed. The number of recorded species and numerous new species show that New Zealand sediments not only are inhabited by a diverse kinorhynch fauna, but Echinoderes, the most speciose genus, still holds much to discover.
The kinorhynch fauna from Portugal has been explored, yielding a new species of the genus Setaphyes (Kinorhyncha: Allomalorhagida). This is the first description of an allomalorhagid species from Portugal. Specimens of the new species were collected at a subtidal muddy beach in Alvor, a village located in the southernmost region of Portugal. Setaphyes algarvensis sp. nov. may be distinguished from its congeners by a unique arrangement of the setae: paired paradorsal setae on segments 2–7 and 9, paradorsal seta on segment 8 unpaired, laterodorsal setae on segments 2–3 and 6–9 in males and 2–9 in females, paralateral setae on segment 1, lateroventral setae on segments 2–10 (two pairs on segment 5), ventrolateral setae on segment 1 in males and 1–3 in females, and ventromedial setae on segments 3–9 in males and 4–9 in females. The diagnostic features of Setaphyes algarvensis sp. nov. are discussed from a comparative perspective with the congener species. Additionally, morphometric analyses of selected features, namely the total trunk length and the relation between the total trunk length and the length of lateral terminal spines, turned out to be useful to distinguish between the new species and Setaphyes kielensis (its most similar congener).
Two novel species of Russula (Russulaceae, Russulales), namely Russula indosenecis A.Ghosh, D.Chakr., K.Das & Buyck sp. nov. and R. pseudosenecis A.Ghosh, D.Chakr., K.Das & Buyck sp. nov. belonging to sect. Ingratae subg. Heterophyllidiae are proposed herein based on their morphological features and nrITS-based phylogenetic inferences. Both species belong to the Asian ʻR. punctipes-senecisʼ complex of sect. Ingratae. The acrid R. indosenecis was collected from subalpine forests associated with Abies densa, whereas the mild R. pseudosenecis associates with tropical forests dominated by Shorea robusta. Both species are distinct from the other species of this species complex in nrITS sequence data and from all other known species in subg. Heterophyllidiae in the strong amyloidity of their suprahilar spot.
The finding of Breviconia andrei sp. nov. in the Russian Bering Sea enabled the clear-cut phylogenetic characterization of the former monotypic genus Breviconia Conroy-Dalton & Huys, 2000 as a monophylum. Comparison of the new species with B. australis (George, 1998) and other members of the subfamily Ancorabolinae Sars, 1909 yielded four autapomorphies that unambiguously support the monophyletic state of Breviconia: (1) an elongated and approximately 90°-curved mandibular gnathobase, (2) reduction of the maxillar endopod, (3) maxillar endites carrying 2 instead of 3 setae, and (4) loss of the minute seta on the maxillipedal claw. For B. andrei sp. nov., two autapomorphies could be detected, namely, (1) the development of dorsal tubercles on the P5-bearing body somite and (2) the remarkable elongation of the first endopodal segment of the first swimming leg that is twice as long as the whole exopod. Of particular interest is the presence of a 3-segmented endopod in the third swimming leg of the male of B. andrei sp. nov. It disproves the current assumption that the Ancorabolinae are characterized by (among others) the derived presence of an only 2-segmented endopod in the male’s third swimming leg.
Megapsyrassa Linsley, 1961 is synonymized with Psyrassa Pascoe, 1866 (Coleoptera: Cerambycidae). Aneflomorpha martini Chemsak and Linsley, 1968 is synonymized with Psyrassa sinaloae Linsley, 1935, and the species is transferred to Aneflomorpha Casey, 1912, new combination. Morphological and chromatic variations in Psyrassa cylindricollis Linsley, 1935 are reported, and a new state record is provided. New records are provided for the following species: Psyrassa atkinsoni (Chemsak and Giesbert, 1986) new combination; P. ebenina Linsley, 1935; and P. nigripes Linsley, 1935. Lastly, four new species of Psyrassa Pascoe, 1866 are described: Psyrassa wappesi García and Santos-Silva, from Mexico (Michoacán); P. sonorensis García and Santos-Silva, from Mexico (Sonora); P. obscuriventris García and Santos-Silva, from Mexico (Jalisco); and P. ocularis García and Santos-Silva, from Guatemala (Zacapa).
