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Three species of Corethrella Coquillett, 1902 from the state of Amazonas, Brazil are described as new to science based on female adult specimens. Corethrella cabocla Feijó, Belchior, Marialva & Pessoa sp. nov. possesses four large setae on the frons between the ventromedial area of ommatidia, a wide clypeus with 1–4 setae, a wing with the apex of R2 basal to the apex of M2 and with a midlength band, and with the abdomen entirely dark brown. Corethrella ielemdei Feijó, Ramires, Lima & Pessoa sp. nov. possesses an elongated coronal suture, four large setae on the frons between the ventromedial area of ommatidia, a clypeus squarish with 42–43 setae, a wing with the apex of R2 basal to the apex of M1 and with a midlength band and dark scales on the basal and subbasal areas of the anterior margin, legs with dark scales, and with the abdomen entirely dark brown. Corethrella menini Feijó, Picelli, Ríos-Velásquez & Pessoa sp. nov. possesses wings with the apex of R2 basal to the apex of M2 and a midlength band, with darker basal scales along all veins, basal band dark scales on C, Sc, R, M, and Cu and the abdomen entirely dark brown. With the addition of the new species, the numbers of frog-biting midges described in the Amazon basin, Brazil and in Neotropical region are now 31, 49 and 80 species, respectively.
Camellia sinensis is one of the major crops grown in Taiwan and has been widely cultivated around the island. Tea leaves are prone to various fungal infections, and leaf spot is considered one of the major diseases in Taiwan tea fields. As part of a survey on fungal species causing leaf spots on tea leaves in Taiwan, 19 fungal strains morphologically similar to the genus Diaporthe were collected. ITS (internal transcribed spacer), tef1-α (translation elongation factor 1-α), tub2 (beta-tubulin), and cal (calmodulin) gene regions were used to construct phylogenetic trees and determine the evolutionary relationships among the collected strains. In total, six Diaporthe species, including one new species, Diaporthe hsinchuensis, were identified as linked with leaf spot of C. sinensis in Taiwan based on both phenotypic characters and phylogeny. These species were further characterized in terms of their pathogenicity, temperature, and pH requirements under laboratory conditions. Diaporthe tulliensis, D. passiflorae, and D. perseae were isolated from C. sinensis for the first time. Furthermore, pathogenicity tests revealed that, with wound inoculation, only D. hongkongensis was pathogenic on tea leaves. This investigation delivers the first assessment of Diaporthe taxa related to leaf spots on tea in Taiwan.
Two common sigalionid species from the Northeast Atlantic, Sthenelais boa (Johnston, 1833) and S. limicola (Ehlers, 1864), have never been revised in detail. Although their validity has never been questioned, a number of taxonomic problems related to Sthenelais Kinberg, 1856 and the later established Fimbriosthenelais Pettibone, 1971 remain unresolved. The validity of F. minor (Pruvot & Racovitza, 1895) has been repeatedly discussed, but no agreement reached. Also the validity of Fimbriosthenelais has been at stake, affecting the generic assignment of Fimbriosthenelais zetlandica (McIntosh, 1876), another species present in the area. Among the investigated species of Sthenelais, some where thought to be synonyms of Eusthenelais hibernica McIntosh, 1876, which led us to also include Eusthenelais McIntosh, 1876. We also re-examined Eusthenelais abyssicola McIntosh, 1879, the only other species attributed to the genus, and confirm that it is indeterminable. In total, we investigated 37 nominal taxa reported from the Northeast Atlantic and as a result we consider only five species to be valid: Sthenelais boa, S. limicola, Fimbriosthenelais zetlandica, F. longipinnis (Grube, 1869) and Eusthenelais hibernica. These genera and species are described and discussed herein and an updated identification key to all Northeast Atlantic species of Sigalioninae Gonzalez et al., 2018 is given.
The classification of the genera belonging to the doryctine tribe Rhaconotini (Braconidae) is updated. The following new taxa are described: Troporhaconotus gen. nov. (with 12 species), Afroipodoryctes subgen. nov. (of Ipodoryctes Granger, 1949) (with three species), Hexarhaconotinus subgen. nov. (of Rhaconotinus Hedqvist, 1965) (with ten species), Bathycentor zimbabwensis sp. nov., Ipodoryctes (Afroipodoryctes) reunionus sp. nov., I. (A.) saintphilippensis sp. nov., Platyspathius (Platyspathius) venezuelicus sp. nov., P. (P.) ranomafanus sp. nov., Rhacontsira haeselbarthi sp. nov., Rh. mozambiquensis sp. nov., Rh. saigonensis sp. nov. and Rh. toamasina sp. nov. The generic status of Euryphrymnus Cameron, 1910 is resurrected. The generic name Aptenobracon Marsh, 1965 is synonymised under Rhaconotus Ruthe, 1854 (syn. nov.); Rhaconotus asiaticus Belokobylskij, 1990 is synonymised under Rh. kerzhneri Belokobylskij, 1985 (syn. nov.). The new name, Rhaconotinus austrochinensis nom. nov., is suggested for the preoccupied name Rhaconotus chinensis Chen & Shi, 2004 (December) not Rhaconotus chinensis Belokobylskij & Chen, 2004 (June). New species contents, in many cases with numerous new combinations, are suggested for the genera Bathycentor Saussure, 1892, Euryphrymnus Cameron, 1910, Ipodoryctes Granger, 1949, Rhaconotinus Hedqvist, 1965 and Rhaconotus. The tribe Leptorhaconotini is synonymised with Rhaconotini based on previously published molecular phylogenetic studies, though we leave this group within the subtribe Leptorhaconotina.
