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Sympetrum fonscolombii is for the first time reported for Tuva (Tyva Republic, Russia), as found in the UbsuNur Depression. New data are provided on Odonata of the Turan or TuranUyuk Depression of Tuva, including the first record of Somatochlora exuberata in Tuva beyond the Todzha Depression and Coenagrion armatum, C. ecornutum, Aeshna juncea, A. grandis, Somatochlora graeseri and Libellula quadrimaculata for the first time reported for the Turan Depression. New distributional data and comments on Ophiogomphus spinicornis Selys, 1878 are added. Somatochlora alpestris found at Lake Oyskoe is for the first time reported for the southern Krasnoyarskiy Kray.
In the period 18th – 31st May 2017, 33 localities were examined for Odonata in the Central Zagros area of Iran: 16 in Markazi Province, 14 in Lorestan Province and 3 in Esfahan Province; in 27 of those localities Odonata were found, 25 species in total. For Markazi Province, only one species, Calopteryx splendens, was previously reported (and also found by us), so of 17 species found there 16 are formally new provincial records (Epallage fatime, Lestes barbarus, Coenagrion ornatum, C. persicum, Enallagma cyathigerum, Ischnura elegans, I. intermedia, I. pumilio, Platycnemis kervillei, Anax imperator, Anaciaeschna isoceles, Caliaeschna microstigma, Libellula depressa, Onychogomphus lefebvrii, Orthetrum brunneum, Sympetrum fonscolombii). Of 17 species
found in Lorestan, 5 are new for this province (L. barbarus, Aeshna mixta, Orthetrum taeniolatum, Sympetrum arenicolor, S. striolatum). Only two species were seen in Esfahan Province, in which little time was spent. Notes on variation and taxonomy are provided for Sympecma paedisca, C. ornatum (considered to be a senior synonym of C. vanbrinkae because of variation in the presumed main diagnostic character), E. cyathigerum, I. elegans, Gomphus schneideri (including discussion of G. amseli), as well as notes on habitats of most species and the general characteristics of the area.
The Odonata fauna of flat marshy areas of the Gulf of Siam coast in Koh Kong Province of Cambodia, containing 55 species, is considered. The published data of 2010- 2013 and new data of 2014 and 2016 on the surroundings of Koh Kong town are compiled and the first data on the area of large swampy Melaleuca forests at Andoung Tuek village are presented as well as some occasional photographic records. Gynacantha bayadera, Lyriothemis mortoni and Pornothemis serrata were for the first time recorded for Cambodia. Mortonagrion falcatum was found unexpectedly abundant at Andoung Tuek.
Results are presented of an odonatological survey of 23 localities in Thala Barivat District of Stung Treng Province, northern Cambodia, on July 26 – August 1, 2016. Most localities were situated in areas of open low deciduous dipterocarp forests on gravel soils, some at hillside areas of tall evergreen dipterocarp forest. The great Mekong River right bank was studied within 7 km downstream of its Nimith (Nimet, Khon Thai, Labak Koun, Khone Pha Pheng) Waterfall. In total, 55 species were found, of which 52 identified to species and three to genus. Two species, Gynacantha saltatrix Martin, 1909 and Macrogomphus matsukii Asahina, 1986, were recorded in Cambodia for the first time. Five obligatory lotic species were found at the Mekong River, namely Dysphaea gloriosa, Prodasineura coerulescens, Burmagomphus asahinai, Nychogomphus duaricus and Onychothemis testacea, including tenerals of P. coerulescens and B. asahinai. Most probably these species breed in the Mekong reach which is enriched with oxygen downstream of the great waterfall cascade.
