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Results of a collecting trip to the Samunsam Wildlife Sanctuary in western Sarawak are presented. Several species are reported from Sarawak for the first time: Elattoneura coomansi, Mortonagrion cf aborense, Macrogomphus phalantus and Pornothemis starrei. Other notable records include Coeliccia species, Prodasineura cf interrupta and Raphismia bispina.
The results of a short collecting trip to Perlis in the northwest
of Peninsular Malaysia are reported. Eighty three species were collected, at least 61 of these are new records for the state, and three species are recorded from Malaysia for the first time: Euphaea
masoni Selys, 1879, Archibasis oscillans (Selys, 1877) and Paracercion calamorum (Ris, 1916). A checklist of the Odonata recorded from Perlis is given in an appendix.
Specimens of Neurothemis disparilis Kirby, 1889, N. fluctuans (Fabricius, 1793), N. fulvia (Drury, 1773), N. ramburii (Brauer, 1866), N. stigmatizans (Fabricius, 1775) and N. terminata Ris, 1911, including their subspecies, were studied with the main focus on the morphology of the vesica spermalis, wing maculation, wing venation, abdominal markings and vulvar scales. The results were compared with species descriptions and directly with type specimens where possible. The vesica spermalis, especially the medial process, is useful at least in separating species groups and supports the traditional differentiation methods using wing maculation and venation. The use of other characters in accessing specific status, coupled with known distribution patterns, is discussed. The following taxonomic changes are proposed: Neurothemis manadensis (Boisduval, 1835) stat. nov., Neurothemis papuensis (Lieftinck, 1942) stat. nov. and Neurothemis taiwanensis sp. nov. is described (27.5.1998, Kenting, Pingtung County/Taiwan, L. M. Juang leg.; holotype is deposited at Taiwan Forestry Research Institute, Taipeh, Taiwan). The type of Polyneura palliata Rambur, 1842 was rediscovered at MNHN and designated as lectotype; a lectotype for Neurothemis nicobarica Brauer, 1867 housed at NHMW is designated. The holotype of Neurothemis incerta Brauer, 1867 was rediscovered and synonymized with N. ramburii.
Records of Odonata collected in the vicinity of the Borneo Highlands Resort on Gunung Penrissen in south western Sarawak in the period 2014-2016 are presented.
Notable records include new species of Telosticta Dow & Orr, 2012, Rhinocypha Rambur, 1842 and Leptogomphus Selys, 1878, as well as the first record of the recently described genus Borneogomphus Karube & Sasamoto, 2014 from Sarawak.
New records of Odonata from Kelantan, Malaysia, with a checklist of species recorded from the state
(2017)
We report here the results from field trips to collect Odonata in the central and northeastern parts of Kelantan state, Peninsular Malaysia. Sixty eight species were collected, and 15 of these are new records for the state. Interesting species collected include Euphaea masoni Selys, 1879 and Leptogomphus tioman Choong, 2016. A checklist of the Odonata recorded from Kelantan with a total 131 confirmed species is given in an appendix.
Drepanosticta kosterini sp. nov. (holotype ♂, from Gunung Penrissen, Kuching Division, Sarawak, Malaysian Borneo, deposited in RMNH) is described from both sexes. It is the sister species of D. actaeon Laidlaw, 1934; a fresh description of the male of D. actaeon and the first description of the female are given, along with discussion of variation in this species. Both D. actaeon and D. kosterini are considered to belong to a species group also including D. rufostigma (Selys, 1886) and a preliminary discussion of variation in this species is given, along with illustrations of both sexes. A neighbour joining COI gene tree for D. actaeon and D. kosterini is presented. The relationships of D. actaeon, D. kosterini and D. rufostigma to other members of the Platystictidae are briefly discussed.
The results of a short collecting trip to Kedah in the northwest of Peninsular Malaysia, made in September 2016, are reported. 64 species were collected, 13–14 of these are new records for the state and 28–29 are new records for the mainland of Kedah. A checklist of the Odonata recorded from Kedah (including Langkawi Island) is given in an appendix. At least 126 species of Odonata are now known from the state.
A survey of Odonata on the Indonesian island of Belitung is reported. The work of Belitung Biodiversity Observer on Odonata is briefly outlined. Sixty four species were recorded during the survey, including two new records for the island. A checklist of the known odonate fauna, consisting of 105 species, of the island is given in an appendix.
We report here the results from two field trips to collect Odonata in the Crocker Range National Park in western Sabah, Borneo, Malaysia. Thirtysix species were collected. Telosticta fugispinosa had not been described at the time of collection, nor had the two Devadatta species. There was no published record of Protosticta species cf kinabaluensis before the 2012 expedition, nor of Drepanosticta species cf crenitis.