We here describe a new Indian helicarionoidean genus, Varadia Bhosale & Raheem gen. nov., containing the single species Varadia amboliensis Bhosale, Thackeray, Muley & Raheem gen. et sp. nov. This new semi-slug is endemic to the northern and central Western Ghats and is primarily a forest-living species. We describe and figure the shell, reproductive system, radula, spermatophore and external morphology of this new species, and detail its known distribution. We explore its relationships to other helicarionoideans using phylogenetic analyses of DNA sequence data for part of the ribosomal RNA gene cluster and discuss the morphology of the new genus in relation to other, primarily South Indian, helicarionoidean taxa. Based on characters of the reproductive system, particularly the male genitalia and the gametolytic sac, we provisionally place Varadia gen. nov. in the Macrochlamydinae (Ariophantidae). This is consistent with the results of our molecular phylogenetic analyses. The combination of large size, broad, densely tuberculated shell lobes and a shell with ca 4 whorls and a disproportionately large body whorl makes V. amboliensis gen. et sp. nov. unique among the helicarionoidean taxa of the Western Ghats. The new semi-slug is also highly distinctive in the morphology of its male genitalia.
Dual-task paradigms encompass a broad range of approaches to measure cognitive load in instructional settings. As a common characteristic, an additional task is implemented alongside a learning task to capture the individual’s unengaged cognitive capacities during the learning process. Measures to determine these capacities are, for instance, reaction times and interval errors on the additional task, while the performance on the learning task is to be maintained. Opposite to retrospectively applied subjective ratings, the continuous assessment within a dual-task paradigm allows to simultaneously monitor changes in the performance related to previously defined tasks. Following the Cognitive Load Theory, these changes in performance correspond to cognitive changes related to the establishment of permanently existing knowledge structures. Yet the current state of research indicates a clear lack of standardization of dual-task paradigms over study settings and task procedures. Typically, dual-task designs are adapted uniquely for each study, albeit with some similarities across different settings and task procedures. These similarities range from the type of modality to the frequency used for the additional task. This results in a lack of validity and comparability between studies due to arbitrarily chosen patterns of frequency without a sound scientific base, potentially confounding variables, or undecided adaptation potentials for future studies. In this paper, the lack of validity and comparability between dual-task settings will be presented, the current taxonomies compared and the future steps for a better standardization and implementation discussed.
Handaoia Seyrig, 1952 is a small genus of Phygadeuontinae currently represented by eleven described species from Madagascar, Tanzania and Europe, and can be recognized by the combination of the distally expanded and ventrally flattened antennal flagellum, complete posterior transverse carina of the mesosternum, isolated ‘pit’ (episternal scrobe) in the mesopleuron, and a single bulla in fore wing vein 2m-cu. Most species have a distinctive combined area basalis and area superomedia on the propodeum. The following six new species from Central and South America are described and illustrated: H. cuscoensis Bordera sp. nov. from Peru, H. fritzi sp. nov. from Brazil, H. mercedensis Bordera sp. nov. from Peru, H. plaumanni sp. nov. from Brazil, H. ruizcancinoi Bordera sp. nov. from Mexico, and H. urceus sp. nov. from Brazil. A key to the New World species is provided.
The genus Oplognathus MacLeay, 1819 is revised based on type material of two of the three described species and scattered additional material from several collections around the world. The diagnostic characters of the genus are confirmed, distinguishing it from other Brazilian Areodina mainly by: quadrangular clypeus with trilobate apex in males, rounded in females, extending beyond labrum in both sexes; mandibles with three distinct teeth; maxillae with six teeth; antenna with 10 antennomeres; 10 elytral striae; mesoventral process present; and asymmetrical parameres. The genus and all three species are redescribed, and the female of Oplognathus bahianus Ohaus, 1912 is described for the first time. We consider Oplognathus helmenreichi var. maculicollis Ohaus, 1914 an unavailable infrasubspecific taxon that is conspecific with Oplognathus helmenreichi Ohaus, 1905; its distribution is updated, and the different spelling of the specific epithet is discussed. A neotype is designated for Oplognathus kirbii MacLeay, 1819 since the holotype is currently considered lost. Additionally, an identification key and a distribution map are included.
The enigmatic millipede assassin bug genus Xenorhyncocoris Miller, 1938 is revised. Previously known species, X. caraboides Miller, 1938, X. princeps Miller, 1949 and X. schoenitzeri Putshkov & Bérenger, 1999, are diagnosed and photographed. A new species, X. attractivus sp. nov., is described based on male and female specimens from northeastern Borneo. The male of Xenorhyncocoris is reported for the first time, revealing the extreme sexual dimorphism present in the genus. The diagnosis of Xenorhyncocoris is extended in order to make it applicable to the new discovery, and a female-based key to species of the genus is updated. Relationships among Xenorhyncocoris and Vilius Stål, 1863, Neozirta Distant, 1919 and Schottus Distant, 1902 are briefly discussed.
In the present paper, we have made a taxonomic revision of the previously monotypic genus Troglostygnopsis Šilhavý, 1974. Based on the revision of diverse material, the genus is rediagnosed and the type species, Troglostygnopsis anophthalma Šilhavý, 1974 is redescribed. The new species Troglostygnopsis kalebi sp. nov. is described from specimens collected in a cave in Chiapas, Mexico, and this species is compared with T. anophthalma. A final discussion on some troglobitic genera of Stygnopsidae is addressed.
