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The overarching aim of this doctoral research was to examine and quantify the spatiotemporal variability in the movements of nomadic ungulates to better understand the possible drivers and characteristics of such movements as well as to examine the particular conservation challenges associated with nomadic movements.
1. Recent research highlights the ecological importance of individual variation in behavioural predictability. Individuals may not only differ in their average expression of a behavioural trait (their behavioural type) and in their ability to adjust behaviour to changing environmental conditions (individual plasticity), but also in their variability around their average behaviour (predictability). However, quantifying behavioural predictability in the wild has been challenging due to limitations of acquiring sufficient repeated behavioural measures.
2. We here demonstrate how common biologging data can be used to detect individual variation in behavioural predictability in the wild and reveal the coexistence of highly predictable individuals along with unpredictable individuals within the same population.
3. We repeatedly quantified two behaviours—daily movement distance and diurnal activity—in 62 female brown bears Ursus arctos tracked across 187 monitoring years. We calculated behavioural predictability over the short term (50 consecutive monitoring days within 1 year) and long term (across monitoring years) as the residual intra-individual variability (rIIV) of behaviour around the behavioural reaction norm. We tested whether predictability varies systematically across average behavioural types and whether it is correlated across functionally distinct behaviours, that is, daily movement distance and amount of diurnal activity.
4. Brown bears showed individual variation in behavioural predictability from predictable to unpredictable individuals. For example, the standard deviation around the average daily movement distance within one monitoring year varied up to fivefold from 1.1 to 5.5 km across individuals. Individual predictability for both daily movement distance and diurnality was conserved across monitoring years. Individual predictability was correlated with behavioural type where individuals which were on average more diurnal and mobile were also more unpredictable in their behaviour. In contrast, more nocturnal individuals moved less and were more predictable in their behaviour. Finally, individual predictability in daily movement distance and diurnality was positively correlated, suggesting that individual predictability may be a quantitative trait in its own regard that could evolve and is underpinned by genetic variation.
5. Unpredictable individuals may cope better with stochastic events and unpredictability may hence be an adaptive behavioural response to increased predation risk.
This paper argues that traces only range over individual semantic types and cannot be type shifted into higher types to circumvent this restriction. The evidence comes from movement targeting positions where DPs must denote properties and the behavior of definite descriptions in these positions. These constraints on possible traces demonstrate that syntactic operations impose active restrictions on permissible semantic types in natural language.
The four novels of Cătălin Dorian Florescu (Wunderzeit, Der kurze Weg nach Hause, Der blinde Masseur, Zaira) analysed here are interpreted in relation to the chronotope, a term developed by Michail Bachtin meaning the connectedness of time and space in narrative. Space knows two opposite dimensions in Florescu’s works: the West (Switzerland, USA) and the East (Romania). The Romanian space is represented by three different images corresponding to three different periods: Romania between the two world wars, during the communist period and after 1989. The main characters are Romanians who leave their country of origin during the communist period, hoping that they will find a better life in the West. After the revolt in 1989, the characters return to the space of their childhood, where they could find themselves and happiness once again. There are analyzed different aspects of the aesthetic space Romania: exotic space, space of discovery of oneself, spiritual space of traditions, but also space of disappointment and of perils. Space is in close connection with movement, the movement of the protagonists from one dimension to another, which is also the basis of the plot.