Records of Odonata from the Lanjak Entimau Wildlife Sanctuary (LEWS) in Sarawak, Malaysian Borneo are presented. Previous records of Odonata from LEWS are critically examined. One hundred and ten species have been recorded within the sanctuary, including three that have yet to be found outside (Drepanosticta adenani, Telosticta iban and "Elattoneura" mauros); records of four more species are regarded as in need of confirmation, those of another six are incorrect. In addition to the three species only known from LEWS, other notable records include: Drepanosticta sbong, Dysphaea lugens, Euphaea sp. cf basalis, Pericnemis kiautarum, Burmagomphus insularis, Gomphidia caesarea, Merogomphus species, Phaenandrogomphus safei, Macromia callisto and Idionyx montana. A fresh illustration of the anal appendages of Drepanosticta sbong in lateral view is provided to make up for inaccuracies in the original illustration. The taxonomy of Phaenandrogomphus safeii is briefly discussed. Zygonyx errans Lieftinck, 1953 is considered a subspecies of Z. ida not Z. iris Additional records from areas adjacent to LEWS are given in an appendix.
The Odonata found during work on the Biodiversity and Ecosystem Function in Tropical Agriculture (BEFTA) Project in Riau Province, Sumatra, Indonesia are reported. Prior to the BEFTA project we are only aware of published records of 37 species of Odonata from Riau Province (these are listed in an appendix). Seventy five species have been recorded during the BEFTA project, including five that have not (Archibasis incisura, Archibasis rebeccae and Pseudagrion williamsoni), or not definitely (Argiocnemis species and Mortonagrion species cf aborense), been recorded in Sumatra before. Macromia dione is recorded for the first time since its description. The number of species now known from Riau Province is 88; 51 of these are reported from the province for the first time here.
Records of Odonata from Gunung Melatai, Nanga Gaat and the Kastima Logging area, all in Sarawak’s Kapit Division, are presented. The most notable records are of Matronoides cyaneipennis Förster, 1897 and Heliogomphus blandulus Lieftinck, 1929. A distribution map for Matronoides cyaneipennis and updated distribution maps for three species from the Coeliccia borneensisgroup are given. Tentative identifications to species of previously published records of Idionyx females are given. The genus Heliogomphus in Borneo, Peninsular Malaysia and Singapore is reviewed and a simple one marker molecular analysis is presented for the genus in this region. Based on reexamination of specimens from the genus and the molecular results, an additional member of the genus is reported from Borneo: H. sp. cf olivaceus Lieftinck, 1961. Although both morphological and molecular results remain incomplete, it does appear likely that there is at least one more species of Heliogomphus present in Borneo than has been recognised until now, and that H. borneensis Lieftinck, 1964 may be a junior synonym of H. kelantanensis (Laidlaw, 1902).
Records of Odonata from the southwest of Sri Aman Division and the extreme east of Serian Division in Sarawak are presented. The sampled areas are interesting not only because they are poorly known for Odonata but also because many are just to the south and west of the Lupar Line which is a division between the ancient Sunda shelf and more recent geological formations. Differences between the odonate faunas on either side of the Lupar Line are discussed. Eightyfive species of Odonata were recorded during the surveys reported on. The single most notable record is that of Coeliccia southwelli Dow & Reels, 2011, which represents a considerable extension to the known range of this species. Other interesting records include Telosticta dupophila (Lieftinck, 1933), T. species cf longigaster Dow & Orr, 2012, Podolestes parvus Dow & Ngiam, 2019 and Heliogomphus species cf olivaceous Lieftinck, 1961. Variation in the markings of Stenagrion dubium (Laidlaw, 1912) across its range is discussed and a gene tree using the COI marker is presented to illustrate the high variability of this species in this marker. However the variability in COI does not appear to be correlated with other characters.