The Smicridea (Smicridea) fasciatella species group occurs from the southwestern USA, throughout Central America, the Greater Antilles islands, and most of South America, except for the Chilean subregion. It is characterized by the phallic apparatus being a simple tube with eversible internal sclerites at the apex. The fasciatella group is composed of 61 species, of which only 11 occur in Brazil, mainly in the Atlantic Forest biome in the southeastern region. In order to reduce the Linnean and Wallacean shortfalls for the Smicridea Brazilian fauna, we diagnose, describe, and illustrate males of six new species in the fasciatella group: Smicridea (Smicridea) blahniki Desiderio, Pes & Hamada sp. nov., S. (Smicridea) brevitruncata Desiderio, Pes & Hamada sp. nov., S. (Smicridea) caaguara Desiderio, Pes & Hamada sp. nov., S. (Smicridea) ipiranga Desiderio, Pes & Hamada sp. nov., S. (Smicridea) jeaneae Desiderio, Pes & Hamada sp. nov., and S. (Smicridea) polyacantha Desiderio, Pes &; Hamada sp. nov. Additionally, we provide distributional data for S. (Smicridea) albosignata Ulmer, 1907, S. (Smicridea) bivittata (Hagen, 1861), S. (Smicridea) erecta Flint, 1974, S. (Smicridea) obliqua Flint, 1974, S. (Smicridea) paranensis Flint, 1983, and S. (Smicridea) sattleri Denning & Sykora, 1968. The number of S. (Smicridea) species in Brazil increases from 21 to 27 and Smicridea is recorded from the states of Acre, Amapá, and Sergipe for the first time.
Comanopa Blocker, 1979 and Gehundra Blocker, 1976 are small and poorly known genera of leafhoppers, previously comprising four and five species, respectively. In this study, two new species are proposed: Comanopa mananciensis sp. nov. from the state of Paraná, Southern Brazil, and Gehundra cristalinensis sp. nov. from the state of Mato Grosso, Centralwestern Brazil. The male of Gehundra sordida (Baker, 1900) is studied by the first time. Detailed descriptions and illustrations of males and females are provided and keys to males of Comanopa and Gehundra are given.
The genus Stenaspis Audinet-Serville, 1834 (Coleoptera: Cerambycidae: Cerambycinae: Trachyderini) is recharacterized by providing additional morphological features to enhance the definition of this genus. Stenaspis plagiata Waterhouse, 1877 is transferred to Crioprosopus Audinet-Serville, 1834, new combination, and the latest key to species of Crioprosopus is modified. A key to the genera of Group III-Stenaspes with “abruptly separated anteocular space,” and a key to species of Stenaspis is provided. Color illustrations of the available species are included. Stenaspis lingafelteri Eya, new species, from Mexico is described.
This paper offers an explanation of each of the 44 scientific names given by Leopold Krüger (1861-1942) to odonate taxa together with that for the names of all the genera into which they are sorted now. But prior to that there is some information about the life and work of this scientist, and in the final part his preferences in odonatological nomenclature are compared with those in the names created by F.M. Brauer and F. Ris and some impressions of his studies on Neuroptera are presented and considerations about his aspirations in his work are given.
Fractipes tayrona new species (Coleoptera: Chrysomelidae: Eumolpinae: Eumolpini) is described from a dry forest on the Caribbean coast of Colombia. This locality is very distant from the localities in southeast Brazil, where the other two known species of Fractipes Bechynĕ were described. Male and female genitalia are described for the first time in this genus, and F. tayrona is compared with holotype photographs of F. rhabdopteroides Bechynĕ and F. secundus Bechynĕ and Springlová de Bechynĕ.
A new species of verrucid barnacle, Gibbosaverruca weijiai sp. nov., is described and illustrated based on specimens collected from a deep-sea seamount, Weijia Guyot, in the West Pacific. This is the third barnacle species reported from Weijia Guyot. The new species is distinguished from its congeners by its extremely long caudal appendage. In terms of shell morphology, G. weijiai sp. nov. is most similar to G. gibbosa and G. sulcata but differs from them by the intermediate articular ridges of its movable tergum and scutum, which are distinctly wider than the axial ridges. The COI and 16S rRNA sequences of the holotype of G. weijiai sp. nov. are presented to support future research, and a key to extant species of the genus Gibbosaverruca Young, 2002 is provided.
The genus Miridiba Reitter, 1902, of phytophagous chafers from Asia, is revised based on external morphological and genital (male and female) characters. In this study, a total fifty-eight species of Miridiba were examined, and the genus is redescribed. Male genital characters of Miridiba, especially the morphology of parameres and endophallus, are studied in depth herein. The female genitalia of Miridiba are studied and described in detail for the first time. According to genital characters, nine genital morphotypes, including fifty-four species, are established under the genus Miridiba. Sixty-seven type specimens are studied. Miridiba gressitti (Frey, 1970) comb. nov., Miridiba borneensis (Moser, 1918) comb. nov., Miridiba coxalis (Arrow, 1944) comb. nov., Miridiba rugaticollis (Moser, 1913) comb. nov., Miridiba nigrescens (Moser, 1916) comb. nov., Miridiba scutata (Reitter, 1902) comb. nov., Miridiba ciliatipennis (Moser, 1903) comb. nov. and Miridiba brunneipennis (Moser, 1916) comb. nov. are transferred from Holotrichia. Holotrichia dalatensis Frey, 1970 is transferred to Miridiba as a synonym of Miridiba sinensis (Hope, 1842). Four junior subjective synonyms are proposed: Miridiba pilosella (Moser, 1908) (= Holotrichia formosana Moser, 1909), Miridiba sinensis (= Holotrichia dalatensis Frey, 1970), Miridiba scutata (= Holotrichia scutulata Dalla Torre, 1912 and Miridiba lassallei Keith, 2010). Miridiba frontalis (Fairmaire, 1886) is resurrected as a valid species. A key to 51 of the 58 valid species of Miridiba is presented.
Ysachron pilosus gen. et sp. nov. is described from the Caribbean region of Colombia, and an updated key to genera of South American Achrysonini is provided. Moreover, the geographical distribution of 45 species of Cerambycidae, belonging to the subfamilies Cerambycinae, Lamiinae and Prioninae, is expanded, including nine new country records for Colombia and a further locality in Colombia for nine species.
