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This paper deals with the taxonomy and faunistics of the genus Diplommatina in Nepal. Altogether 16 species are reported, seven of which are new to science viz. D. abiesiana sp. nov., D. fistulata sp. nov., D. godawariensis sp. nov., D. maipokhariensis sp. nov., D. salgharica sp. nov., D. shivapuriensis sp. nov. and D. syabrubesiensis sp. nov. Information on nine previously reported species is provided: D. exserta Godwin-Austen, 1886, D. folliculus (L. Pfeiffer, 1846), D. miriensis Godwin-Austen, 1917, D. munipurensis Godwin-Austen, 1892, D. oviformis Fulton, 1901, D. pachycheilus Benson, 1857, D. regularis Fulton, 1901, D. silvicola Godwin-Austen, 1886 and D. sperata W.T. Blanford, 1862. Although D. canarica was once reported from Nepal, it is not treated here as it is an endemic of the Western Ghats. A dichotomous identification key for all Nepalese species is presented.
The monotypic Neotropical genus Ectophasiopsis Townsend, 1915 (Diptera, Tachinidae, Phasiinae) is revised, with the addition of two species (one new and another transferred species), and a redefinition of the genus, accompanied by photographs and drawings of specimens and male terminalia. A new combination is proposed, Ectophasiopsis gradata (Wiedemann, 1830) comb. nov., previously Trichopoda Berthold, 1827, and a new species Ectophasiopsis ypiranga sp. nov. is described. A key for the genera of the “Trichopoda typica” subgroup sensu Sabrosky (1950), as well as a key to species of Ectophasiopsis is given. The geographical range of the genus and the host list are updated.
Nine individuals of Apatidelia from Zhejiang Province, China were examined and their barcode sequences were generated and analyzed. A new species, A. morsei Xu & Sun sp. nov., is described and illustrated. The larva, male and female of A. acuminata Leng & Yang, 1998 and the male and female of A. morsei Xu & Sun sp. nov. are associated by mtCOI gene sequences. The male of A. acuminata Leng & Yang, 1998 is re-described and re-illustrated, and the female and the larva of the same species are also described and illustrated. Females and larvae of the genus are here reported for the first time.
Four new species of Nyctonympha Thomson, 1868 are described: N. antonkozlovi sp. nov. and N. sinjaevi sp. nov. from Colombia, N. birai sp. nov. from Venezuela and N. mariahelenae sp. nov. from Brazil (Rondônia), Bolivia and Peru. Nyctonympha flavipes Aurivillius, 1990 is formally excluded from the Peruvian and Brazilian fauna. A provisional key to species of Nyctonympha is provided.
Japanese species of the genus Intybia are revised taxonomically, with the examination of the endophallic structure. Eight species, including one new species Intybia donan sp. nov. from Yonagunijima, are recognized. All species are described or redescribed with a key and figures. The endophallic structure contains one primary sclerite (gonoporal piece), three secondary sclerites (ligula, semigonoporal piece, and spinous plate) in some species, and a membranous basal area densely covered with many spines (spinous area). Based on the structures of the endophallus, the Japanese members of the genus are divided into two species groups (the histrio and pelegrini groups). The pelegrini species group is furthermore subdivided into three subgroups (subgroups 1–3). New distributional records are as follows: I. histrio from Hachijô-jima and Tanega-shima; I. niponica from Sakhalin and I. takaraensis from Tokuno-shima and Amami-Ôshima.
The genus Koiulus gen. nov. and its type-species, Koiulus interruptus gen. et sp. nov., are described from the Russian Far East. The new genus is compared with other genera of Mongoliulidae, in particular with Ussuriiulus Golovatch, 1980, also from the Russian Far East, with which it shares the absence of ozopores from individual body rings distributed along the body, a condition so far otherwise unknown in the superorder Juliformia. A synoptic table of genera and a list of species of Mongoliulidae are presented.
A new species of vermetid gastropod belonging to the genus Novastoa Finlay, 1926, N. rapaitiensis sp. nov., is described from French Polynesia and the Great Barrier Reef, based on morpho-anatomical and molecular data, increasing the recognized extant diversity of this genus from five to six species. The new species is characterized by the largest operculum in the genus to date, with a conspicuous spindle-shaped mammilla that readily distinguishes this species from its congeners. Based on available data, members of the studied populations of N. rapaitiensis sp. nov. are interpreted as a single species, although slight morphological and color differences exist between localities separated by up to 7000 km. The larval development of N. rapaitiensis sp. nov. is inferred to be direct, raising new questions about genetic connectivity and dispersal trajectories across a vast geographic range. Additional molecular data may prove helpful in refining our current knowledge on the morphological variability within this species and verifying the degree of cryptic diversity in this genus.
Tortonian teleost otoliths from northern Italy: taxonomic synthesis and stratigraphic significance
(2017)
The Tortonian fish otoliths of northern Italy have been studied for more than a century and represent one of the best known otolith-based teleost faunas in the Miocene of the Mediterranean Basin. Yet with the growing knowledge on Recent otoliths, an updated taxonomic overview of this fauna is needed. Moreover, new material from hemipelagic Tortonian marls sampled at nine localities is described herein, revealing 109 taxa of which 88 are recognised at species level. Four of these are new: Coryphaenoides biobtusus sp. nov., “Merluccius” rattazzii sp. nov., Neobythites auriculatus sp. nov. and Lesueurigobius stironensis sp. nov. The compilation of previously studied and newly acquired material revealed a total of 118 nominal Tortonian species. At generic level, the fauna is characterised by many modern forms; more than 90% can be assigned to present day genera. At species level, however, more than half of the represented taxa are extinct. Based on the fossil otolith record, the Tortonian fauna of the Mediterranean is most similar to that of the Langhian (Badenian) of the Central Paratethys by sharing many extinct Miocene species, but it is also very close to that of the Pliocene Mediterranean, by sharing many modern Atlantic-Mediterranean forms. The Tortonian fauna is further characterised by many species that are apparently confined to the upper Miocene, resulting in a unique combination of its taxonomic composition.
To date, six species of the Australian endemic millipede genus Boreohesperus have been recognized: all have highly localized distributions, consistent with being short-range endemic species, and all are from the Cape Range and Pilbara region of Western Australia. In this paper, we describe three new species, B. alcyonis sp. nov., B. psittacinus sp. nov., and B. vascellus sp. nov., each from a different island in the Kimberley region of north-western Australia.
Contributions to the knowledge of the mite genus Stigmaeus Koch, 1836 (Acari: Stigmaeidae) of Turkey
(2017)
Based on the mite specimens collected within the scope of a study on Erzincan (Turkey) mite biodiversity, two species of the genus Stigmaeus are described and illustrated here: S. bifurcus sp. nov. as new to science and S. miandoabiensis Bagheri & Zarei, 2012 as a new record for Turkey. Some morphological abnormalities in the new species are noted. The deutonymph of S. miandoabiensis is described for the first time in this study. Discovery of this stage from soil and litter under Pinus sylvestris in Turkey adds more data to our knowledge of the species.
Oromia is the largest National Regional State of Ethiopia. Here we present the first comprehensive checklist of its birds. A total of 804 bird species has been recorded, 601 of them confirmed (443) or assumed (158) to be breeding birds. At least 561 are all-year residents (and 31 more potentially so), at least 73 are Afrotropical migrants and visitors (and 44 more potentially so), and 184 are Palaearctic migrants and visitors (and eight more potentially so). Three species are endemic to Oromia, 18 to Ethiopia and 43 to the Horn of Africa. 170 Oromia bird species are biome restricted: 57 to the Afrotropical Highlands biome, 95 to the Somali-Masai biome, and 18 to the Sudan-Guinea Savanna biome. 26 species of birds of Oromia are Threatened (three Critically Endangered, nine Endangered, and 14 Vulnerable), and an additional 20 species are classified as Near Threatened. International regulations affect 225 Ethiopian bird species. Of the polytypic species, a total of 620 subspecies can be taxonomically assigned to Oromia. 11 of them are endemic to Oromia, 71 to Ethiopia and 150 to the Horn of Africa. This very high diversity underlines the high responsibility of the Oromia National Regional State for bird conservation.