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Three new species of the family Pyrgodesmidae Silvestri, 1896 from the Udzungwa Mountains are described: Cryptocorypha geminiramus sp. nov., Cryptocorypha cactifer sp. nov., and Cryptocorypha exovo sp. nov. Several additional, unidentifiable pyrgodesmids from the Udzungwa Mountains are recorded. Elythesmus enghoffi Hoffman, 1978 (Cryptodesmidae Karsch, 1880), is recorded from the East Usambara (type locality), West Usambara, Uluguru, Nguru (Kanga) and Udzungwa Mountains, and its variation in size and body ring numbers is analyzed. The surface sculpture of pro- and metazonites is discussed.
Following a review of the circumscription of the genus Cladoceras Bremek. (Rubiaceae) in relation to Tarenna Gaertn., the new species Cladoceras rovumense I.Darbysh., J.E.Burrows & Q.Luke sp. nov. is described from the dry forests of the Rovuma Centre of Plant Endemism (CoE) in southeast Tanzania and northeast Mozambique. This species has previously been known as Tarenna sp. 53, following the revision of African Tarenna by Jérôme Degreef. A comparison to Cladoceras subcapitatum (K.Schum. & K.Krause) Bremek., the only other member of this genus as currently circumscribed, is provided. The new species is assessed as Endangered under the criteria of the IUCN Red List. New records for Mozambique of two further Rovuma CoE endemics are recorded: Celosia patentiloba C.C.Towns. (Amaranthaceae) and Cordia fissistyla Vollesen (Boraginaceae), both of which are globally threatened.
Two new species of shrubs in the genus Lasianthus Jack (Rubiaceae), L. naikii V.S.Dang & Vuong sp. nov. and L. sonlangensis V.S.Dang, Vuong & Quan sp. nov. from Kon Chu Rang Nature Reserve of Gia Lai Province in southern Vietnam, are described and illustrated. The similarities and differences of two new species with the morphologically closest allies are presented. Descriptions, illustrations, colour plates and vernacular names are provided.
The original description of Rivudiva trichobasis Lugo-Ortiz & McCafferty, 1998 is short, with few illustrations. The lack of information resulted in a broad specific diagnosis, with emphasis on the spine-like setae on the antenna. Our hypothesis is that the lack of information resulted in many species being hidden behind R. trichobasis name. We evaluated the species R. coveloae (Traver, 1971) and R. venezuelensis (Traver, 1943) considering the new evidence. After analyzing the paratype of R. trichobasis and records of the species in Brazil, the hidden-species hypothesis was corroborated. Five new species were identified among the published records of R. trichobasis of which four are described here: R. amazona sp. nov. (Roraima State), R. oxum sp. nov. (Rondônia State), R. uiara sp. nov. (Amazonas State), R. naia sp. nov. (Roraima State). The fifth species, Rivudiva sp. X from Maranhão State, could not be described due to the poor conservation of the specimen and is therefore left in open nomenclature. Records from Espírito Santo State (Brazil) and from Paraguay are treated as putative and must be evaluated considering the new evidence. Rivudiva venezuelensis and Rivudiva coveloae are diagnosed and illustrated based on type material. After analyzing these two species, we hypothesize that only R. coveloae likely belongs to the genus Rivudiva. However, information on the nymphal stage is needed to corroborate this hypothesis.
Perinereis anderssoni Kinberg, 1865 originally described from Brazil has been attributed with a wide distribution on the American Atlantic coasts, ranging from Brazil to Bermuda, due to the synonymy of Nereis bairdii Webster, 1884 and Nereis (Perinereis) melanocephala McIntosh, 1885. In this paper, the synonymy of Nereis bairdii with Perinereis anderssoni is rejected based on a re-examination of the syntypes of N. bairdii which are found to contain two species requiring designation of a lectotype and paralectotypes here redescribed as Perinereis bairdii (Webster, 1884) and the remaining specimens are here described as Perinereis websteri sp. nov. The synonymy of Nereis (Perinereis) melanocephala with P. anderssoni is rejected and the synonymy with P. bairdii proposed by previous authors is accepted here. A description of P. anderssoni, a redescription of P. floridana, and a description of the males of P. cariboea with additional material are provided for comparison. A key to identifying all Atlantic Perinereis species is also included.