Records of Odonata from Sarawak's Bintulu Division are presented. One hundred and sixtysix (or more) species are listed, of which three (Oligoaeschna amata (Förster, 1903), O. buehri (Förster, 1903) and Oligoaeschna (?) species) had not previously been recorded in Sarawak and Macromia species cf dione Lieftinck, 1971 had not previously been recorded from Borneo. Additionally this is the first published record of Prodasineura tenebricosa Lieftinck, 1937 from Sarawak, although it had been found at another location in the state prior to its discovery in Bintulu, and the first published record of Phyllothemis raymondi Lieftinck, 1950 from Borneo, although specimens from Kalimantan are present in the collections of the Naturalis Biodiversity Center, Leiden. Other notable records include "Elattoneura" longispina Lieftinck, 1937, Pericnemis dowi Orr & Hämäläinen, 2013, Linaeschna polli Martin, 1909, Burmagomphus arthuri Lieftinck, 1953, Heliogomphus borneensis Lieftinck, 1964, Merogomphus femoralis Laidlaw, 1931, Chlorogomphus species, Macromidia genialis erratica Lieftinck, 1948 and Tetrathemis flavescens Kirby, 1889. Altogether there do not appear to be any previous published records from Bintulu Division of 52 of the species listed in this paper. The status of Rhyothemis fulgens Kirby, 1889 is discussed and illustrations of its anal appendages and those of R. pygmaea (Brauer, 1867) are provided. Illustrations of the anal appendages of Tetrathemis hyalina Kirby, 1889 and T. irregularis Brauer, 1868 are also provided. Identification problems when using COIbased DNA barcoding with some species of Archibasis are discussed, and the nuclear marker ITS is shown to be a successful alternative in these cases; COI and ITS gene trees for part of the genus are included. Some COI data for Macromia species are published and the marker is used to identify larvae of M. corycia Laidlaw, 1922 and establish the relationship of another Macromia larva with M. dione, a very poorly known species from Sumatra. Macromia euterpe Laidlaw, 1915 is considered to be the same species as M. westwoodii Selys, 1874 and dropped from the list of species known from Sarawak, however formal combination of the two species is left for a peer reviewed publication. A detailed list of previously unpublished specimens from the locations covered is given in an appendix. Concise checklists for two of the locations covered Similajau National Park (54 species) and the Bukit Mina Wildlife Corridor (84 species) are given in another appendix.
Records of Odonata collected in Gunong Mulu National Park in Sarawak are presented. Between 2005 and 2019, in 12 surveys that lasted between one week and five months, 163 species were collected. The collections from Gunong Mulu National Park are of importance for the taxonomic study of dragonflies and damselflies in Borneo; several species have been described based on material collected in the Park.
We report here the results from field trips to collect Odonata in the northeastern parts of Kelantan state and the north of Terengganu state, Peninsular Malaysia. Eighty four species were collected, and four of these are new records for the state Kelantan and 10 are new records for the state of Terengganu. Notable records obtained from the field trips were Euphaea masoni, Coeliccia sameerae, Pseudagrion ?lalakense, Leptogomphus tioman and Macromia cupricincta. Checklists for Kelantan (140 species) and Terengganu (132 species) are given in an appendix.
Previously unpublished records of Odonata from the states of Negeri Sembilan and Selangor in Peninsular Malaysia are presented. One hundred and eight species are listed, of which 77 were collected in Negeri Sembilan and 87 in Selangor. Fifteen of the species recorded from Negeri Sembilan and seven of those recorded from Selangor appear to be first records for the respective state. Notable records include Drepanosticta sp. cf hamadryas Laidlaw, 1931, Rhinocypha pelops Laidlaw, 1936, Acrogomphus ?malayanus Laidlaw, 1925, Heliogomphus kelantanensis (Laidlaw, 1902); Onychogomphus duaricus Fraser, 1924, Macromia cupricincta Fraser, 1924, Idionyx montana Karsch, 1891, Chalybeothemis chini Dow, Choong & Orr, 2007 and Hylaeothemis clementia
Ris, 1909 Ris, 1909. Provisional checklists of the Odonata known from Selangor plus the
Federal Territory of Kuala Lumpur (171 species) and for Negeri Sembilan (116 species)
are given in appendices.
Odonata recorded during two brief sampling trips to the Kapit Town area in Kapit Division, Sarawak in 2020 are reported on. Seventy nine species were recorded, of which at least six (Onychargia atrocyana Selys, 1865, Ischnura senegalensis (Rambur, 1842), Pseudagrion lalakense Orr & van Tol, 2001, Megalogomphus borneensis (Laidlaw, 1914), Agrionoptera insignis (Rambur, 1842) and Hydrobasileus croceus (Brauer, 1867)) are first records for the division, bringing the total number of Odonata known from the division to 160. Notable records include Coeliccia kenyah Dow, 2010, Teinobasis laidlawi Kimmins, 1936, Burmagomphus insularis Laidlaw, 1914, Leptogomphus sp. cf coomansi Laidlaw, 1936 and Macromia callisto Laidlaw, 1922. Remarks are made on the habitat preferences of Agrionoptera insignis and Camacinia gigantea (Brauer, 1867). A highly atypical population of Neurothemis Brauer, 1867 (species unclear at present) is reported.