Results of an odonatological survey of Mondulkiri Province of Cambodia, at the foothills and Central Plateau of the Annamese Mts. in June 8 – June 17, 2014 are presented. Of 106 Odonata species met (46 zygopterans, 60 anisopterans), 97 were identified to previously known named species, of which 15 are reported for the first time for Cambodia, namely Mnais mneme Ris, 1916, Rhinocypha seducta Hämäläinen et Karube, 2001, Philoganga loringae Fraser, 1927, Rhinagrion hainanense Wilson et Reels, 2001, Amphiallagma parvum (Selys, 1876), Ceriagrion chaoi Asahina, 1967, Paracercion malayanum (Selys, 1876), Prodasineura doisuthepensis Hoess, 2007, Protosticta grandis Asahina, 1985, Tetracanthagyna waterhousei McLachlan, 1898, Gomphidia kruegeri Martin, 1904, Heliogomphus chaoi Karube, 2004, Leptogomphus baolocensis Karube, 2001, Microgomphus jurzitzai Karube, 2000, Onychothemis culminicola Förster, 1904, and two species for which the specimens collected on this trip were described as new subspecies: Indolestes gracilis expressior Kosterin, 2015, Coeliccia poungyi dasha Kosterin, 2016. Five species collected on this trip have been described elsewhere as new to science, namely Onychargia priydak Kosterin, 2015, Prodasineura hoffmanni Kosterin, 2015, Asiagomphus reinhardti Kosterin et Yokoi, 2016, Euthygomphus schorri Kosterin, 2016 and Risiophlebia guentheri Kosterin, 2015. So the total number of the first country records of named species made on this trip is 22. Still four species found may be undescribed. The number of named species recorded in Cambodia has reached 178. Remarks on taxonomy and variation of Euphaea masoni Selys, 1879, E. ochracea Selys, 1859, Aciagrion approximans (Selys, 1876), and Lamelligomphus castor Lieftinck, 1941 are provided. Characters of mature males of R. seducta are updated. Notes on habitats and habits of some species are provided. Onychothemis culminicola and O. testacea Laidlaw, 1902 seem to exclude each other at rivers, the former occupying smaller and more elevated ones; a putative hybrid male was observed. General notes on the area and field impressions are briefly outlined.
The Tyva Republic, within the Russian Federation, has acquired its present name in 1991. It was formerly known as the Tuvinian Autonomous Region within the USSR. Its common names in the past centuries were Uryankhaiskii Krai or Soyotskii Krai. In modern Russian, it is known under the name Tuva (not Tyva), which will be used further in the text. This country, for example, embraces the sources of the great Siberian river Yenisei and its capital, the town of Kyzyl, represents the geographical centre of Asia. To date, this vast and clear cut territory was hitherto almost unexplored in odonatological respect. In particular, it was not visited by the outstanding Siberian odonatologist B. F. Belyshev. Scarce materials from the recent Tuvinian territory were mentioned by Valle (1942), who considered it to be Mongolian. In the only paper concerning the dragonflies of Tuva in particular (two localities in its very south) 14 species were reported (Zaika & Kosterin 1990). The "transitional form between Ophiogomphus serpentinus and O. reductus” considered in that work, however, turned out to be the Chinese-Mongolian species Ophiogomphus spinicornis (Kosterin, 1999). Finally, one of us published two works on the general lentic fauna of the Ubsu-Nur Hollow in general (Zaika, 1996) and Lake Tere-Khol' in particular (Zaika, 1999), where 26 and 9 species were listed, respectively. The latter work was based on larval identification. Because this is difficult, this information should be taken with caution. The presence of species such as Ischnura pumilio and Cordulia aenea was not confirmed by records of adults and may be erroneous.
More Odonata found at the Cardamonean foothills in Koh Kong Province of Cambodia in 2014-2018
(2018)
Still unpublished data on Odonata of the coastal foothills of the Cardamom Mts. in Koh Kong Province, SW Cambodia obtained in 2014-2018 are provided. Tetrathemis flavescens Kirby, 1889 is for the first time reported for Cambodia and Risiophlebia guentheri Kosterin, 2015 for SW Cambodia. The photographic records from this area of Gynacantha basiguttata Selys, 1882, Gynacantha demeter Ris, 1911, Heliaeschna crassa Krüger, 1899, Amphithemis curvistyla Selys, 1891 and Pornothemis serrata Krüger, 1902 have been confirmed by specimens. The known local faunas of two neighbouring rivulets in Koh Kong Province, rich in Odonata, are updated and summarised.
New data on Odonata of the Preah Sihanouk Province of Cambodia obtained in March 2017 and November 2018 are presented. The presence of Onychargia atrocyana Selys, 1865 in Cambodia is confirmed. Twentyone species are added to the known fauna of the Kbal Chhay Waterfall environs, 19 species to that of Ream Peninsula and 4 to that of Koh Rong Island. The total number of species registered for Kampong Saom Peninsula amounts to 74. The presented data are rather of historical importance since most of the remaining forest has been quickly and irreversibly logged a few months ago. The validity of Gynacantha demeter Ris, 1911 as a species distinct from G. dohrnii Krüger, 1899 is doubted.
Phnom Tumpor is a scarcely accessible basalt table mountain in the Cardamom Mts. in Pursat Province of Cambodia. On top surface it bears tall evergreen forest (ca 1100 m a.s.l.), concealing a slow rivulet, O'Gran, being a chain of deep pools. It was examined odonatologically on March 1418th, 2019. Six common species were recorded in dry and burnt scrub on the Phnom Tumpor slopes and ten on the forested upper surface at O'Gran, among them Polycanthagyna erythromelas (Selys, 1891) and Macromia sp. cf. pinratani Asahina, 1987 for the first time in Cambodia. The peculiarities of the males of Coeliccia kazukoae Asahina, 1984 from Phnom Tumpor and the problem of distinguishing females of M. pinratani and M. moorei are discussed.
