590 Tiere (Zoologie)
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The blind and mostly endogean julid genus Stygiiulus Verhoeff, 1929 stat. nov. is reviewed to include ten species: S. ausugi (Manfredi, 1953) comb. nov., S. fimbriatus (Strasser, 1971) comb. et stat. nov., S. gentianae (Strasser, 1971) comb. et stat. nov., S. illyricus (Verhoeff, 1929) comb. nov., S. insularis (Strasser, 1938) comb. nov., S. maximus (Verhoeff, 1929) comb. nov., S. montellensis (Verhoeff, 1930) comb. nov., S. rotundatus (Strasser, 1962) comb. et stat. nov., S. seewaldi (Strasser, 1967) comb. nov., and S. tobias (Berlese, 1886) comb. nov. The distinctiveness of the genus from Typhloiulus is proven by both morphological and molecular data. The monotypic genus Alpityphlus Strasser, 1967 is here treated as a junior subjective synonym of Stygiiulus syn. nov. Stygiiulus fimbriatus comb. et stat. nov. and S. gentianae comb. et stat. nov., originally described as subspecies of S. ausugi comb. nov., are here erected to full species, while the subspecies S. illyricus stygis (Verhoeff, 1933) comb. et syn. nov. is considered to be a junior subjective synonym of the typical S. illyricus comb. nov. Lectotypes are formally designated for S. gentianae comb. et stat. nov. and S. rotundatus comb. et stat. nov., to stabilize the nomenclature of the two species under Article 74.1 of the ICZN. The phylogeny of the genus, its distribution patterns, and the modified mouthparts in some of its species are discussed.
The Caucasian leucogeorgiinine genera Archileucogeorgia Lohmander, 1936 and Leucogeorgia Verhoeff, 1930 are revised, with Leucogeorgia being considered as a senior subjective synonym of Archileucogeorgia, syn. nov. The following new combinations are thus warranted: Leucogeorgia abchasica (Lohmander, 1936) and L. satunini (Lohmander, 1936), both comb. nov. ex Archileucogeorgia. All four previously described species, viz., L. longipes Verhoeff, 1930, L. abchasica, L. satunini and L. rediviva Golovatch, 1983, are redescribed based on new material, partly also on the type series, with a lectotype being designated for L. longipes. In addition, eleven new species of Leucogeorgia are described, both with normal (L. borealis sp. nov., L. gioi sp. nov., L. golovatchi sp. nov., L. lobata sp. nov., L. oculata sp. nov. and L. prometheus sp. nov.) and modified mouthparts (L. caudata sp. nov., L. mystax sp. nov., L. profunda sp. nov., L. redivivoides sp. nov. and L. turbanovi sp. nov., all clearly troglobionts). Additionally, a new monotypic genus, Martvilia gen. nov., is erected, with M. parva gen. et sp. nov. as the type species, another presumed troglobiont. An identification key to both genera and all 16 species of Caucasian Leucogeorgiini is presented. Several other members of this tribe are also re-examined, including a syntype male of Telsonius nycteridonis Strasser, 1976, from Greece, herewith designated as the lectotype. Troglomorphisms and mouthpart modifications, as well as the distributions and relationships within both Leucogeorgia and Leucogeorgiini, and a Leucogeorgiini species richness estimate for the western Caucasus are discussed.