This paper describes new or little known Amphipoda collected from Hydrozoa, Bryozoa, Porifera or kelp along the Chilean fjord region. The following species have been found and most of them were redescribed and illustrated: Sunamphitoe femorata (Krøyer, 1845); Caprella cf. equilibra Say, 1818; Haplocheira barbimana robusta K.H. Barnard, 1932; Epimeria (Metepimeria) acanthurus (Schellenberg, 1931); Labriphimedia vespuccii K.H. Barnard, 1931; Leucothoe kawesqari Esquete & Aldea, 2015; Podocerus cf. danae (Stebbing, 1888); Ligulodactylus macrocheir (Schellenberg, 1926) and Torometopa cf. crassicornis (Schellenberg, 1931). One species in this contribution is new to science: Liouvillea rocagloria sp. nov.
Oxyethira Eaton, 1873 is one of the most diverse genera of Hydroptilidae, comprising over 240 species distributed in all biogeographical regions. Here three new species of Oxyethira (Trichoglene) Neboiss, 1977 are described and illustrated from male specimens collected in New Caledonia: O. (Trichoglene) hamus sp. nov., recognized by the hook-shaped apex of the long inferior appendages in lateral view and by the posterior margin of segment IX with a trilobed appearance in ventral view; O. (Trichoglene) rectangulata sp. nov., recognized by the rectangular shape of the inferior appendages, which are totally fused and with two pairs of small setae on the inner face; and O. (Trichoglene) spiralis sp. nov., recognized by the strongly curvilinear shape of the subgenital process in dorsal and lateral views and by the long process spiralling around the ejaculatory duct at the phallus apex.
Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs
(2022)
The concept of the previously monogeneric subfamily Pseudocetherinae (Hemiptera: Heteroptera: Reduviidae) is revised and expanded. We here transfer Gerbelius Distant, 1903, Kayanocoris Miller, 1954, Microvarus Jeannel, 1917, Paragerbelius Miller, 1958, and Voconia Stål, 1866 from Reduviinae to Pseudocetherinae and treat Kayanocoris, Microvarus, Paragerbelius, and Pseudocethera Villiers, 1963 as junior synonyms of Voconia, resulting in new combinations for Voconia conradti (Jeannel, 1917) comb. nov., V. ifana (Villiers, 1963) comb. nov., V. monodi (Villiers, 1963) comb. nov., V. motoensis (Schouteden, 1929) comb. nov., V. ornata (Distant, 1903) comb. nov., V. schoutedeni (Villiers, 1964) comb. nov., V. typica (Miller, 1958) comb. nov., and V. wegneri (Miller, 1954) comb. nov. We also describe 23 new species of Voconia: V. bakeri sp. nov., V. bracata sp. nov., V. brachycephala sp. nov., V. chrysoptera sp. nov., V. coronata sp. nov., V. decorata sp. nov., V. dolichocephala sp. nov., V. fasciata sp. nov., V. grandioculata sp. nov., V. hemera sp. nov., V. isosceles sp. nov., V. laosensis sp. nov., V. lasiosoma sp. nov., V. lirophleps sp. nov., V. loki sp. nov., V. mexicana sp. nov., V. minima sp. nov., V. nyx sp. nov., V. smithae sp. nov., V. tridens sp. nov., V. trinidadensis sp. nov., V. tuberculata sp. nov., and V. vittata sp. nov. Lectotypes are designated for Gerbelius confluens Distant, 1903, G. typicus Distant, 1903, V. conradti comb. nov., V. ornata comb. nov., and V. pallidipes Stål, 1866. A revised diagnosis and description of Pseudocetherinae are provided along with photographs of the species and of the male genitalia of 13 pseudocetherine and five closely related reduviine species. An identification key to the two genera of Pseudocetherinae as well as a key to species of Voconia are presented. A phylogenetic hypothesis is proposed for the relationships of Pseudocetherinae using parsimony analyses of 77 morphological characters.
Contrary to what its name suggests, Nemesia arboricola is not strictly arboreal in habit. Here we compare female specimens of N. arboricola collected from arboreal and terrestrial nests. We furthermore describe the male of N. arboricola for the first time as well as two recently discovered species of Nemesia (N. maltensis sp. nov. and N. cominensis sp. nov.). Nemesia maltensis is described from both sexes, N. cominensis is described from the female and a juvenile male specimen. For N. cominensis we discuss the sexual dimorphism of juvenile male and female spiders. Field observations and laboratory observations show remarkable features of the Maltese Nemesia fauna that are unknown from Nemesia species found elsewhere. Particularly, the arboreal dwellings of N. arboricola and the absence of a trapdoor to close off the burrow entrance in N. maltensis appear to be exceptional. The composition of the Maltese Nemesia fauna, as located in the central Mediterranean, is finally discussed in relation to the different Nemesia species-complexes occurring in the eastern and western Mediterranean basin.