Se describen para las Antillas cuatro especies nuevas de Lasioglossum Curtis pertenecientes al taxon Habralictellus Moure y Hurd (Hymenoptera: Halictidae): Lasioglossum (Habralictellus) cuba, Lasioglossum (Habralictellus) darlingtoni, Lasioglossum (Habralictellus) jagibbsi y Lasioglossum (Habralictellus) laupackeri. Se utiliza como subgénero para mantener la estabilidad taxonómica. Se presenta la historia y el estado actual del conocimiento de Habralictellus, que aún no ha encontrado un lugar justificado y aceptado, al unísono por los especialistas, en la nomenclatura zoológica. Se ofrece un listado de las especies conocidas, sexo y distribución.
Susamyia Ziegler & Gilasian gen. nov. and its type species Susamyia mira Ziegler & Gilasian gen. et sp. nov. are described from southwestern Karkheh National Park and southeastern Jazmourian Wetland of Iran. Drawings of terminalia and head of the male as well as images of habitus, head and egg are provided. Susamyia gen. nov. is incorporated into the identification key to the Palearctic tachinid genera of Tschorsnig & Richter (1998). The systematic position of Susamyia gen. nov. in the tribe Goniini (Exoristinae) is discussed in light of molecular analyses based on the ribosomal genes 16S, 18S and 28S, and the egg type of females. The morphology of the microtype eggs was studied and compared with that of other Goniini, but proved too unreliable to be included in the phylogenetic analysis. Susamyia gen. nov. is believed to be most closely related to the goniine genera Gymnophryxe Villeneuve, 1922, Palesisa Villeneuve, 1929 and Prosopodopsis Townsend, 1926.
Despite their importance as potential biological control agents, species of the campoplegine genus Campoplex Gravenhorst, 1829 are hard to identify. Previous works provided short descriptions or poor illustrations of crucial characters, meaning it is often impossible to distinguish closely related species. We provide illustrations to identify species of the Campoplex difformis group and redescriptions of and illustrations for C. difformis (Gmelin, 1790), C. capitator Aubert, 1960, C. dubitator Horstmann, 1985, C. formosanae Horstmann, 2012, and C. unicingulatus (Schmiedeknecht, 1909). In addition, the taxonomic status of C. difformis is clarified; a lectotype is designated for C. capitator in Aubert’s collection in Lausanne and the host record for this species on Ancylis mitterbacheriana (Denis & Schiffermüller, 1775) is queried; Campoplex corsicator Aubert, 1960 stat. nov. is removed from synonymy with Campoplex tibialis (Szépligeti, 1916) and redescribed.
An illustrated checklist of the Mollusca of Galicia Bank, a large and deep seamount off the NW Iberian Peninsula, is provided. The studied material was collected in 8 samples of Seamount 1 cruise (1987), 7 samples of ECOMARG 0709 (2009) and 36 samples of BANGAL 0711 (2011), between 615 and 1768 m. A total of 212 species are known to occur at the Galicia Bank (1 Monoplacophora, 7 Solenogastres, 3 Polyplacophora, 132 Gastropoda, 54 Bivalvia, 6 Scaphopoda, and 9 Cephalopoda), 21 of which from previous studies only. Four species are described as new, 34 species are first record in Spanish waters and another 20 species first record for the Northern Spanish waters. Over 7500 specimens, representing 104 species, were collected alive, and 87 species were represented by empty shells only. Only 53 species were detected in both Seamount 1 and BANGAL 0711; most of the species are rare and more species can be expected if exploration is continued. There is a marked difference in species composition between the summit platform (615‒1000 m) and the deeper part below 1500 m, with some genera (e.g., Colus and Limopsis) represented by alternative species. Endemism, if any, is very low and most of the species are widespread.
Ophiuroidea is the largest class among extant echinoderms, with over 2000 described species assigned to 33 families. Here, the first identification key to the recently revised classification was developed, and revised morphological descriptions were derived from it, expanding the previous short diagnoses. The key was built by analyzing internal and external skeletal characters of predominantly the type species of each family, including at least two mutually exclusive attributes per family. Various numeric and multistate characters were used to create a traditional as well as an interactive key using the DELTA and Xper software programs. Illustrations (SEM and digital photos) are included in the key to facilitate the assessment of character states by users. Not only is it the first identification key to the families, according to the recently proposed new classification and the examined species, but this interactive key also assists users in understanding the family level taxonomy of brittle stars. The interactive key allows new characters and states to be added, when more species will have been analyzed, without the need to reconfigure the complete key (as may be necessary with conventional keys).
The planthopper genus Augilina Melichar, 1914, is recorded in China for the first time. Two new species of the genus Augilina, A. tetraina Chen & Gong sp. nov. and A. triaina Chen & Gong sp. nov., are described and illustrated from South China. The genus now has a total of four described species. New generic characteristics are proposed and photographs of the new species are provided. A checklist and a key to the species of Augilina are also included.
A new Brazilian leafhopper genus of the subfamily Neocoelidiinae is proposed: Takiyaella gen. nov. based on T. sexguttata (Chiamolera & Cavichioli, 2003) gen. et comb. nov. and five new species: T. anomala gen. et sp. nov., T. coelhomarquesae gen. et sp. nov., and T. daniela gen. et sp. nov. from the state of Rio de Janeiro; T. cavichiolii gen. et sp. nov. from the state of Paraná, and T. mejdalanii gen. et sp. nov. from the state of Minas Gerais. A distribution map and identification key to Takiyaella species are provided.
Three new South American leafhopper genera of the tribe Gyponini are proposed: Beltrana gen. nov. based on Beltrana reticulata gen. et sp. nov. from French Guiana, Fulana gen. nov. based on Fulana brasiliensis gen. et sp. nov. from Brazil, and Sicrana gen. nov. based on Sicrana plana gen. et sp. nov. from Brazil and Ecuador. Diagnoses, detailed descriptions, and illustrations are provided for each taxon, as well as comparisons with closely related genera. In addition, the following synonyms are proposed: Freytagana DeLong, 1975 as a junior synonym of Marganana DeLong, 1948 and F. gibsoni DeLong, 1975 as a junior synonym of M. (Marganana) mexicana DeLong & Freytag, 1963. Chilella DeLong & Freytag, 1967 is transferred from Gyponini to Selenomorphini Evans, 1974.