Two new species of the Rhyacophila nigrocephala species group, R. voluta sp. nov. and R. linguiformis sp. nov., are described, diagnosed and illustrated. Rhyacophila voluta sp. nov. is similar to R. pentagona Malicky & Sun, 2002 in male genitalia, but can be diagnosed by the inferior appendages of the same length as the complex of preanal appendages and dorsal lobe of segment IX, fused base of the basal segments of inferior appendages, and slightly incised distal margin of apical segment of inferior appendage. Rhyacophila linguiformis sp. nov. is similar to R. rima Sun & Yang, 1995 and R. esorima Mey, 1996 in male genitalia, but can be diagnosed by the progressively narrowed complex of preanal appendages and the dorsal lobe of segment X, the short anal sclerites, and the small gap between the upper and lower lobes of apical segment of inferior appendages. An updated checklist of 23 species recorded from China is presented.
Nomenclatural changes are made in three previously described genera in the planthopper tribe Hemisphaeriini (Hemiptera: Fulgoromorpha: Issidae: Issinae), viz Gergithus Stål, 1870, Mongoliana Distant, 1909 and Hemisphaeroides Melichar, 1903. In addition, a new genus, Gnezdilovius gen. nov., with Gergithus lineatus Kato, 1933 as its type species, is described for 40 species formerly included in Gergithus, and the generic characteristics of the latter genus is revised. One new species, Gergithus frontilongus sp. nov. from China (Yunnan), is described and illustrated. One additional Gergithus species, previously misidentified as G. signatifrons Melichar, 1906 from Siberut Island, is mentioned and illustrated. Gergithus contusus Walker, 1851 is transferred to Mongoliana and Hemisphaerius atromaculatus Distant, 1916 and H. fuscoclypeatus Distant, 1916 are transferred to Hemisphaeroides. Checklists for all four genera are provided detailing the nomenclatural changes and a key to the 19 genera of Hemisphaeriini is provided. Morphological diversity and distribution of the genera are briefly discussed.
The genus Austrotinodes contains 55 species, largely distributed in the Neotropics, with a few species occurring in the Australian Region. In Brazil, 10 species have been recorded, mainly in the southern and southeastern regions. Herein, we describe 13 new species from Brazil, all of them named in memory of great Brazilian scientists: Austrotinodes absaberi sp. nov., A. adolfolutzi sp. nov., A. berthalutzae sp. nov., A. chagasi sp. nov., A. costalimai sp. nov., A. cruzi sp. nov., A. donagrazielae sp. nov., A. gusmaoi sp. nov., A. lattesi sp. nov., A. lenti sp. nov., A. santosdumonti sp. nov., A. vanzolinii sp. nov. and A. vitalbrazili sp. nov. Additionally, we give new distribution records for A. amazonensis Flint & Denning, 1989, A. longispinum Thomson & Holzenthal, 2010, A. paraguayensis Flint, 1983 and A. taquaralis Thomson & Holzenthal, 2010. We also provide new diagnoses and illustrations for A. amazonensis and A. paraguayensis, to facilitate identification of those species.
Two new species of hangingflies, Terrobittacus rostratus sp. nov. and Terrobittacus angustus sp. nov., are described and illustrated from Yunnan, southwestern China, increasing the species number of Terrobittacus Tan & Hua, 2009 to six. Terrobittacus rostratus sp. nov. differs from its congeners by wings devoid of markings and epandrial appendages slightly longer than half the length of the gonocoxites. Terrobittacus angustus sp. nov. can be recognized by wing markings along R5 distally. A key to species of the genus is updated to include the two new species.
Six species of Anastatus Motschulsky, 1859 (Hymenoptera: Eupelmidae) were described from China in Chinese by J.K. Sheng and coauthors in 1997 and 1998: A. dexingensis, A. flavipes, A. fulloi, A. huangi, A. meilingensis and A. shichengensis. This represents almost half the species of Anastatus recorded from China, but no keys were given to differentiate the species and the original descriptions included only simple line drawings to illustrate the species. Because recognition of these species is critical prior to clarifying the Anastatus fauna of China and of the eastern Palaearctic and Oriental regions, we have redescribed the six species in detail in English based on original type material, illustrating the species through macrophotography of type material and providing a key to differentiate females of the species.
The family Ammotheidae is the most diversified group of the class Pycnogonida, with 297 species described in 20 genera. Its monophyly and intergeneric relationships have been highly debated in previous studies. Here, we investigated the phylogeny of Ammotheidae using specimens from poorly studied areas. We sequenced the mitochondrial gene encoding the first subunit of cytochrome c oxidase (CO1) from 104 specimens. The complete nuclear 18S rRNA gene was sequenced from a selection of 80 taxa to provide further phylogenetic signal. The base composition in CO1 shows a higher heterogeneity in Ammotheidae than in other families, which may explain their apparent polyphyly in the CO1 tree. Although deeper nodes of the tree receive no statistical support, Ammotheidae was found to be monophyletic and divided into two clades, here defined as distinct subfamilies: Achelinae comprises the genera Achelia Hodge, 1864, Ammothella Verrill, 1900, Nymphopsis Haswell, 1884 and Tanystylum Miers, 1879; and Ammotheinae includes the genera Ammothea Leach, 1814, Acheliana Arnaud, 1971, Cilunculus Loman, 1908, Sericosura Fry & Hedgpeth, 1969 and also Teratonotum gen. nov., including so far only the type species Ammothella stauromata Child, 1982. The species Cilunculus gracilis Nakamura & Child, 1991 is reassigned to Ammothella, forming the binomen Ammothella gracilis (Nakamura & Child, 1991) comb. nov. Additional taxonomic re-arrangements are suggested for the genera Achelia, Acheliana, Ammothella and Cilunculus.
With about 250 species, the genus Blaps Fabricius, 1775 is one of the most diverse genera of darkling beetles (Coleoptera: Tenebrionidae: Tenebrioninae: Blaptini: Blaptina). In this study, we provide new insights on the evolutionary relationships of Blaps species using a combined molecular and morphological dataset encompassing 69 distinct Blaps species and subspecies (105 specimens in total, all belonging to the subgenus Blaps), four other representatives of the tribe Blaptini (from the subtribes Gnaptorina, Gnaptorinina and Prosodina) and 12 outgroup species. Five new species of Blaps are also described within the subgenus Blaps: B. effeminata sp. nov. from Libya, B. intermedia sp. nov. from Morocco, B. maldesi sp. nov. from Algeria, B. nitiduloides sp. nov. from Algeria and Tunisia and B. teocchii sp. nov. from Tunisia. The results of the phylogenetic analyses indicate that the genus Blaps is likely paraphyletic; the two highlighted clades are morphologically distinct and correspond to groups previously referred to as sections (I and II) within the subgenus Blaps. This suggests the need for more phylogenetic studies in order to clarify the status of the various genera and subgenera belonging to the tribe Blaptini.
In this review (third part), several species within the Nassarius pauperus complex from the eastern Indian Ocean and western Pacific are treated, including a revised concept of Nassa paupera Gould, 1850, type species of the genus Reticunassa Iredale, 1936. In the most recent taxonomic revision, several species had been synonymized with Nassarius pauperus (Gould, 1850), despite distinctive differences among these species in shell morphology. We sequenced a fragment of the mitochondrial COI and the nuclear 28S genes of all available Nassarius pauperus complex species. Automatic barcode gap discovery and reciprocal monophyly were applied to propose species delimitation hypotheses and to support a new concept of the genus. Using morphological and molecular characters, Reticunassa is elevated to full genus rank. Six new species are described: Reticunassa visayaensis sp. nov., R. poppeorum sp. nov., R. annabolteae sp. nov., R. goliath sp. nov., R. intrudens sp. nov. and R. thailandensis sp. nov., while R. tringa (Souverbie, 1864) is recognized as a valid species.
In Southeast Asia, bats of the genus Tylonycteris Peters, 1872 have traditionally been classified into two wide-ranging species, T. pachypus (Temminck, 1840) and T. robustula Thomas, 1915. Our comparative phylogeographic analyses based on two mitochondrial and seven nuclear genes, combined with our multivariate morphological analyses, show that these species actually represent cryptic species complexes that share a similar biogeographic history in three major regions, i.e., Sundaland, southern Indochina, and northern Indochina. Our molecular dating estimates suggest that Pleistocene climatic oscillations and sea level changes have repeatedly isolated ancestral populations of Tylonycteris spp. in distant bamboo forest refugia. The analyses indicate, however, that populations of the T. pachypus complex were less affected by forest fragmentation in mainland Southeast Asia than those of the T. robustula complex. Accordingly, we propose several taxonomic changes within the genus Tylonycteris: the species T. fulvida and T. malayana are revalidated, and a new species, T. tonkinensis Tu, Csorba, Ruedi & Hassanin sp. nov., endemic to northern Indochina, is described.