Odonatological results of a trip across the southern Far East of Russia, from nearly the border of North Korea to Khabarovsk from July 1st to 16th, 2014, are presented. In total, 50 species were recorded, including those with limited presence in Russia, such as Paracercion calamorum, P. hieroglyphicum, P. plagiosum (for this species the 3rd Russian locality is reported), Pseudocopera tokyoensis, Stylurus annulatus (2nd Russian locality), Sinictinogomphus clavatus; Trigomphus citimus, Macromia daimoji (3rd Russian locality), M. manchurica, Deielia phaon, Lyriothemis pachygastra (2nd Russian locality). For S. annulatus, M. manchurica and D. phaon the northernmost known localities in the world are reported. D. phaon, earlier reported from one locality in Russia, have been found in five localities in Primorye and for the first time reported for Khabarovskiy Kray in general and Bol'shekhekhtsirskiy State Nature Reserve in particular. A trend of mutual exclusion of two abundant Shaogomphus postocularis epophthalmus and S. schmidti is supposed: the lowermost Ussuri River / Amur River, respectively. Variation in Paracercion spp. and Macromia amphigena fraenata is discussed.
Phnom Kulen is a small and low plateau in the northern Cambodia still partly covered with evergreen forests and isolated from similar habitats by the Cambodian Lowland at least for 60 km. A preliminary checklist of its Odonata is provided, including 97 species. Burmagomphus sp. cf. willamsoni and Macromia callisto are for the first time reported for Cambodia.
Lake Tonle Sap in NW Cambodia is the largest freshwater lake in Southeast Asia and one of the most productive freshwater ecosystems in the world, so its banks are a home for ca 1,5 million people. It serves as a natural reservoir of the excess water of the Mekong River and cyclically changes its area from 2,500 km2 in May to 16,000 km2 in October. Its banks are naturally occupied by temporarily inundated forest and scrub, at present mostly replaced by rice fields. The present day semiaquatic vegetation of the lake is to a large extent formed by invasive plant species. The hitherto existing data on Odonata of the lake are very scarce. The author briefly examined the bank and floodplain at the NW part of the lake in February/March 2017, June and November 2018 and December 2019. Five main localities studied are described and illustrated
in detail. In total 41 odonate species of four families (22 in Libellulidae) were found. Most of them are common and widespread lentic species but Macrogomphus phalantus is a species hitherto known only by few specimens from swamped forests of Borneo and Sumatra; its Tonle Sap population was earlier described by the author as the subspecies M. phalantus jayavarman Kosterin, 2019. The earlier published report by Seehausen et al. (2016) of Sinictinogomphus clavatus (not found by the author) was a considerable extension of the known species’ range to the south. Six species were found in all main examined localities and ten only in one of them. At any season at the lake immediate bank (that is water front at the lowest level), Brachythemis contaminata predominates overwhelmingly, Orthetrum sabina and Crocothemis servilia are numerous, two damselfly species, Pseudagrion microcephalum and P. rubriceps, invariably occur at floating vegetation (mostly water hyacinth), and Trithemis pallidinervis, Urothemis signata, Rhyothemis phyllis, R. variegata and Tholymis tillarga are common at bushes. Agriocnemis nana, Ceriagrion praetermissum, Ischnura senegalensis, Macrogomphus phalantus, and Aethriamanta aethra were occasionally met at the lake bank. Other 26 species were found, with different occurrence and quantity, on the lake floodplain. Variation of the male occiput coloration of Amphiallagma parvum is commented.
Prey Long (Prey Lang) Forest is the largest remaining lowland rainforest in Indochina, shared by Stung Treng, Preah Vihear, Kampong Thom and Kratie Provinces of Cambodia, which has been persisted until present because of the lack of roads. It includes patches of unique evergreen swamp forests. Odonata of Prey Long forest, including Cheum Takong forest swamp, was briefly examined in December 2019, while the already deforested area was examined in June 2018. The former examination resulted in 40 species, the latter in 34 species, 60 species in total. Two species, Copera chantaburii Asahina, 1984 and Burmagomphus williamsoni Förster, 1914, are for the first time reported for Cambodia from the deforested area (Chey Saen District of Preah Vihear Province). The swamped forest of Cheum Takong provided 17 species, 5 of which were not found elsewhere in the considered area, 3 are rare and 4 generally Sondaic. Prey Long Forest should be re-examined in the rainy season soon after the road to Spong village is constructed.