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620
619
Nine species of Navasoleon (Neuroptera: Myrmeleontidae: Nemoleontini) are recognized in the present work with the description of seven new species: N. amazonas Stange, N. brasiliensis Miller, N. egeri Stange, N. lithophilus Miller, N. lotti Stange, N. tarsalis Miller, and N. venezolanus Stange. Navasoleon bosqui (Navás) is designated a nomen dubium. A key to the species is provided as well as to the genera of Nemoleontini with closing pretarsal claws. The larvae of two species are described and keyed with biological notes. Biological data presented in Miller and Stange (1985), erroneously attributed to N. bolivianus (Banks), has now been corrected to N. lithophilus. Included in this treatise are 71 color photographs including photos of male and female terminalia, male genitalia, and two species of larvae.
618
A new species of leaf insect, Phyllium (Phyllium) letiranti Cumming and Teemsma, new species (Phasmida: Phylliidae), is described from a series of males, females, and eggs from Peleng Island, Indonesia. This new species is the first record of the family Phylliidae on the island and is here differentiated from congeners. Keys to males, females, and eggs of the Phyllium species of Sulawesi and Peleng islands are included within.
617
Hippodamia variegata (Goeze) (Coleoptera: Coccinellidae), a Palearctic lady beetle that has become established in North America, is reported for the first time from the state of Iowa, U.S.A. The earliest specimen of H. variegata from Iowa was collected in 2009, with two subsequent specimens in 2012. Collection records of an advancing, non-native species like H. variegata are important for determining comparative rates of geographic spread across various regions and understanding its invasion dynamics.
616
Based on morphological features of the adult female, the Brazilian soft scale, Mesolecanium ferum Hempel, 1920 (= Toumeyella ferum) (Hemiptera: Coccomorpha: Coccidae) is transferred to the genus Neotoumeyella Kondo and Williams, 2009, as Neotoumeyella ferum (Hempel), comb. nov. The genus Neotoumeyella is rediagnosed in order to accommodate the unique features of M. ferum and an updated key to the six species of the genus is provided. An English translation of the Spanish redescription of M. ferum (as T. ferum) by Granara de Willink (2012) is also provided, with the author’s comments and interpretations.
615
A combination of five characters supports the arrangement of three species in the Anthocharis sara complex (Lepidoptera: Pieridae: Pierinae: Anthocharidini) consisting of Anthocharis sara Lucas, 1852 (Sara orangetip), Anthocharis thoosa Scudder, 1878 (southwestern orangetip), and Anthocharis julia W.H. Edwards, 1872 (Julia orangetip). These five characters include adult phenotypes, fifth instar larval coloration, pupal cone curvature and color, number of overwintering cycles, and interspecific contact zone behavior. This taxonomic treatment is generated from extensive life history studies from 12 western U.S. states through most of the sara complex distribution.
614
The Republic of Panama currently has 358 recorded species of Trichoptera. Herein we add 25 new country records for Panama. The newly recorded taxa increase Panama’s total known caddisfly fauna to 383 species, distributed among 15 families and 52 genera. These results are part of an ongoing effort to characterize the caddisfly fauna of Panama, and to evaluate that country’s major watersheds (cuencas).
613
There have been 136 species of microcaddisflies (Trichoptera: Hydroptilidae) recorded from the Republic of Panama. Herein we describe seven new species from the Mount Totumas Cloud Forest and Biological Reserve, in the upper reaches of the Río Chiriquí Viejo watershed (Costatrichia devestiva, C. dietrichi, Metrichia brocha, M. calla, Neotrichia atopa, Ochrotrichia anticheirion, and Rhyacopsyche totuma). These results are part of an ongoing effort to characterize the aquatic insect fauna of Panama, and to evaluate that country’s major watersheds.
612
Through this survey, four species of aphelinid, one species of encyrtid, and one species of signiphorid wasp (Hymenoptera: Chalcidoidea) were collected and identified as follows: Aphytis japonicus DeBach and Azim, Encarsia berlesei (Howard), Marietta carnesi (Howard), Pteroptrix sp. 2, Arrhenophagus chionaspidis Aurivillius, and Chartocerus subaenus (Förster). Among these, C. subaenus is recorded newly from the Korean parasitoid fauna. Also, three species of aphelinids and one species of signiphorid are newly added as parasitic wasps associated with Pseudaulacaspis cockerelli (Cooley) (Hemiptera: Coccoidea) from Korea. In this paper, the list of parasitoid species of P. cockerelli that occur in Korea is updated and a brief diagnosis and photographs of these species are provided.