This paper describes and illustrates a new genus and a new species belonging to the family Nesticidae based on morphology and supported by molecular data. The new genus, Kryptonesticus gen. nov., groups eight species spread from Bulgaria and Turkey to Croatia, including Montenegro, Bosnia and Herzegovina and Crete. As a result, seven new combinations are proposed: K. eremita (Simon, 1879) comb. nov., K. arenstorffi (Kulczyński, 1914) comb. nov., K. fagei (Kratochvíl, 1933) comb. nov., K. beroni (Deltshev, 1977) comb. nov., K. beshkovi (Deltshev, 1979) comb. nov., K. henderickxi (Bosselaers, 1998) comb. nov. and K. dimensis (López-Pancorbo, Kunt & Ribera, 2013) comb. nov., all ex Nesticus. Kryptonesticus deelemanae gen. et sp. nov. is described on the basis of both sexes and its phylogenetic relationships with closely related species are discussed based on morphological and molecular data (the cox1, rrn and H3 genes). In addition, the species of this new genus (except for K. eremita) are clear candidates for protection: they have highly restricted ranges and some of them show a high degree of adaptation to the subterranean environment.
Umborotula bogorensis (Weber, 1890) is a freshwater sponge species that is recorded occasionally, mainly on islands and peninsulas of Australasia. Less than 10 records with morphological descriptions and illustrations have been published so far, and the most recent record is dated 1978. A list of the few voucher specimens from museum collections is provided here together with the rich unpublished Sasaki collection from Japan, Korea, and Taiwan, recently deposited in a Japanese museum. The present new record from Northeast Thailand enlarges the geographic range of U. bogorensis to the Indochina mainland. A comparison of historical data vs present Thai records is performed by morpho-analysis(SEM) as well as biogeographic, ecological and climatic data. Results show low variability in shape and size of the diagnostic morphotraits in populations scattered over the wide geographic range. Here we also formally accept the new taxonomic status (rank elevation) of the previous suborder Spongillina as a new order Spongillida. The presence of this potentially threatened species in the Sakaerat Biosphere Reserve, together with its possible long-term persistence in the Bogor Botanical Garden, may support its conservation. Only a census of the known, extremely scattered populations will define the status of this species.
Pukupuku arunachalensis sp. nov. (Coleoptera, Scarabaeidae, Rutelinae) from Arunachal Pradesh, India
(2017)
A new species of the genus Pukupuku Muramoto, 2006 is described from the state of Arunachal Pradesh (Northeast India): Pukupuku arunachalensis sp. nov. Relevant diagnostic characters are illustrated and compared with closely related species, Pukupuku curtus (Arrow, 1919) and Pukupuku katsurai (Muramoto, 2002). The distribution of the new species is mapped.
The fossil sirenian material from the upper Oligocene Linz Sands of Upper Austria is reviewed and re-described in detail following a recent approach on the invalidity of the genus Halitherium Kaup, 1838. This morphological study provides the first evidence for the synonymy of “Halitherium” cristolii Fitzinger 1842, “H.” abeli Spillmann, 1959 and “H.” pergense (Toula, 1899), supporting the hypothesis that only a single species inhabited the late Oligocene shores of present-day Upper Austria. In the course of the taxonomic revision of the “Halitherium” species-complex, this taxon is now assigned to the new genus Lentiarenium Voss gen. nov. It represents a more derived sirenian compared to Eocene and early Oligocene taxa distributed across Central Europe and North Africa, which is in accordance with the stratigraphical data. An updated inventory list of all identifiable and referable skeletal material is provided, including a detailed synonymy list for the new taxonomic combination.
The Australian endemic staphylinoid plant bug genus Carvalhoma Slater & Gross, 1977 is revised. The genus is redescribed and its systematic position within Cylapinae is discussed. Carvalhoma malcolmae Slater & Gross, 1977 and C. taplini Slater & Gross, 1977 are redescribed. Three species, C. ovatum sp. nov., C. parvum sp. nov., and C. weiri sp. nov., are described as new to science. A key to species, digital habitus images, SEM images, drawings of male and female genitalia and distribution map are provided.
The genus-group taxon Megatrigon Johnson, 1898, stat. nov., is revised and treated as a valid genus within the Merodontini (= Eumerini). Extensive diagnoses are given for the genus and for its three constituent species groups: argenteus group [11 spp.], nivalis group [monotypic], sexfasciatus group [3 spp.]. Five new generic combinations are proposed within Megatrigon: M. argenteus (Walker, 1852) comb. nov., M. flavimarginatus (Hull, 1964) comb. nov., M. jacobi (Hervé-Bazin, 1913) comb. nov., M. nivalis (Hull, 1964) comb. nov. and M. ochreatus (Hull, 1964) comb. nov. All species of the argenteus group are revised and nine new species are described: Megatrigon apiformis sp. nov., M. argentifrons sp. nov., M. argentimaculatus sp. nov., M. cooksoni sp. nov., M. immaculatus sp. nov., M. magnicornis sp. nov., M. natalensis sp. nov., M. sexmaculatus sp. nov., M. tabanoides sp. nov. Within the sexfasciatus group, M. jacobi (Hervé-Bazin, 1913) comb. nov. is treated as a senior synonym of Eumerus connexus Hull, 1964 syn. nov., but no further work is done at the species level due to insufficient material.
A new species of Charinus is described from Israel and new localities for C. ioanniticus are reported. Charinus israelensis sp. nov. is a cave dwelling species with extremely small median eyes, no median tubercle and reduced lateral eyes. It is similar to C. ioanniticus, which occurs in nearby areas, but can be differentiated by the shape of the carapace, the number of pedipalp spines and the development of the eyes. A detailed comparison is made between the two species, including pictures, drawings and scanning electron micrographs. Charinus ioanniticus is reported here from several new localities in Israel and Turkey. Identification keys to the Charinus species groups and to the species of the bengalensis group are provided.
Magnacarina gen. nov. from Mexico is described. Hapalopus aldanus West, 2000 from Nayarit, is transferred to the new genus with an emended diagnosis creating the new combination Magnacarina aldana comb. nov. Three new species are described: Magnacarina moderata Locht, Mendoza & Medina sp. nov. from Nayarit and Sinaloa; Magnacarina primaverensis Mendoza & Locht sp. nov. and Magnacarina cancer Mendoza & Locht sp. nov., both from Jalisco. Magnacarina gen. nov. is characterized by an unusual bifid palpal bulb, and has a primary projection located in the central area of the palpal bulb and directed retrolaterally; this projection possesses the prolateral superior and retrolateral keels. Next to the primary projection is a secondary projection, which may be short or long, ending in the prolateral inferior and apical keel surrounding the sperm pore. This secondary projection may have prolateral accessory keels and is diagnosed by possessing a nodule of inwardly curled megaspines, located in the basal ventro-retrolateral region of metatarsi I in adult males. Additionally, male tibiae I possess three apophyses. Females of Magnacarina gen. nov. have a single reduced and strongly sclerotized spermatheca, with an apical lobe projecting ventrally, and with a uterus externus that is longer and wider than the spermatheca.
Endogeophilus ichnusae gen. et sp. nov. (Chilopoda: Geophilidae sensu stricto) is described based on three specimens from two localities in south-western Sardinia, examined by light and scanning electron microscopy. The new centipede resembles the rare Ibero-Pyrenean genus Galliophilus Ribaut & Brolemann, 1927 in some features, especially in the forcipular segment, and the temperate European species Geophilus electricus (Linnaeus, 1758) in other features, especially in the ultimate leg-bearing segment. However, the true affinities of E. ichnusae gen. et sp. nov. are uncertain, because the new species departs significantly from the majority of geophilids for the higher number of legs (91–107 pairs in the specimens examined), the slender trunk segments (the sternites being longer than wide), the relatively stout legs (the tarsus being only about twice as long as wide) and the very short setae (≤ 15 mm) scattered on the body surface. All these features are probably derived and suggest adaptation to a more strictly endogeic habit than other geophilids.