New species, synonymy, new records, and taxonomic notes in American Cerambycidae (Coleoptera)
(2022)
Cotyclytus parumnotatus (Zajciw, 1963) (Coleoptera: Cerambycidae: Cerambycinae: Clytini) is revalidated and a neotype is designated. Anatinomma Bates, 1892 (Cerambycinae: Hesperophanini) is reported as a neuter gender and two species-group names are corrected: A. brevicorne Fisher, 1944 and A. insulare Chemsak and Linsley, 1964; the latter is newly recorded from Honduras. Pachymerola vitticollis Bates, 1892 (Cerambycinae: Hyboderini) is recorded for the first time in the Mexican state of Jalisco, and chromatic variation is reported. Smodicum dinellii Bruch, 1911 (Cerambycinae: Smodicini) is recorded from Bolivia; and Corcovado bezarki Martins and Galileo, 2008 (Lamiinae: Hemilophini) is recorded from Belize. Parhippopsis Breuning, 1973 (Lamiinae: Agapanthiini) is synonymized with Rosalba Thomson, 1864 (Lamiinae: Apomecynini) and Rosalba columbiana (Breuning, 1973) is a new combination. Neocompsa flavoquadripunctata Botero and Santos-Silva, new species (Cerambycinae: Neoibidionini), is described from Mexico (Jalisco); Temnopis spiculata Botero and Santos-Silva, new species (Cerambycinae: Oemini) is described from Bolivia (Beni); Trichohippopsis basilaris Botero and Santos-Silva, new species (Lamiinae: Agapanthiini) and Anobrium bicolor Botero and Santos-Silva, new species (Lamiinae: Pteropliini) are described from French Guiana.
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The western Indian Ocean genus Paracephaelis is revised. Sixteen species are endemic to Madagascar, 12 of which are new for science: P. aristata sp. nov., P. bardotiae sp. nov., P. capitulifera sp. nov., P. gautieri sp. nov., P. grandifructa sp. nov., P. longipedicellata sp. nov., P. orientalis sp. nov., P. pauciflora sp. nov., P. ranirisonii sp. nov., P. russata sp. nov., P. sambavensis sp. nov., and P. seyrigii sp. nov. Paracephaelis trichantha occurs on the east coast of continental Africa and on the islands of the Aldabra Group. One other species is newly described from the Comoros, P. comorensis sp. nov. Paracephaelis is characterized by sessile inflorescences, pubescent inflorescence and flower parts, ovules arranged at the periphery of the placenta, laterally flattened seeds with entire endosperm and a shallow elongate hilum, and pollen with supratectal elements. Of the Malagasy endemics, five species are assessed as Critically Endangered with P. sambavensis sp. nov. possibly Extinct in the Wild, three are Endangered, four are Vulnerable, one is Near Threatened, and three are Least Concern. Outside Madagascar, P. trichantha and P. comorensis sp. nov. are assessed as Vulnerable.
The present study aims to resolve the taxonomic confusion involving several taxa within Mycetophagidae Leach, 1815, originating from the introduction of the genus Atritomus Reitter, 1877, and then by its subsequent controversial interpretation. A detailed overview of the taxonomic and nomenclatural history of the taxa previously linked to Atritomus is provided. The authors propose the introduction of Stereophilus Biscaccianti, Audisio & Esser gen. nov. for Atritomus filicornis Reitter, 1887, and the restoration of Entoxylon Ancey, 1869 at the genus rank, together with some rectifications regarding the authorship and the date of publication of both Entoxylon and its type species, E. abeillei Ancey, 1869. Moreover, the Ethiopian species Atritomus vicinus Grouvelle, 1908 is herein transferred to the genus Typhaeola Ganglbauer, 1899 based on the examination of the holotype. The following new combinations are proposed: Entoxylon baudii (Seidlitz, 1889) comb. nov. (from Esarcus Reiche, 1864), Entoxylon besucheti (Dajoz, 1964) comb. nov. (from Esarcus subg. Entoxylon), Entoxylon franzi (Dajoz, 1964) comb. nov. (from Esarcus subg. Entoxylon), Entoxylon inexpectatus (Dajoz, 1964) comb. nov. (from Esarcus subg. Entoxylon), Entoxylon martini (Reitter, 1887) comb. nov. (from Esarcus), Stereophilus filicornis (Reitter, 1887) gen. et comb. nov. (from Atritomus), Typhaeola vicina (Grouvelle, 1908) comb. nov. (from Atritomus).
