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This paper revises the genus Ganelius Benesh, which is endemic to Madagascar, in the stag beetle tribe Figulini Burmeister (Coleoptera: Lucanidae: Lucaninae). Species in the genus are striking in their highly sexually dimorphic mandibles, a character that rarely occurs in the tribe. The genus was previously comprised of four species, but three of these names were problematic or questionably applied. A lectotype is designated for G. madagascariensis Laporte, a neotype is designated for Ganelius oberndorferi Nonfried, and Nigidius passaliformis Benesh is newly synonymized with G. oberndorferi. Two new species, G. gnamptus Paulsen and G. zombi Paulsen, are described from western Madagascar. The identity of the overlooked Ganelius nageli (Kriesche) is fixed through a neotype designation, and the species is moved to the new genus Agnelius, which is distinguished from Ganelius by a lack of sexual dimorphism, serrate protibiae, and a more flattened body.
Generic placement of Nearctic species of Ochodaeinae has lagged behind changes in generic concepts in the group. In order to place Nearctic species into the appropriate genus-level taxa, several nomenclatural changes are made. The new generic name Xenochodaeus is proposed for species with an elongate, subparallel sulcus on the propygidium and a longitudinally impressed mentum. The genus contains 6 species. Xenochodaeus americanus (Westwood), new combination, is valid and is removed from synonymy with X. musculus (Say), new combination. A neotype is designated for Odontaeus musculus. Ochodaeus opacus LeConte is synonymized under X. americanus. Other new combinations include Xenochodaeus luscinus (Howden), X. planifrons (Schaeffer), X. simplex (LeConte), and X. ulkei (Horn). The new generic name Cucochodaeus is proposed for species having the propygidial sulcus absent, stridulatory peg absent, and possessing 9 antennomeres. The genus contains 1 species, C. sparsus (LeConte), new combination. Ochodaeus mandibularis Linell is placed into synonymy with C. sparsus. Ochodaeus gnatho Fall is transferred to the genus Codocera Eschscholtz, resulting in C. gnatho (Fall), new combination. Two new combinations result from Nearctic species transferred to Neochodaeus Nikolajev: N. repandus (Fall) and N. striatus (LeConte). Three Nearctic species are transferred to Parochodaeus Nikolajev, resulting in new combinations: Parochodaeus californicus (Horn), P. duplex (LeConte), and P. peninsularis (Horn). A lectotype is designated for Ochodaeus kansanus Fall, and that same specimen is designated as the neotype of Ochodaeus duplex (LeConte), objectively synonymizing O. kansanus with O. duplex. A checklist of Nearctic species and key to Nearctic genera are provided.
The monotypic aesaline genus Lucanobium Howden and Lawrence (Coleoptera: Lucanidae) was previously known only from Venezuela. A second species is here described as new from French Guiana, extending the range of the genus approximately 1800 km to the southeast. The generic description of Lucanobium is updated with respect to the discovery of a second species.
The Neotropical members of the genus Aesalus Fabricius (Coleoptera: Lucanidae: Aesalinae) are transferred to a new genus, Trogellus, due to their morphological dissimilarity and molecular divergence from the Old World species of Aesalus, and the new genus is revised. A neotype is designated for A. trogoides Albers and a lectotype for A. neotropicalis Bates. Aesalus smithi Bates is placed into synonymy with Aesalus trogoides, new synonymy. Two new combinations result from the transfer of species formerly in Aesalus: Trogellus trogoides from Mexico and Trogellus neotropicalis from Guatemala. In addition to the two known species, eight new species are described: T. catrachitus, T. chapinitus, T. hawksi, T. herrmanni, T. maesi, T. narizotus, T. ticiticus, and T. trifinius. This brings the total number of species of Central American aesalines to ten. A phylogeny of Aesalini based on two gene regions of ribosomal DNA is presented. Due to clear morphological differences and large molecular divergence between species groups, two additional new subgenera of Trogellus are proposed: Mayaesalus and Trogoides.
Multiple sources of confusion surround the identity of Ochodaeus grandiceps Fairmaire, 1897, from Sichuan, China (Coleoptera: Ochodaeidae). Herein the type specimen is illustrated to solve these taxonomic issues. Examination of the holotype indicates that the species must be transferred to the genus Nothochodaeus Nikolajev, 2005, resulting in a new combination. The genus Mimochodaeus Nikolajev, 2009, based on a misidentifi cation of O. grandiceps as its purported type species, is discussed. Finally, Cuban specimens had been treated erroneously as belonging to O. grandiceps, and so the species had remained undescribed. A new species, Parochodaeus perdidus, is described to accommodate them.
Adults of three known species of Ochodaeus Dejean (Coleoptera: Scarabaeoidea: Ochodaeidae) in Madagascar have a strongly granulate pronotum. Three new species are described herein that have a smooth or weakly granulate pronotum: Ochodaeus modopunctatus, O. polypollicatus, and O. meandrus. A key to the species of Ochodaeus from Madagascar is provided.