Two new species of the millipede genus Anamastigona are described, A. cypria sp. nov. and A. strasseri sp. nov., both from the island of Cyprus. The new species are considered to be most similar to A. terraesanctae Golovatch & Makarov, 2011 from Israel, the three of them forming a species group which is briefly characterized. An updated key to all 20 recognized species of the genus Anamastigona is provided and a short overview of the taxonomy of the genus is given.
Four new species of the genus Coecobrya, C. gejianbangi sp. nov., C. annulata sp. nov., C. ciliata sp. nov., and C. oculata sp. nov., are described from Guangxi caves as the representative of the genus in China. Coecobrya oculata sp. nov. of the boneti-group has 1+1 eyes and a serrate outer edge of the unguiculus. The other three species, devoid of eyes and with a tiny outer tooth on the unguiculus, are assigned here to the tenebricosa-group, assuming that the large tooth on the unguiculus is transformed into a tiny one in cave-obligate species. Clypeal chaetae in Entomobryoidea are systematically surveyed for the first time, and are found to be well diversified at species level. They have a potential taxonomical value in discriminating taxa of morphologically conserved groups.
We revise the Southeast Asian Pholcus bicornutus group in which males are characterized by a unique pair of horns on their ocular area, each of which carries at its tip a brush of hairs. In two species, the two hair brushes are ‘glued’ or ‘waxed’ together by an unidentified substance into a very consistently curved and pointed single median tip. In the other five species known, the hairs are unglued. We present a first revision of ocular modifications in Pholcidae and identify twenty supposedly independent origins. Most cases are in Pholcinae, and all but one case are limited to the male, suggesting sexual selection as the main driving force in the evolution of ocular modifications in Pholcidae. Previously, the Pholcus bicornutus group consisted of four species limited to the Philippines. We describe four new species, including three species from the Philippines (P. olangapo Huber, sp. nov.; P. kawit Huber, sp. nov.; P. baguio Huber, sp. nov.) and the first representative from outside the Philippines (P. mulu Huber, sp. nov. from Sarawak, NE Borneo) and provide new records and SEM data for three previously described species.
By taking Flavalona gen. nov. out of Alona s.l. (Cladocera: Anomopoda: Chydoridae), the last major clade has now been removed from this polyphyletic assemblage. Flavalona gen. nov. is a monophylum defined by having three, rarely two connected head pores and slit-shaped, rarely rounded lateral head pores. Postabdomen rather long, distally narrowed, with robust marginal denticles and weakly developed lateral fascicles of setules. End-claw weakly curved and with short basal spine. Male postabdomen with gonopores opening at the end of a penis-like outgrowth. Trunk limbs: exopodite of P2 with seta; inner portion of P4 with flaming-torch shaped setae; P5 with filter plate of three setae; P6 a large simple lobe. The relationship of the new genus with other Aloninae remains to be determined. A key to the 11 species of the genus is provided and a discussion of their geographic distribution and habitat type is given.
Three new genera of Odontopygidae are described, all based on new species from the Udzungwa mountains, Tanzania, and all monotypic: Casuariverpa gen. nov. (type species: C. scarpa gen. et sp. nov.), Yia gen. nov. (type species: Y. geminispina gen. et sp. nov.), and Utiliverpa gen. nov. (type species: U. decapsulatrix gen. et sp. nov.). Similarities and differences between the new genera and other genera are discussed.
Knowledge about the taxonomy and distribution of the monogeneric polychaete family Longosomatidae Hartman, 1944 is limited in all parts of the world. A new species of the genus Heterospio Ehlers, 1874 (Annelida, Longosomatidae) is described from the Indian Ocean. Heterospio indica sp. nov. is mainly characterised by the conspicuous flattening of the distal half of capillary chaetae located in anterior elongated segments. Several body characters of high taxonomic relevance in the genus are examined under the SEM and discussed. The 18S SSU rDNA and COI genes were sequenced and represent the first sequencing of a species of Heterospio, and the sequences have been deposited in GenBank.
Two species of the genus Prionopetalum Attems, 1909, are recorded from the Udzungwa Mountains: P. asperginis sp. nov. and P. kraepelini (Attems, 1896). Prionopetalum stuhlmanni Attems, 1914, is synonymized under P. kraepelini. Odontopyge fasciata Attems, 1896, is transferred from Prionopetalum to Aquattuor Frederiksen, 2013, and new illustrations are given. A new illustrated key to species of Prionopetalum is provided.
The New World Clistopyga chaconi species group is revised. Eleven species are described as new: C. amazonica sp. nov., C. cinnamoptera sp. nov., C. cuscoensis sp. nov., C. hayesiana sp. nov., C. melanoptera sp. nov., C. misionensis sp. nov., C. mocaguae sp. nov., C. orellanae sp. nov., C. porteri sp. nov., C. rondoniae sp. nov. and C. yabuquensis sp. nov. Additional morphological data are provided for the previously known species, C. caramba Castillo & Sääksjärvi and C. chaconi Gauld. An illustrated identification key to all species of the group is provided. The Clistopyga chaconi species group appears to be most diverse at the Andean and Amazonian interface in western South America.
Two new species of Tetrastigma from Thailand, T. calcicola Kochaiph. & Trias-Blasi sp. nov. and T. jaichagunii C.L.Li ex Kochaiph. & Trias-Blasi sp. nov. are described and illustrated. Tetrastigma calcicola sp. nov. is a slender climber restricted to the open areas on limestone mountains at high elevation in the northern part of Thailand. The other species, T. jaichagunii sp. nov., is similar to T. harmandii Planch., but differs from it by having more densely verrucose young branches, broader leaflets, 4-lobed thick discs, bigger globose berries and oblongoid seeds. This species occurs along streams or in forest margins in evergreen forest and it is widely distributed in several parts of Thailand.
New leaf- and litter-dwelling species of the genus Pholcus from Southeast Asia (Araneae, Pholcidae)
(2016)
We describe eight new species of the genus Pholcus, and document their microhabitats. Four species are assigned to the previously described Pholcus ethagala group: P. tanahrata Huber sp. nov., P. uludong Huber sp. nov., and P. bukittimah Huber sp. nov. from the Malay Peninsula, and P. barisan Huber sp. nov. from Sumatra. These species are all litter-dwellers that build domed sheet webs on the undersides of large dead leaves on the ground. The other four species are assigned to newly created species groups: the P. tambunan group with two species from northern Borneo: P. tambunan Huber sp. nov. and P. bario Huber sp. nov.; and the P. domingo group with two species from the Philippines, Mindanao: P. domingo Huber sp. nov. and P. matutum Huber sp. nov. These latter four species are leafdwellers that build barely visible silk platforms tightly attached to the undersides of live leaves. The main rationale for this paper is to provide part of the taxonomic and natural history background for upcoming phylogenetic and evolutionary (microhabitat shifts) analyses.
The Southeast Asian Pholcus halabala species group is revised and re-delimited, based mainly on field observations (life color pattern, web design, position of egg-sac when carried by female, microhabitat) and ultrastructure (silk spigots, modifications of male cheliceral apophyses). The core group includes six leafdwelling species that have distinctive color patterns in life specimens (black and white or yellowish abdominal marks, dark pattern on posterior half of carapace) and build round to oval silk platforms on the undersides of leaves. Seven further species are tentatively assigned to the group pending further study. Several species originally assigned to the Pholcus halabala group are transferred to three newly proposed species groups, the Ph. krabi, Ph. buatong, and Ph. andulau groups. Nine species are newly described, four in the Ph. halabala group (Ph. khaolek Huber, sp. nov.; Ph. kuhapimuk Huber, sp. nov.; Ph. lintang Huber, sp. nov.; Ph. ubin Huber, sp. nov.); three in the Ph. krabi group (Ph. kipungit Huber, sp. nov.; Ph. krabi Huber, sp. nov.; Ph. narathiwat Huber, sp. nov.); one in the Ph. buatong group (Ph. buatong Huber, sp. nov.); and one in the Ph. andulau group (Ph. lambir Huber, sp. nov.). The females of Ph. satun Huber, 2011 and Ph. schwendingeri Huber, 2011 (both members of the buatong group) are newly described.