The taxonomy of Micronectidae in Vietnam is reviewed. Based on our cumulative collections during 2001–2020, 11 new species of Micronecta Kirkaldy, 1897 are documented and described in this study: M. acuminata sp. nov., M. arcuata sp. nov., M. caperata sp. nov., M. clavata sp. nov., M. cultellata sp. nov., M. fulvopicta sp. nov., M. nieseri sp. nov., M. pingae sp. nov., M. sinuata sp. nov., M. undulata sp. nov., and M. vietnamica sp. nov. Nine species are recorded from Vietnam for the first time: M. decorata Lundblad, 1933, M. drepani Nieser, 2000, M. erythra Nieser, Chen & Yang, 2005, M. fugitans Breddin, 1905, M. johorensis Fernando, 1964, M. melanochroa Nieser, Chen & Yang, 2005, M. ornitheia Nieser, Chen & Yang, 2005, M. sahlbergi (Jakovlev, 1881), and M. tuwanoni Nieser, Chen, Leksawasdi, Thanyakam & Duangsupa, 2004. Thus, in this study, two genera and 37 species of Micronectidae are now reported from Vietnam: Synaptonecta Lundblad, 1933 with only one species, S. issa (Distant, 1910), and Micronecta with 36 species. A key to all species occurring in Vietnam, subgeneric diagnoses, and a key to all current subgenera of Micronecta are provided.
The genus Stephostethus LeConte, 1878 of minute brown scavenger beetles (Coleoptera: Latridiidae: Latridiinae) is recorded from Taiwan for the first time, which extends the southernmost boundary of this genus in the Palaearctic realm. Two new species, S. taiwanus sp. nov. and S. yuanfengensis sp. nov., and one new record, S. chinensis (Reitter, 1877), are described in this study. A taxonomic key to the three species of this genus in Taiwan is also provided. The distributions, habitats and collecting methods of these three species, as well as the structure of the genital segments of this genus are discussed.
Grasstrees (Xanthorrhoea) are a group of monocotyledonous plants endemic to Australia, with a characteristic crown of long and narrow leaves. In the present study, the grasstree-associated weevils of the genus Paratranes Zimmerman, 1994 are reviewed and two species are recognised: P. monopticus (Pascoe, 1870) and P. zimmermani sp. nov. A lectotype for Tranes xanthorrhoeae Lea, 1898, which was recently synonymised with P. monopticus, is herein designated, and the validity of this synonymy is confirmed. Descriptions and diagnoses of the species are supplemented with photographs of their habitus and salient structures as well as a map of their distributions.
We describe and illustrate a new Neotropical predaceous midge, Parabezzia carlae Huerta, Spinelli and Grogan, new species (Diptera: Ceratopogonidae) from males collected by Malaise trap in La Union, Ayutla de los Libres, state of Guerrero, Mexico. We also report a second record of P. alexanderi Wirth from Veracruz, Mexico. A key to the known species in Mexico is included.
A new spider genus (Araneae: Linyphiidae: Erigoninae) from a tropical montane cloud forest of Mexico
(2021)
A new genus and species of spider (Araneae, Linyphiidae, Erigoninae) from a tropical montane cloud forest of Mexico is described from both male and female specimens, Xim trenzado gen. et sp. nov. A phylogenetic parsimony analysis situates Xim gen. nov. as a distinct genus among the distal Erigoninae. Xim gen. nov. is sister to a clade including Ceratinopsis, Tutaibo and Sphecozone, but differs from those genera by having a high cymbium, large paracymbium, short straight embolus, male cheliceral stridulatory striae widely and evenly spaced, both sexes with a post-ocular lobe, male with two series of prolateral macrosetae on femur I, and the female by having strongly oblong, u-shaped spermathecae.
Acroterius gen. nov. (Araneae: Linyphiidae: Linyphiinae) with twelve new species from Yunnan, China
(2021)
Acroterius gen. nov. is described from Yuannan, together with twelve new species: A. absentus gen. et sp. nov. (♂); A. brevis gen. et sp. nov. (♂/♀); A. camur gen. et sp. nov. (♂/♀); A. circinatus gen. et sp. nov. (♂/♀); A. hamatus gen. et sp. nov. (♀); A. inversus gen. et sp. nov. (♀); A. latus gen. et sp. nov. (♂/♀); A. longidentatus gen. et sp. nov. (♀); A. longimultus gen. et sp. nov. (♂/♀); A. longiprojectus gen. et sp. nov. (♀); A. ovatus gen. et sp. nov. (♀), and A. parvus gen. et sp. nov. (♀). Detailed descriptions of genitalic characters and somatic features by means of photographs and line drawings, as well as comparisons with closely related species, are provided.
For each of the 534 species of the millipede order Chordeumatida known from Europe, available information on taxonomy, distribution and habitat is summarized, and the distribution in 50 × 50 km UTM/MGRS squares is shown on a map. Comparisons between Chordeumatida and the equally-sized order Julida are made with respect to distribution patterns and history of exploration.
Two subspecies of Cosmodela duponti (Dejean) (Coleoptera: Cicindelidae) are elevated to species rank: Cosmodela barmanica (Gestro) and C. indica (Fleutiaux). The lectotypes of all the above-mentioned species are designated as well. Short redescriptions of the three species are provided together with a key, and illustrations of their habitus.
The pharciceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea)
(2021)
The ammonoids of the suborder Pharciceratina from the Red Ironstone Formation of the area around Dillenburg (eastern Rhenish Mountains) are revised, mainly based on historical collections stored in the Museum für Naturkunde, Berlin. The genus Evopharciceras gen. nov. and the following species are newly described: Maenioceras ornatum sp. nov., Pharciceras beyrichi sp. nov., Pharciceras kruegeri sp. nov., Pharciceras ferrum sp. nov., Evopharciceras formosum gen. et sp. nov., Extropharciceras metallicum sp. nov., Lunupharciceras kochi sp. nov., Stenopharciceras lotzi sp. nov., Pluripharciceras ahlburgi sp. nov. and Sandbergeroceras archiaci sp. nov. Neotypes are proposed for the species Extropharciceras becheri (von Buch, 1832) and Sandbergeroceras costatum (d’Archiac & de Verneuil, 1842). The stratigraphic distribution of the genera is discussed; they are assigned to three assemblages: (1) Maenioceras terebratum Zone (early Givetian; two species), (2) Pseudoprobeloceras pernai Zone (latest Givetian; fifteen species) and (3) Sandbergeroceras costatum Zone (early Frasnian; three species).
The tribe Platynini (Coleoptera: Carabidae: Harpalinae) is revised for New Zealand. Eight genera and forty-three species are recognized. Four genera and sixteen species are described as new: Ctenognathus davidsoni Larochelle and Larivière new species, Ctenognathus earlyi Larochelle and Larivière new species, Ctenognathus garnerae Larochelle and Larivière new species, Ctenognathus hoarei Larochelle and Larivière new species, Ctenognathus kaikoura Larochelle and Larivière new species, Ctenognathus marieclaudiae Larochelle new species, Ctenognathus perumalae Larochelle and Larivière new species, Ctenognathus takahe Larochelle and Larivière new species, Ctenognathus tawanui Larochelle and Larivière new species, Ctenognathus tepaki Larochelle and Larivière new species, Ctenognathus urewera Larochelle and Larivière new species, Kiwiplatynus Larochelle and Larivière new genus, Kiwiplatynus taranaki Larochelle and Larivière new species, Kupeplatynus Larochelle and Larivière new genus, Maoriplatynus Larochelle and Larivière new genus, Maoriplatynus marrisi Larochelle and Larivière new species, Prosphodrus mangamuka Larochelle and Larivière new species, Prosphodrus sirvidi Larochelle and Larivière new species, Prosphodrus waimana Larochelle and Larivière new species, Tuiplatynus Larochelle and Larivière new genus. Lectotypes are designated for twelve taxa: Anchomenus adamsi Broun, 1886, Anchomenus colensonis White, 1846, Anchomenus feredayi Bates, 1874, Anchomenus helmsi Sharp, 1881, Anchomenus intermedius Broun, 1908, Anchomenus macrocoelis Broun, 1908, Anchomenus munroi Broun, 1893, Anchomenus sophronitis Broun, 1908, Colpodes crenatus Chaudoir, 1878, Colpodes neozelandicus Chaudoir, 1878, Ctenognathus littorellus Broun, 1908, and Ctenognathus pictonensis Sharp, 1886.Six new combinations are established: Ctenognathus bidens (Chaudoir, 1878) = Kiwiplatynus bidens (Chaudoir, 1878); Ctenognathus crenatus (Chaudoir, 1878) = Kupeplatynus crenatus (Chaudoir, 1878); Ctenognathus lucifugus (Broun, 1886) = Kupeplatynus lucifugus (Broun, 1886); Ctenognathus sulcitarsis (Broun, 1880) = Kupeplatynus sulcitarsis (Broun, 1880); Ctenognathus libitus (Broun, 1914) = Tuiplatynus libitus (Broun, 1914); Ctenognathus sophronitis (Broun, 1908) = Tuiplatynus sophronitis (Broun, 1908).Fifteen new synonymies are established: Ctenognathus littorellus Broun, 1908 = Ctenognathus adamsi(Broun, 1886); Anchomenus parabilis Broun, 1880 = Ctenognathus cardiophorus (Chaudoir, 1878); Anchomenus integratus Broun, 1908 = Ctenognathus colensonis (White, 1846); Anchomenus macrocoelis Broun, 1908 = Ctenognathus edwardsii (Bates, 1874); Ctenognathus actochares Broun, 1894 = Ctenognathus elevatus(White, 1846); Anchomenus punctulatus Broun, 1877, Anchomenus montivagus Broun, 1880, Anchomenus perrugithorax Broun, 1880, Anchomenus politulus Broun, 1880, Anchomenus suborbithorax Broun, 1880, and Colpodes neozelandicus Chaudoir, 1878 = Ctenognathus novaezelandiae (Fairmaire, 1843); Ctenognathus simmondsi Broun, 1912 = Ctenognathus pictonensis Sharp, 1886; Anchomenus (Platynus) cheesemani Broun, 1880 and Calathus deformipes Broun, 1880 = Kupeplatynus crenatus (Chaudoir, 1878); Anchomenus munroi Broun, 1893 = Kupeplatynus lucifugus (Broun, 1886). Ctenognathus elevatus (White, 1846), previously synonymized with Ctenognathus novaezelandiae (Fairmaire, 1843), is reinstated as full species. A revision of all taxa is provided. Descriptions, identification keys, illustrations of male genitalia, habitus photos, distributional data and maps are given. Extensive information on ecology, biology, dispersal power, and collecting techniques is included for each species.
This paper is the conclusion of the “Spaghetti Project” aiming to revise French species of Terebellidae sensu lato (s.l.) belonging to the five families: Polycirridae, Telothelepodidae, Terebellidae sensu stricto (s.s.), Thelepodidae and Trichobranchidae. During this project, 41 species were observed, 31 of them new for science: eight species of Polycirridae, eleven species of Terebellidae s.s., three species of Thelepodidae and nine species of Trichobranchidae. We provide a comprehensive key for all European species of terebellids with a focus on the important diagnostic characters for each family. Finally, we discuss issues on taxonomy, biodiversity and cryptic and pseudo-cryptic species of polychaetes in European waters, based on results obtained during this project.