611
The Scarabaeus subgenus Scarabaeolus Balthasar, 1965 (Scarabaeidae: Scarabaeinae: Scarabaeini) is defined and nine new species are described, six from the Republic of South Africa [S. (S.) fragilis sp. n., S. (S.) krugeri sp. n., S. (S.) lizleri sp. n., S. (S.) orientalis sp. n., S. (S.) rugosipennis sp. n., S. (S.) similis sp. n.] and one each from Angola [S. (S.) cunene sp. n.], Namibia [S. (S.) namibensis sp. n.] and Kenya [S. (S.) werneri sp. n.], bringing the number of recorded species up to 41. Species accounts listing original descriptions, subsequent accounts, type localities, type repositories, and geographic distributions are provided for all the species. The status of S. (S.) reichei Waterhouse, 1890 and of S. (S.) bohemani Harold, 1868, both formerly synonymized and reinstated, are discussed. It is concluded that S. (S.) reichei is synonymous with S. (S.) canaliculatus Fairmaire, 1888, whereas S. (S.) bohemani differs from S. (S.) palemo Olivier, 1789 in the color of antennal club and shape of paramere tips, and is upheld pending availability of molecular data.
610
609
A new Mexican species of Ochraethes Chevrolat, 1860 (Coleoptera, Cerambycidae, Cerambycinae, Clytini) is described: Ochraethes skillmani Wappes, Santos-Silva and Botero. Plocaederus mirim Martins and Monné, 2002 (Cerambycini) is redescribed and its female is figured for the first time. New geographical records in Plocaederus Dejean, 1835 are also provided.
608
A taxonomic revision of Panamanian species of the genus Dasymutilla Ashmead (Hymenoptera, Mutillidae) is presented and a key for the six species is given, all recognized from both sexes. Dasymutilla colorado Cambra, Williams and Quintero sp. nov., from central and eastern Panama, is described and illustrated. Sex associations permitted us to make the following five synonymies: D. sleipniri Manley and Pitts, 2007 (male) under D. phya (Cameron, 1895) (female); D. deyrollesi Mickel, 1937 (male) and Sphaerophthama [sic.] temaxensis Cameron, 1895 under Dasymutilla araneoides (Smith, 1862) (female); D. ionothorax Manley and Pitts, 2007 (male) under Dasymutilla spilota Manley and Pitts, 2007 (female); and D. guanacaste Manley and Pitts, 2007 (male) under D. paradoxa (Gerstaecker, 1874) (female). Seasonal flight activity for Dasymutilla from six years of continuous malaise trappings in Barro Colorado Island is presented.
607
Se registra por primera vez el ácaro rojo de las palmas, Raoiella indica Hirst (Acari: Tenuipalpidae), en Guatemala. Este ácaro invasivo se encontró en los departamentos de Petén e Izabal infestando palma de coco (Cocos nucifera), palma de Navidad (Adonidia merrillii) y palmera robeleni (Phoenix roebelenii).
606
En este estudio se describen dos nuevas especies de hemípteros del género Aleuropleurocelus (Hemiptera: Aleyrodidae): Aleuropleurocelus mexicanus Carapia y Sánchez sp. nov., encontrada en los estados de Coahuila, México, Puebla, Sonora, y Tlaxcala en hojas de Quercus agrifolia Née y Quercus sp., y Aleuropleurocelus anahuac Carapia y Sánchez sp. nov. encontrada en los estados de México, Morelos y Puebla en el envés de hojas de Quercus. Se proporcionan microfotografías de estructuras morfológicas de puparios y se discute la separación con otras especies ovales de Aleuropleurocelus.
605
An early, overlooked description of the genus Atelodesmis (Coleoptera, Cerambycidae, Lamiinae) is attributed to Chevrolat (in Duponchel and Chevrolat 1841), with A. mannerheimii Duponchel and Chevrolat, 1841 as its type species, and the genus redescribed. Atelodesmis Buquet, 1857, is a junior synonym and primary homonym of Atelodesmis Chevrolat. Atelodesmis hirticornis Buquet, 1857 and A. vestita Buquet, 1857 are synonymized with A. mannerheimii. Fallaxdesmis is described as a new genus with Atelodesmis unicolor Buquet, 1857 as type species. Atelodesmis piperita Bates, 1855 is transferred to Eupogonius and newly recorded for the state of Oaxaca, Mexico. A new species for Mexico and Guatemala, Eupogonius giesberti, is described. The following new combinations are established: Fallaxdesmis unicolor (Buquet, 1857) and Eupogonius piperita (Bates, 1855). Illustrations of A. mannerheimii, Fallaxdesmis unicolor, and the holotypes of A. hirticornis, A. vestita, A. unicolor, A. piperita and Eupogonius giesberti are included.