Two small, ground and litter-dwelling pholcid species from northern Borneo are described as representatives of a new genus, Hantu gen. nov.: H. kapit gen. et sp. nov. and H. niah gen. et sp. nov. Previous cladistic analyses suggested a closer relationship with the genera Savarna Huber, 2005 and Khorata Huber, 2005 (mainland Southeast Asia) than with the geographically closer genus Aetana Huber, 2005 (Borneo and Philippines to Fiji). Since the two species do not share any of the synapomorphies of Khorata and Savarna while having several synapomorphies on their own (ventral apophysis on male palpal coxa; male palpal trochanter apophysis with small teeth or scales; spines on male femora 1; high density of vertical hairs on male femora; presence of scape on epigynum), they are here proposed as representing a new genus.
We revise the Panjange nigrifrons group in Borneo and document an unexpected diversity in western Sarawak forests. Five species occur within 80 km from Kuching, each species being known from its type locality only. Further species occur east until Niah, but the genus seems to be absent from Sabah. We contrast this with another pholcid genus (Aetana Huber, 2005), which is diverse in Sabah and westward until Niah, but does not seem to occur in central and western Sarawak. Five species are newly described: Panjange kapit Huber, sp. nov., Panjange kubah Huber, sp. nov., Panjange niah Huber, sp. nov., Panjange pueh Huber, sp. nov., Panjange seowi Huber, sp. nov.; Panjange tahai (Huber, 2011) comb. nov. is transferred from Pholcus.
A new genus in the tribe Issini (Hemiptera: Fulgoromorpha: Issidae) is described from South China: Orbita Meng & Wang, gen. nov., its type species Orbita parallelodroma Meng & Wang, gen. et sp. nov. (China: Fujian) is described and illustrated. In addition, one new species of Neokodaiana Yang, N. minensis sp. nov. is described and illustrated from the same locality as the new genus. A key to the genera of Issini from China and neighbouring countries is provided.
The new genus Geotypodon gen. nov. is described. It includes two species from the Udzungwa Mountains: G. millemanus gen. et sp. nov. (type species) and G. submontanus gen. et sp. nov., one species from nearby Iringa: G. iringensis gen. et sp. nov., and 18 previously described species hitherto incorrectly assigned to Odontopyge Brandt, 1841.
Two species of Tomoceridae were found near Dalat, southern Vietnam. Tomocerus ocreatus Denis, 1948 is redescribed based on a neotype specimen. Previous records of Tomocerus ocreatus in non-type localities are reevaluated. A new species Tomocerina annamitica sp. nov. is described. The new species is mainly characterized by its small body size, pointed tenent hair, compound dental spines and the absence of intermediate teeth on mucro.
Two new species of the genus Sectonema from natural habitats of northern Vietnam are studied. This paper includes their descriptions, measurements, line illustrations, and light microscope (LM) and scanning electron microscope (SEM) pictures. Sectonema tropicum sp. nov. is characterized by a 2.56– 3.24 mm long body, 19–21 μm broad lip region, odontostyle 20–21 μm long at its ventral side, 730– 834 μm long neck, pharyngeal expansion occupying 52–59% of total neck length, uterus a simple tube-like structure 150–242 μm long or 1.2–2.5 times the body diameter, pars refringens vaginae present, V = 48–52, short (31–40 μm, c = 70–91, c’ = 0.5–0.6) and rounded tail, 91–97 μm long spicules, and only one weakly developed ventromedian supplement. Sectonema vietnamense sp. nov. is characterized by its slender (a = 33–49) and 2.71–4.25 mm long body, 14–16 μm broad lip region, odontostyle 8–9 μm long at its ventral side, 716–918 μm long neck, pharyngeal expansion occupying 63–67% of total neck length, uterus simple and 209–242 μm long or 2.5–2.9 times the corresponding body diameter, pars refringens vaginae absent, V = 54, short (34–39 μm, c = 70–115, c’ = 0.6–0.8) and rounded tail, 59–75 μm long spicules, and three or four irregularly spaced ventromedian supplements bearing hiatus. Both species are also characterized by their nearly continuous lip region, an atypical feature in this genus. Molecular analysis of S. tropicum sp. nov. confirms that Sectonema is a natural (monophyletic) taxon, very close to Metaporcelaimus.
The Lycocerus hanatanii species group is revised, with the addition of seven taxa: L. araticollis (Fairmaire, 1897), L. nigripennis (Pic, 1938), L. griseopubens (Pic, 1928), L. yitingi Hsiao & Okushima sp. nov., L. aurantiacus Hsiao & Okushima sp. nov., L. evangelium Hsiao & Okushima sp. nov. and L. kintaroi Hsiao & Okushima sp. nov. Supplementary descriptions of the males of L. araticollis and L. griseopubens are provided. Lycocerus nigripennis (Pic, 1938) and L. pictus (Wittmer, 1983) are redescribed in detail. Each species is provided with photos or illustrations of genitalia of both sexes and abdominal ventrite VII of the female if available. Distribution maps and a key to the species of the L. hanatanii species group are presented. In addition, the monophyly of the L. hanatanii species group is supported based on a morphological phylogenetic analysis.
Fifteen new species of Begonia L. from Sumatra are described and illustrated, in Begonia sect. Bracteibegonia (B. beludruvenea M.Hughes sp. nov. and B. jackiana M.Hughes sp. nov.), B. sect. Petermannia (B. harauensis Girm. sp. nov.), B. sect. Platycentrum (B. leuserensis M.Hughes sp. nov.), B. sect. Reichenheimia (B. fl uvialis M.Hughes sp. nov., B. halabanensis M.Hughes sp. nov., B. karangputihensis Girm. sp. nov., B. kemumuensis M.Hughes sp. nov., B. korthalsiana Miq. ex M.Hughes sp. nov., B. kudoensis Girm. sp. nov., B. lilliputana M.Hughes sp. nov., B. olivacea Ardi sp. nov., B. raoensis M.Hughes sp. nov., B. simolapensis Ardi sp. nov.) and B. sect. Sphenanthera (B. pseudoscottii Girm. sp. nov.). Using the International Union for the Conservation of Nature criteria, 6 are considered to be Least Concern, 5 Vulnerable and 4 Data Deficient. A key to 58 of the 63 currently accepted Begonia species in Sumatra is provided.
The taxonomy of the deep-water homolid crabs Moloha grandperrini Guinot & Richer de Forges, 1995 and M. alisae Guinot & Richer de Forges, 1995 is re-examined, and the types redescribed and figured. Moloha alisae is reported from South Africa for the first time. A new species with an inflated carapace, M. tumida sp. nov., is also described from southern India and compared with its closest congeners.
The genus Savarna Huber, 2005 was previously one of the most poorly known Pholcinae genera. Less than 20 specimens (representing four nominal species) were available worldwide; nothing was known about ultrastructure, natural history, or relationships. We present the first SEM data, supporting the position of the genus in Pholcinae outside the Pholcus group of genera and weakly suggesting a closer relationship with the genera Khorata Huber, 2005, Spermophorides Wunderlich, 1992, and two undescribed species of unknown affinity from Borneo. We provide the first data about microhabitat, web structure, and reaction to disturbance. We clarify the type locality of Savarna tessellata (Simon, 1901) (“Jalor, Biserat”) and describe topotypical material. We describe the previously unknown male of Spermophora miser Bristowe, 1952 and transfer the species (that was previously considered incertae sedis) to Savarna as Savarna miser (Bristowe, 1952) comb. nov. Savarna baso (Roewer, 1963) is newly synonymized with S. miser. We describe the most northern species in the genus, Savarna kaeo sp. nov., and provide amendments to the descriptions of all previously described species.
The genus Aquattuor Frederiksen, 2013 is revised. A. denticulatus Frederiksen, 2013 (type species) from the East Usambara Mts, Tanzania, is redescribed, and six new species are described: A. claudiahempae sp. nov. from Mt. Kilimanjaro, Tanzania, and five species from the Udzungwa Mts, Tanzania: A. longipala Enghoff sp. nov., A. major Enghoff sp. nov., A. stereosathe Enghoff sp. nov., A. submajor Enghoff sp. nov., and A. udzungwensis Enghoff sp. nov.