In an elevational gradient, the mountain top generally presents a reduced species diversity. However, it is there where we often find microendemic and quite often still undescribed species. That prediction is very common in underexplored Neotropical mountains, like those of the Caparaó National Park – a protected area that includes the highest peak of the Atlantic Forest, a megadiverse domain. Up in its top, we found a dwarf frog of the genus Physalaemus (Anura, Leptodactylidae, Leiuperinae), belonging to the P. signifer clade. After an integrative (morphological, bioacoustical, and genetic) analysis, we were able to describe it as a new species and found it to be sister to P. maculiventris. Due to its very restricted distribution at a site with extreme environmental conditions (which includes fires and frosts) and current instability in national environmental policy, we suggest this to be classified as an endangered species. A brief description of its natural history and the description of the species itself will now enable its proper conservation status categorization and the future planning for conservation actions.
Thirty-four species of dacine fruit flies (Diptera: Tephritidae) were recorded in Bangladesh, based on field surveys carried out between 2013 and 2020. Five species are reported in Bangladesh for the first time: Bactrocera aethriobasis (Hardy), B. limbifera (Bezzi), B. melania (Hardy and Adachi), B. nigrifemorata Li and Wang, and Dacus jacobi David and Sachin. The attraction of B. nigrifemorata to cue-lure is a new male lure association. An annotated checklist and illustrated key to the species are provided.
The planthopper genus Arcofaciella Fennah, 1956 (Hemiptera: Fulgoromorpha: Delphacidae) is reviewed. Four species are recognized, of which A. obflexa Guo & Liang, 2005 and A. verrucosa Fennah, 1956 are redescribed, and one new species, A. indiana sp. nov., is described. Habitus photos for adults and illustrations of male genitalia (excluding A. penangensis (Muir, 1919)) are given. A key for identifying the species of Arcofaciella is also provided.
The genus Onychelmis Hinton, 1941 was for a long time regarded as a small taxon with only three known species distributed in the Andes. A study of new material from Ecuador, using morphological and molecular data, has resulted in the discovery of five new species: Onychelmis lenkae sp. nov., O. lobata sp. nov., O. minor sp. nov., O. onorei sp. nov. and O. splendida sp. nov. We also revised the entire genus and redescribed the three known species, O. longicollis (Sharp, 1882), O. leleupi Delève, 1968 and O. whiteheadi Spangler & Santiago, 1991. Habitus photographs of adults are provided, together with line drawings of male and female genitalia, and schematic illustrations of the distribution of femoral tomentum for each species. DNA sequences for barcoding the COI mtDNA fragment were used to support species delimitation and to suggest possible relationships among species. The revision includes a key to adults of all species of Onychelmis and notes on the biogeography of the genus, with an updated distribution map.
Three new species of Mesacanthion Filipjev, 1927 were found along Patagonian coasts (Argentina). Mesacanthion bifidum sp. nov. is characterized by short labial and cephalic setae, onchia of equal size, spicule arcuate, and gubernaculum with caudal apophysis, ending in two teeth. The species is related to M. virile (Ditlevsen, 1930) De Coninck & Schuurmans Stekhoven, 1933. However, the spicules and gubernaculum of both species are different in shape. Mesacanthion longigubernaculum sp. nov. is characterized by its long and slender body, striated cuticle, relatively long cephalic and cervical setae, onchia of different sizes, amphidial fovea lentil-shaped, spicule arcuate, gubernaculum surrounding the spicule, and tail conical-cylindrical with terminal setae. Mesacanthion sanantoniensis sp. nov. is characterized by its long and stout body, striated cuticle, long cephalic setae, onchia of different sizes, amphidial fovea pouch-shaped, spicule arcuate, gubernaculum with dorsal apophysis, and tail conical without terminal setae. Following the key of Jeong et al. (2019), the last two species are related to M. pali Wieser, 1959 and M. longissimesetosum Wieser, 1953, so we provide a key to differentiate the four species.
Two new species of the genus Eponisiella Emeljanov, 1984 are described and illustrated from China (Guizhou and Shandong Provinces). These are E. dafangensis sp. nov. and E. shandongensis sp. nov., giving the genus eight species in total. A key to the species of Eponisiella is provided as well as a map of their geographic distribution, which is briefly discussed.
A recent phylogenetic analysis has shown that the genus Stephanopis comprises several different lineages of bark-dweller crab spiders. The ones with South American distribution that present a close relationship with other Neotropical genera like Epicadus, Onocolus, Rejanellus and Epicadinus were recovered in a single clade, with good support and stability. Here, we present a taxonomic review of Stephanopis species attributed to the ‘pentacantha clade’, proposing the new genus Kryptochroma Machado gen. nov. to accommodate them. New distribution records are provided and the following species are described for the first time: Kryptochroma gigas Machado & Viecelli gen. et sp. nov., Kryptochroma hilaris Machado & Teixeira gen. et sp. nov., Kryptochroma quadrata Machado & Viecelli sp. nov. and Kryptochroma septata Machado & Teixeira sp. nov. The species Stephanopis borgmeyeri is considered a nomen dubium, Stephanopis aheneus is a junior synonym of Epicadus tuberculatus, Stephanopis quimiliensis is transferred to Ulocymus and Stephanopis stelloides is transferred to Epicadus, being Epicadus stelloides the senior synonym of Stephanopis salobrensis, Stephanopis trilobata and Epicadus caudatus.