Chvalaea australis sp. nov. is described and illustrated, representing the first species of Chvalaea Papp & Földvári, 2002 from the Australasian Region. A discussion of the geographic distribution and the possible relationship among the species of the genus is provided.
Two new species of the family Nemouridae of the genus Indonemoura Baumann, 1975, Indonemoura yingjiangensis Bai & Qian sp. nov. and Indonemoura longihamata Bai & Qian sp. nov. are described from Yunnan Province, southwest of China. The morphological characteristics of the two new species are compared to related taxa.
A new species of the genus Caucaseuma Strasser, 1970, Caucaseuma strasseri Antić sp. nov., is described from a cave in the Western Caucasus, representing the eighth species of the genus, and the fourth presumed troglobiontic Caucaseuma. An updated key to and a distribution map of all eight species of Caucaseuma is presented, including new records. In addition, the cave millipede fauna of the Caucasus is briefly discussed, with the inclusion of the most relevant references.
The pseudoscorpion (Arachnida: Pseudoscorpiones) fauna of mainland Ecuador, excluding the Galápagos Islands, is poorly known, with only 41 described species in 9 families. The family Syarinidae has a pantropical distribution and presently comprises ca 120 species in 17 valid genera that are found in leaf litter and subterranean habitats, mostly in tropical and subtropical climates. Four syarinid species have been recorded from Ecuador, including the Galápagos, in two widespread genera, Ideobisium and Ideoblothrus, but field collections suggest that these pseudoscorpions are common and abundant in all forest systems across the country. Here, we review field collections of syarinids from mainland Ecuador and describe five new species in these genera: Ideobisium kichwa sp. nov. (Napo Province, Colonso Chalupas Natural Reserve), I. sonqo sp. nov. (Napo Province, Colonso Chalupas Natural Reserve), I. susanae sp. nov. (Napo Province, Jatun Sacha Natural Reserve), Ideoblothrus nadineae sp. nov. (Napo Province, Colonso Chalupas Natural Reserve) and I. safinai sp. nov. (Pichincha Province, Otongachi Natural Reserve) based on morphology. These species seem to have narrow distributions and we therefore restrict the ranges of two species previously recorded from Ecuador (Ideobisium crassimanum Balzan, 1892 and Ideoblothrus costaricensis (Beier, 1931)) to their countries of origin, which is Costa Rica and Venezuela, respectively.
The genus Seticornuta Morley, 1913 currently comprises nine described species; here, nine new species are described: S. anchanchu sp. nov., S. carinata sp. nov., S. cuckoo sp. nov., S. curupira sp. nov., S. flava sp. nov., S. muqui sp. nov., S. nigroflava sp. nov., S. quilmes sp. nov., and S. rufa sp. nov. The genus is redescribed to encompass the features found in Neotropical species and distinctions between the species in the New World and Old World are presented. The genus is recorded for the first time for Argentina, Bolivia, Ecuador, Guatemala and Peru.
The hoverfly genus Eumerus Meigen, 1822 (Eristalinae: Merodontini) comprises 250+ described species, of which 36 are reported from the Iberian Peninsula. The high species diversity linked to the low degree of morphological differentiation between some species, which is even lower in females, leads to a high taxonomic complexity in this genus. The aim of this work is to confirm the morphological and molecular validity of an undescribed species of Eumerus, which is widespread in the Iberian Peninsula. The new species is described and compared with similar species. The genitalia of the new species are similar to those of Eumerus clavatus Becker, 1923 and Eumerus uncipes Rondani, 1850, but also share some features with Eumerus nudus Loew, 1848. The COI-5’ barcode is provided for the new taxon and analysed together with those of other named Eumerus sequences/species publicly available online. In the light of the morphology and barcoding data, the systematic position of the new species is discussed.