Four new species of the genus Caccothryptus (Coleoptera: Limnichidae) are described: C. taiwanus from Taiwan; C. orion from Okinawa; C. tibetanus and C. chayuensis from Tibet. All the species belong to the testudo species group (sensu Hernando & Ribera 2014). Additional specimen data and an updated species list are also given, and C. testudo Champion, 1923 is newly recorded from Thailand.
A new genus, Campodesmoides gen. nov., is described to only encompass C. corniger sp. nov., from Cameroon. This genus and species is distinguished from the few known species of the small western African family Campodesmidae, all currently in Campodesmus, by the much longer antennae and legs, the normal pore formula with ozopores borne on porosteles, and the suberect and distally twisted gonopod, coupled with peculiar horns on a few anterior postcollum segments. A new Campodesmus is also described, C. alobatus sp. nov., from Ivory Coast, which differs from congeners primarily in the lack of a dorsal/lateral lobe on the otherwise usual and strongly subcircular gonopod telopodite, albeit the latter is not directed mesad, but held subparallel to the main body axis.
A new species of Rhaphiostylis, R. minima sp. nov. from Liberia and Ivory Coast, is described and illustrated here. It is characterised by its comparatively small size, an ovate to lanceolate leaf blade with long drip tip and a partly glabrous ovary that is in shape in between those of R. elegans Engl. and R. preussii Engl. Rhaphiostylis elegans is lectotypified and illustrated here.
Based on intensive collecting from various sites in Sweden, the genus Dasyhelea Kieffer, 1911 was reviewed and the number of species now known from the country increased from five to twenty. Among the new species recorded there are two species described as new to science, D. dominiakae sp. nov. and D. gothlandica sp. nov., both in the subgenus Dicryptoscena Enderlein, 1936. The following subgenera are now documented from Sweden: Dasyhelea, Dicryptoscena, Pseudoculicoides Malloch, 1915, Prokempia Kieffer, 1913 and Sebessia Remm, 1979, the two latter subgenera being reported for the first time.
Two new species of jumping spiders, Thiratoscirtus oberleuthneri (♂) and Th. lamboji (♀), are described from Gabon, one of the least explored areas of the Afrotropics. Both species live in rainforest, at the forest fl oor. They are members of a very poorly known subfamily of salticids, the Thiratoscirtinae.
The new ctenid genus Afroneutria is proposed to include five species: Afroneutria velox (Blackwall, 1865) comb. nov. (type species), A. immortalis (Arts, 1912) comb. nov., A. erythrochelis (Simon, 1876) comb. nov., A. quadrimaculata sp. nov. and A. hybrida sp. nov., all collected in central to southeast Africa. Afroneutria species can be distinguished from the remaining Ctenidae by the presence of a synapomorphic laminar projection on the embolus tip. The genus can also be distinguished by the presence of the following set of characters: elongated cup-shaped median apophysis and elongated and laminar embolus on the male palp; large lateral projections on the epigynum and the spermathecae divided into two parts, dorsal and ventral, in the vulva.
The diagnosis of the Formicidae is revised, including five new, unreversed apomorphies, of which one is a unique synapomorphy. The first global male-based key to all subfamilies is provided and illustrated, and all ant subfamilies are diagnosed for males on a global scale for the first time. Three lineages of “basal ants” are assessed in detail: the Amblyoponinae, Leptanillinae, and Martialinae. The males of Martialis heureka (Martialinae) and Apomyrma (Amblyoponinae) are described. The Martialinae and Leptanillinae are diagnosed based on males, and additional diagnostic traits for the male of Amblyoponinae and worker of Martialis are provided. The placement of Scyphodon and Noonilla in the Formicidae and Leptanillinae is confirmed. Morphological characters of the Amblyoponinae, the Leptanillinae, and the Martialinae are contrasted, and potentially homologous apomorphies are signaled.
A new species of Paracrobeles, P. kelsodunensis sp. nov. is described from the Kelso Dunes area, Mojave National Preserve, southern California. Paracrobeles kelsodunensis sp. nov. is particularly characterised by a body length of 469–626 μm in females and 463–569 μm in males; lateral field with four incisures, extending almost to tail terminus; three pairs of asymmetrical lips, separated by U-shaped primary axils with two long guarding processes, each lip usually with four tines along its margin; three long labial probolae, deeply bifurcated, with slender prongs without tines; metastegostom with a strong anteriorly directed dorsal tooth; pharyngeal corpus anteriorly spindle-shaped, posteriorly elongate bulbous with dilated lumen; spermatheca 24–87 μm long; postvulval uterine sac 60–133 μm long; vulva in a sunken area; spicules 33–38 μm long; and male tail with a 5–8 μm long mucro. The generic diagnosis is emended on the basis of recently described species and a key to the species of Paracrobeles is provided.
We report here on recent collections of freshwater crabs from Antsiranana Province, northern Madagascar. The specimens belong to three species, one of which is new to science and is described here. This raises the number of species of freshwater crabs found in Madagascar to 17. All are endemic to the island and all belong to the Afrotropical family Potamonautidae Bott, 1970. The new species, Foza manonae sp. nov., is compared to the other species in this genus, and an updated key is provided. It is distinguished from the other three congeners by characters of the male first gonopod, sternum, carapace, and cheliped. The conservation status of the Malagasy freshwater crab fauna is summarized and discussed in light of the new material reported on here belonging to two other species, Madagapotamon humberti Bott, 1965 and Foza ambohitra Cumberlidge & Meyer, 2009.
A new Mexican species of Ochraethes Chevrolat, 1860 (Coleoptera, Cerambycidae, Cerambycinae, Clytini) is described: Ochraethes skillmani Wappes, Santos-Silva and Botero. Plocaederus mirim Martins and Monné, 2002 (Cerambycini) is redescribed and its female is figured for the first time. New geographical records in Plocaederus Dejean, 1835 are also provided.
A taxonomic revision of Panamanian species of the genus Dasymutilla Ashmead (Hymenoptera, Mutillidae) is presented and a key for the six species is given, all recognized from both sexes. Dasymutilla colorado Cambra, Williams and Quintero sp. nov., from central and eastern Panama, is described and illustrated. Sex associations permitted us to make the following five synonymies: D. sleipniri Manley and Pitts, 2007 (male) under D. phya (Cameron, 1895) (female); D. deyrollesi Mickel, 1937 (male) and Sphaerophthama [sic.] temaxensis Cameron, 1895 under Dasymutilla araneoides (Smith, 1862) (female); D. ionothorax Manley and Pitts, 2007 (male) under Dasymutilla spilota Manley and Pitts, 2007 (female); and D. guanacaste Manley and Pitts, 2007 (male) under D. paradoxa (Gerstaecker, 1874) (female). Seasonal flight activity for Dasymutilla from six years of continuous malaise trappings in Barro Colorado Island is presented.
An early, overlooked description of the genus Atelodesmis (Coleoptera, Cerambycidae, Lamiinae) is attributed to Chevrolat (in Duponchel and Chevrolat 1841), with A. mannerheimii Duponchel and Chevrolat, 1841 as its type species, and the genus redescribed. Atelodesmis Buquet, 1857, is a junior synonym and primary homonym of Atelodesmis Chevrolat. Atelodesmis hirticornis Buquet, 1857 and A. vestita Buquet, 1857 are synonymized with A. mannerheimii. Fallaxdesmis is described as a new genus with Atelodesmis unicolor Buquet, 1857 as type species. Atelodesmis piperita Bates, 1855 is transferred to Eupogonius and newly recorded for the state of Oaxaca, Mexico. A new species for Mexico and Guatemala, Eupogonius giesberti, is described. The following new combinations are established: Fallaxdesmis unicolor (Buquet, 1857) and Eupogonius piperita (Bates, 1855). Illustrations of A. mannerheimii, Fallaxdesmis unicolor, and the holotypes of A. hirticornis, A. vestita, A. unicolor, A. piperita and Eupogonius giesberti are included.
A new cryptic species, Aleochara (Xenochara) castaneimarmotae Klimaszewski, Webster, and Brunke, new species (Coleoptera: Staphylinidae: Aleocharinae), associated with Marmota monax (Linnaeus) burrows and caves in eastern North America, is described and illustrated. A key to Canadian species of subgenus Xenochara Mulsant and Rey and revised distributions of the taxonomically difficult fumata species group are provided. Aleochara quadrata Sharp is recorded from Washington and Oregon for the first time.