Four new Colombian species of the spider genus Medionops Sánchez-Ruiz & Brescovit, 2017 are described and illustrated: M. carolinae sp. nov. (male and female) from Boyacá department, M. cauca sp. nov. (male) from Cauca department, M. luiscarlosi sp. nov. (male and female) from Caldas department and M. santarosa sp. nov. (male) from Risaralda department. Additionally, the distribution of the type species of M. blades Sánchez-Ruiz & Brescovit, 2017, previously known only from Bogotá, Cundinamarca department, is extended to several localities in the Boyacá department. An updated identification key for all Medionops species is provided.
Six new species of the genus Caccothryptus (Coleoptera: Limnichidae) are described from the Himalayan region: C. brendelli sp. nov., C. arakawae sp. nov., C. championi sp. nov., C. larryi sp. nov., C. tardarsauceae sp. nov. and C. abboti sp. nov. Photographs of type specimens with relevant morphological characters to distinguish species are provided.
The whip spider family Charinidae Quintero, 1986 is the most speciose and widely distributed in the arachnid order Amblypygi Thorell, 1883. It comprises three genera and 95 species distributed across all tropical continents and the eastern Mediterranean. Despite recent advances in the taxonomy of the family, a global revision of all its species, necessary to advance understanding of its systematics, biogeography and evolution, has never been conducted. In the present contribution, the family is revised in its entirety for the first time, including all previous names and 33 new species, 24 in the genus Charinus Simon, 1892: C. alagoanus sp. nov., C. apiaca sp. nov., C. carinae sp. nov., C. carioca sp. nov., C. carvalhoi sp. nov., C. cearensis sp. nov., C. diamantinus sp. nov., C. euclidesi sp. nov., C. goitaca sp. nov., C. guayaquil sp. nov., C. imperialis sp. nov., C. loko sp. nov., C. magalhaesi sp. nov., C. miskito sp. nov., C. mocoa sp. nov., C. monasticus sp. nov., C. palikur sp. nov., C. perquerens sp. nov., C. puri sp. nov., C. renneri sp. nov., C. sooretama sp. nov., C. souzai sp. nov., C. susuwa sp. nov., C. una sp. nov.; eight in the genus Sarax Simon, 1892: S. bilua sp. nov., S. dunni sp. nov., S. gravelyi sp. nov., S. indochinensis sp. nov., S. lembeh sp. nov., S. palau sp. nov., S. rahmadii sp. nov., S. tiomanensis sp. nov.; and one in the genus Weygoldtia Miranda et al., 2018: W. consonensis sp. nov. Taxonomic keys to the 132 species (excluding four nomina dubia) are presented and several taxonomic rearrangements implemented. Four subspecies are elevated to species level: Charinus cavernicolus Weygoldt, 2006, C. elegans Weygoldt, 2006, C. longipes Weygoldt, 2006, and Sarax bispinosus (Nair, 1934). Sarax batuensis Roewer, 1962 is removed from synonymy with Sarax buxtoni (Gravely, 1915) and S. buxtoni newly synonymized with Sarax rimosus (Simon, 1901). Stygophrynus moultoni Gravely, 1915 is transferred to Sarax, resulting in Sarax moultoni (Gravely, 1915) comb. nov. Ten species are transferred from Charinus to Sarax, resulting in new combinations: S. abbatei (Delle Cave, 1986) comb. nov., S. bengalensis (Gravely, 1911) comb. nov., S. dhofarensis (Weygoldt, Pohl & Polak, 2002) comb. nov., S. ioanniticus (Kritscher, 1959) comb. nov., S. israelensis (Miranda et al., 2016) comb. nov., S. omanensis (Delle Cave, Gardner & Weygoldt, 2009) comb. nov., S. pakistanus (Weygoldt, 2005) comb. nov., S. seychellarum (Kraepelin, 1898) comb. nov., S. socotranus (Weygoldt, Pohl & Polak, 2002) comb. nov. and S. stygochthobius (Weygoldt & Van Damme, 2004) comb. nov.
The Chinese species of the genus Chinoperla Zwick, 1980 are reviewed. One species from Hainan is described as new to science: C. changjiangensis sp. nov. Chinoperla gorohovi Sivec & Stark, 2010 is reported from China for the first time, with a redescripion and color images of the male, and the first description of the female and egg. Chinoperla nigrifrons (Banks, 1939) is redescribed and illustrated, synonymy of C. furcomacula (Wu, 1973) is confirmed. Taxonomic relationships within the studied species are discussed. A provisional key to the six known species of Chinoperla for China is presented.
The Neotropical D. agenor species group of the subgenus Luederwaldtinia Martínez, 1951 is taxonomically reviewed. After examination of specimens in collections, the D. agenor species group will consist of seventeen species, a leap from the former nine species: D. agenor (Harold, 1869), D. amplicollis (Harold, 1869), D. belus (Harold, 1880), D. centralis (Harold, 1869), D. deyrollei (Harold, 1869), D. sagittarius (Harold, 1869) stat. rev., D. tristis (Luederwaldt, 1923), D. triquetrus (Luederwaldt, 1923), D. validipilosus (Luederwaldt, 1931), D. fornicatus (Luederwaldt, 1931) [transferred to D. lucasi species group and to which D. bosqi (Pereira, 1941) is considered to be a new synonym], D. inachoides (Felsche, 1901) [transferred from the group fisus (Selenocopris)], D. simplicicornis (Luederwaldt, 1935) [transferred from the group fisus (Selenocopris)], D. subaeneus (Castelnau, 1840) [transferred from the D. carbonarius species group], D. rafanunezi sp. nov. from Mato Grosso, D. henripittieri sp. nov. from Venezuela and D. enioi sp. nov. from Guatemala and Costa Rica. This review was based on external morphology and characters of the male genitalia (aedeagus and internal sac). Descriptions of new species and redescriptions were made after examining the type-specimens of the already described species. Lectotype designations, species revalidations, redescriptions and new species descriptions are provided.