This paper revises the genus Ganelius Benesh, which is endemic to Madagascar, in the stag beetle tribe Figulini Burmeister (Coleoptera: Lucanidae: Lucaninae). Species in the genus are striking in their highly sexually dimorphic mandibles, a character that rarely occurs in the tribe. The genus was previously comprised of four species, but three of these names were problematic or questionably applied. A lectotype is designated for G. madagascariensis Laporte, a neotype is designated for Ganelius oberndorferi Nonfried, and Nigidius passaliformis Benesh is newly synonymized with G. oberndorferi. Two new species, G. gnamptus Paulsen and G. zombi Paulsen, are described from western Madagascar. The identity of the overlooked Ganelius nageli (Kriesche) is fixed through a neotype designation, and the species is moved to the new genus Agnelius, which is distinguished from Ganelius by a lack of sexual dimorphism, serrate protibiae, and a more flattened body.
Adesmus martinsi (Coleoptera, Cerambycidae, Lamiinae, Hemilophini), a new species from Bolivia, is described, illustrated, and included in a previous key. The new species displays gender dimorphism in the pubescent pattern and in anatomical structure. Thus, Adesmus becomes the second genus recorded in the Hemilophini to have visual chromatic dimorphism.
Lantana camara L., considered among the world’s worst invaders is in identity crisis and contentiously referred as Lantana camara L. (sensu lato). Taxonomic ambiguity in L. camara L. (sensu lato), a species complex is one of the grim caveats behind incompetence of its management efforts. Recognizing the extent of variability within the complex, we aim to highlight the need to circumscribe its composition to bring effective management and control efforts into practice. There is a need for clear terminology to examine weedy, naturalized and/or invasive complex constituents that have been placed under the contentious umbrella of 'L. camara L. (sensu lato)'. The time is ripe for invasion ecologists, cytogeneticists and conservationists to collaboratively focus on disentangling the complex and integrate their knowledge and expertise into management and control programs.
Two new genera and species of tiger beetles from Baltic amber (Coleoptera: Carabidae: Cicindelinae)
(2017)
Two fossil tiger beetle species (Coleoptera, Carabidae, Cicindelinae) are described from Eocene Baltic amber using light microscopic and X-ray microscopic techniques. Both species are considered representatives of the subtribe Iresina Rivalier, 1971 due to the shared combination of character states: glabrous head, six labral and four suborbital setae, and glabrous pronotum. Palaeopronyssiformia groehni Wiesner, Will, and Schmidt, new genus, new species, is characterized by a glabrous and furrowed head with six labral setae, large eyes, presence of two supraorbital setae on each side, mandibles with two teeth of the incisor region, and a glabrous and furrowed pronotum. Palaeoiresina cassolai Wiesner, Will, and Schmidt, new genus, new species, is characterized by a unicolored, undentated labrum, mandibles with two teeth of the incisor region, glabrous head with six labral setae, two clypeal setae, two supraorbital setae on each side, and a glabrous pronotum, mesepisternum, mesepimeron, and metepisternum. The species described here represent the only known tiger beetle fossils preserved in Baltic amber.
New Bolivian Rhinotragini (Coleoptera, Cerambycidae, Cerambycinae) are described: three species of Phygopoda Thomson, 1864 (P. longiscopifera sp. nov., P. boliviensis sp. nov. and P. chaquensis sp. nov.); and one species of Phygopoides Peñaherrera-Leiva and Tavakilian, 2007 (P. maxwelli sp. nov.). Two Brazilian species of Neophygopoda Melzer, 1933 are transferred to the genus Phygopoda: P. exilis (Melzer, 1933) comb. nov. and P. agdae (Martins, Galileo and Santos-Silva, 2015) comb. nov. All the species are illustrated, and a key to the Bolivian species of Phygopoda and host flower records are provided.
A primitive subfamily of false click beetles (Coleoptera: Eucnemidae: Phlegoninae) distributed primarily in the Neotropical region is revised. Euryphlegon gen. nov. is described from Belize in Central America. New species include: Phlegon chiriquiensis sp. nov. (Panama), Phlegon panamensis sp. nov. (Panama), Euryphlegon jacqueschassaini sp. nov. (Panama) and Euryphlegon parallelus sp. nov. (Belize). Phlegon herculeanus (Lacordaire) stat. res. is resurrected from synonymy with Phlegon buqueti Laporte. One new combination is proposed: Euryphlegon degallieri (Muona) (Phlegon). Based on a number of observed external character traits, Euryphlegon is placed in Orodotini Muona, 1993 within Macraulacinae Fleutiaux, 1922. Identifi cation keys are provided for species of Phlegon and Euryphlegon in the Neotropical region. The relationships among Phlegon, Euryphlegon, Euryptychus LeConte and other groups within Echthrogasterini Cobos, 1964 and Orodotini are discussed.
Abstract. Six new species of Coleoxestia Aurivillius, 1912 (Cerambycidae, Cerambycinae, Cerambycini) are described: C. clarkei from Bolivia; C. chemsaki and C. eyai from Peru; C. fragosoi and C. rafaeli from Brazil; and C. hovorei from Ecuador. Coleoxestia rachelae Eya and Chemsak is also newly recorded from Peru (new country record). Dorsal, ventral and lateral illustrations along with illustrations of various other structures are provided for each of the new species. In some cases, existing key couplets by Eya and Chemsak (2005) or Martins and Monné (2005) are modifi ed to help separate a new species from similar existing species. A reproduction of the general descriptive terminology for the head, prothorax and antennomeres of Coleoxestia, from Fragoso (1993), is also included in the illustrations.
The New World genus Chariessa Forster (Coleoptera: Cleroidea: Cleridae) is revised and includes C. catalina Opitz, new species, C. elegans Horn, C. dichroa (LeConte), C. floridana Schaeffer, C. pilosa (Forster), C. texana Wolcott, C. ramicornis Perty, C. vestita (Chevrolat), and C. duponti (Spinola). Enoplium pilosa var. marginata Say is synonymized with Chariessa pilosa Forster. Lectotypes are designated for C. pilosa (Forster), C. ramicornis Perty, and C. vestita (Chevrolat). Available information indicates that Chariessa adult and immature individuals are predatory on lignicolous insects with a particular affinity for cerambycids and buprestids that infest species of oak. It is postulated that Pleistocene speciation generated the North American components of Chariessa with more ancient southern species generated during the Middle Tertiary; after closures of the Middle American portals and orogeny of the South American Andes. Included in this treatise is a discussion of natural history, key to species, narratives of zoogeography and phylogeny, one diagram of a phylogenetic tree, 35 line drawings, eight SEM micrographs, twelve habitus photographs, nine photographs of male genitalia, and five distributional maps.
A detailed study of the holotype of Sphecomyrma canadensis Wilson, 1985 (Hymenoptera: Formicidae) from Canadian amber has led to the conclusion that the specimen belongs to a new genus, here named Boltonimecia gen.n. Since the taxonomy of stem-group ants is not well understood, in order to find the taxonomic position of this genus, it is necessary to review the classifi cation of stem-group ants in a study of their relation to crown-group ants. In the absence of data for traditional taxonomic approaches, a statistical study was done based on a morphometric analysis of antennae. Scape elongation is believed to play an important role in the evolution of eusociality in ants; however, this hypothesis has never been confirmed statistically. The statistical analysis presented herein lends support to the view that antennal morphology reliably distinguishes stem-group ants from crown-group ants, to determine whether a species belongs to one or the other group. This, in turn, may indicate a relationship exists between eusociality and scape elongation. A review of Cretaceous records of ants is made and the higher classification of Formicidae with definitions of stem and crown groups is proposed. Newly obtained data are discussed focusing particularly on the origin, evolution and diversity of ants.
Dipropus tequesta Johnson new species (Coleoptera: Elateridae) is described from southern Florida. Dipropus fuscus (LeConte) is a new synonym of D. soleatus (Say), and D. granosus (Fall) is a new synonym of D. asper (LeConte). The fl ightless female of D. asper is described and provides the fi rst report of brachyptery and endogean habits in Dipropus. A key to the species of the eastern United States and a new checklist of species for the country are provided.
Una nueva especie del género Eugnosta Hübner, E. lankinae n.sp. (Lepidoptera: Tortricidae), es descrita para la Provincia de Elqui, Chile. Se entregan los caracteres morfológicos del adulto y las fi guras de la genitalia masculina. Además se discute su posición sistemática con dos especies congéneres Neotropicales.
The taxonomic history of the rhinotragine genera Phygopoda Thomson, 1864 and Pseudophygopoda Tavakilian and Peñaherrera-Leiva, 2007 (Coleoptera: Cerambycidae: Cerambycinae) are discussed, and evidence is presented to suggest that some recent taxonomic changes made by Carelli and Monné (2015) were unjustified. Consequently, Phygopoda nigritarsis Gounelle, 1911 is moved to the genus Neophygopoda Melzer, 1933, creating the new combination Neophygopoda nigritarsis, the genera Panamapoda Clarke, 2014 and Paraphygopoda Clarke, 2014 are revalidated, and the species Paraphygopoda viridimicans (Fisher, 1952) and Paraphygopoda nappae Clarke, 2014 are also revalidated.
Chiquitano gen. nov. Chiquitano volcanesensis sp. nov., Compsibidion achiraensis sp. nov. and Compsibidion amboroensis sp. nov. (Coleoptera: Cerambycidae: Cerambycinae: Neoibidionini) are described from Bolivia. Notes on Rhysium Pascoe, 1866 and Rhysium bimaculatum Pascoe, 1866 are provided, and Brechmoidion separatum Martins and Galileo, 2007 is transferred to Rhysium. Keys to species of Compsibidion Thomson, 1864, Brechmoidion Martins, 1969 and Rhysium Pascoe, 1866 are also provided.
We describe a new butterfl y species, Anaeomorpha mirifi ca Simon and Willmott, n. sp. (Lepidoptera, Nymphalidae, Charaxinae), from premontane rain forest of the Chocó region of northwestern Ecuador. This represents the second known species and the fi rst record outside of the Amazon basin for this taxonomically and biologically enigmatic genus. Although the two species are not known to be sympatric, we identifi ed 21 characters in the external color pattern, wing shape and male genitalia that together support distinct species status. Most notably, the new species possesses an ocellus in the ventral hind wing tornus, a character which occurs in the Neotropical Charaxinae only in the genus Prepona Boisduval, 1836. A mean divergence of 6.8% in the COI ‘barcodes’ between the two species underlines their taxonomic distinctness.
A new species of Ctilodes Murray, 1864 (Coleoptera: Nitidulidae) was recovered in material from Vietnam during a large study of Carpophilinae. A description and detailed diagnosis of Ctilodes clinei Powell and Duffy new species is presented here along with a key to identify all currently known species of Ctilodes.
Species descriptions, keys to genera and species, and geographical distributions are presented for 43 species of the family Bruchidae (Coleoptera: Chrysomeloidea) for Chile. Of these species, seven are described as new:
Acanthoscelides aricae sp. nov., Lithraeus chillan sp. nov., L. comptus sp. nov., L. elguetai sp. nov., L. limari sp. nov., L. lonquimay sp. nov., and L. penai sp. nov. Eight species are endemic to Chile. A list of true host plants and floral records for those with known host associations is presented. Habitus photographs and drawings of pertinent body parts, including male genitalia, are provided. References pertaining to the previously described species are listed.
A new genus and two new species of Argentine Rhinotragini
(Coleoptera, Cerambycidae, Cerambycinae)
(2017)
Rhopalessa irwini sp.nov. and Rhinion parkeri gen. nov., sp. nov. (Coleoptera, Cerambycidae, Cerambycinae, Rhinotragini) are described from Argentina. A key to species of Rhopalessa is provided.
The female of Callichroma magnifi cum Napp and Martins, 2009 (Coleoptera: Cerambycidae: Cerambycinae: Callichromatini) is described and illustrated for the fi rst time. The geographical distribution of the species is expanded to the Colombian Caribbean and information on the collection site is provided.
A new species of Autostichidae, Eraina beechei nov. sp., is described from specimens collected in the Andean mountains of Ñuble, Biobío. Diagnostic characters for the species are presented; photographs of adults, illustrations of male and female genitalia, and a key for identifi cation of Eraina species are provided.
The species of Dipropus Germar (Coleoptera: Elateridae) of the west-central region of North America are taxonomically reviewed. Historical records of D. approximatus (Candèze), D. ferreus (LeConte), D. simplex (LeConte) and D. soleatus (Say) in the region are based on misidentifi cations. Dipropus pericu new species is described from Baja California Sur. Dipropus reinae new species, D. sonora new species and D. yaqui new species are described from southern Arizona, New Mexico and Sonora. Dipropus warneri new species is described from Arizona, New Mexico and Texas. Dipropus sus (Candèze) is reported from Morelos, Sonora and Sinaloa as new state records. Keys to the species of the region are provided. The historical value of a specimen of Elater soleatus Say in the Dejean collection is discussed and its lectotype designated. Ischiodontus oblitus Candèze is treated as an objective synonym of Dipropus soleatus.
Two new species of Eburiini (Coleoptera, Cerambycidae, Cerambycinae) are described from Mexico (Guerrero): Susuacanga boteroi; and Eburia (Eburia) girouxae. Both are included in previous keys.
Epimelitta Bates, 1870 (Coleoptera, Cerambycidae) is redescribed with two species: Epimelitta scoparia (Klug, 1825) and Epimelitta rufiventris Bates, 1870; Epimelitta meliponica Bates, 1870 syn. nov. and Epimelitta acutipennis Fisher, 1947 syn. nov. are considered junior synonyms of E. scoparia. Exepimelitta gen. nov. is described with five species: Exepimelitta mimica (Bates, 1873), Ex. nigerrima (Bates, 1892), Ex. consobrina (Melzer, 1931) (=Epimelitta nigerrima var. flavipubescens Fisher, 1947, syn. nov.), Ex. lestradei (Peñaherrera-Leiva and Tavakilian, 2003) and Ex. windsori sp. nov. Charisia Champion, 1892 is revalidated and redescribed with six species: Charisia euphrosyne (Newman, 1840), C. mneme (Newman, 1841), C. melanaria Gounelle, 1911, C. ornaticollis Zajciw, 1973, C. bleuzeni (Peñaherrera-Leiva and Tavakilian, 2003) and C. durantoni (Peñaherrera-Leiva and Tavakilian, 2003).
Erratamelitta gen. nov. is described with two species: Erratamelitta erato (Newman, 1840) (= Epimelitta bicolor (Bates, 1873), syn. nov.) and Er. eliasi sp. nov.. Adepimelitta gen. nov. is described with two species: Adepimelitta debilis (Gounelle, 1911) and Ad. eupheme (Lameere, 1884). A brief synopsis of the genus Acorethra Bates, 1873 is presented, with two species included: Acorethra chrysaspis Bates, 1873, revalidated, and Ac. aureofasciata Gounelle, 1911. All species are illustrated (including genitalia when available); and keys to the genera, and their species, are provided.
Taxonomic supplement (2001 to 2015) to the catalogue of New Zealand Carabidae (Insecta: Coleoptera)
(2016)
A supplement to the “Carabidae (Insecta: Coleoptera): catalogue” published by Larochelle and Larivière (2001; Fauna of New Zealand 43), is provided. A total of 99 genera and 547 species and subspecies belonging to 21 tribes and 8 subfamilies of Carabidae (including Cicindelini and Rhysodini), is recorded for New Zealand. Endemism is high, with 93% of the species and subspecies as well as 62% of the genera recorded from this country only. Changes to the 2001 catalogue are documented. The synonymy and type locality of taxa described between 2001 and 2015 are also provided.
The description of a new species of Kessleria (Lepidoptera: Yponomeutidae) is presented, based on male and female specimens obtained from laboratory rearing of material collected on their host, Maytenus boaria, of the central region of Chile.
Two Neoibidionini from Bolivia are described as new: Neocompsa schneppi sp. nov. and Tropidion nancyae sp. nov. The new species are compared to closely related species and previous keys by Martins and Galileo (2007, 2009) are modifi ed to include them. Illustrations of dorsal, ventral and lateral habitus, as well as head structures are included for both species. Comments are also included on color and pattern variation of each species.