604
The Trichoptera of Panama V. Descriptions of new species, new country records, and a synonymy
(2018)
The Río Chiriqui basin is one of 52 major watershed areas, or cuencas, in the Republic of Panama. It occurs in western Panama, discharges into the Pacific Ocean, and includes portions of Volcán Barú on its northern extent. The Río Caldera occupies the northernmost subbasin of this basin. Two of its tributaries, Quebrada Grande and Quebrada Jaramillo, occur in close proximity and on opposite sides, and have different recent geologic histories and current land use patterns. During the course of investigating the caddisfly fauna of these two drainages, three new species of the microcaddisfly genus Neotrichia Morton (Trichoptera: Hydroptilidae) were identified: N. collierorum and N. anzuelo from Quebrada Jaramillo and N. tatianae from Quebrada Grande. These are described and figured herein. We also add one new genus (Rhyacopsyche Mueller) and five new country records (Hydroptila paschia Mosely, Metrichia ancora Bueno-Soria and Holzenthal, Ochrotrichia jolandae BuenoSoria and Holzenthal, Rhyacopsyche obliqua Flint, and Chimarra (Curgia) maritza Flint) for Panama. Finally, we designate Ochrotrichia abrelata Harris and Armitage, 2015 as a junior synonym of Ochrotrichia jolandae Bueno-Soria and Holzenthal, 2008. This synonymy and the newly recorded species and country records increase Panama’s known trichopteran fauna to 350 species, distributed among 15 families and 52 genera.
603
After publication of Blahnik and Holzenthal (2017), it was noticed that a large portion of the text had been accidentally removed from the "Phylogenetic and evolutionary comments" section during the proofing stage. The beginning of the deleted section completes the sentence on line 6 of page 129, which begins "The species included in the subgenus...". The Insecta Mundi editorial staff apologizes for this oversight. In order to provide context for the deleted excerpt, the entire "Phylogenetic and evolutionary comments" section is reproduced here, with the deleted text reincorporated. Insecta Mundi has also released a revised version of the Blahnik and Holzenthal (2017) manuscript, with this error corrected. However, the revised version is merely for convenience, and not an official peerreviewed article. Anyone wishing to reference the findings of Blahnik and Holzenthal (2017) should cite the original 2017 manuscript or this erratum. The references and figure plates cited in this section have also been reproduced here. ...
602 [korr. Vers.]
Species of Mortoniella are revised for the northern and Andean part of the South American continent, including the countries of Bolivia, Peru, Ecuador, Colombia, Venezuela, and Guyana. All previously described species from the region are reillustrated and redescribed, except for Mortoniella santiaga Sykora, 1999 and M. quinuas Harper and Turcotte, 1985, whose types could not be located, and M. tranquilla Martynov, 1912, whose type is based on a female specimen and thus is currently unidentifiable. Included in the revision are 35 described species and 59 new species. Mortoniella similis Sykora, 1999 is considered a junior synonym of M. roldani Flint, 1991, and M. macuta (Botosaneanu, 1998) is considered a junior synonym of M. limona (Flint, 1981). A new subgenus, Nanotrichia, is recognized to accommodate species previously referred to as members of the ormina and velasquezi groups. Mexitrichia pacuara Flint, 1974 is designated the type species for the subgenus. Species previously referred to as members of the bilineata and leroda species groups are retained in the nominate subgenus, along with additional taxa not previously placed to species group, and treated within a number of subgroups. Previously described species of M. (Mortoniella) which are redescribed and reillustrated include: M. angulata Flint, 1963; M. apiculata Flint, 1963; M. atenuata (Flint, 1963); M. bifurcata Sykora, 1999; M. bilineata Ulmer, 1906; M. bolivica (Schmid, 1958); M. chicana Sykora, 1999; M. denticulata Sykora, 1999; M. elongata (Flint, 1963); M. enchrysa Flint, 1991; M. flinti Sykora, 1999; M. foersteri (Schmid, 1964); M. hodgesi Flint, 1963; M. iridescens Flint, 1991; M. leei (Flint, 1974); M. limona (Flint, 1981); M. marini (Rueda Martín and Gibon, 2008); M. paralineata Sykora, 1999; M. paraenchrysa Sykora, 1999; M. pocita (Flint, 1983); M. punensis (Flint, 1983); M. roldani Flint, 1991; M. simla (Flint, 1974); M. spinulata (Flint, 1991); M. squamata Sykora, 1999; M. unilineata Sykora, 1999; and M. wygodzinskii (Schmid, 1958). New species described in M. (Mortoniella), followed by their respective areas of distribution, include: M. acutiterga (Ecuador); M. adamsae (Peru); M. akrogeneios (Ecuador); M. applanata (Peru); M. auricularis (Colombia); M. barinasi (Venezuela); M. biramosa (Venezuela); M. bothrops (Peru); M. brevis (Ecuador, Venezuela); M. bulbosa (Peru); M. catherinae (Peru); M. chalalan (Peru); M. cornuta (Peru); M. cressae (Venezuela); M. croca (Peru); M. curtispina (Venezuela); M. curvistylus (Ecuador); M. dentiterga (Ecuador); M. dinotes (Peru); M. draconis (Ecuador); M. emarginata (Ecuador, Colombia); M. esrossi (Colombia); M. flexuosa (Colombia); M. furcula (Ecuador); M. gilli (Ecuador); M. gracilis (Venezuela); M. grandiloba (Venezuela); M. guyanensis (Guyana); M. hamata (Colombia); M. langleyae (Ecuador); M. longiterga (Ecuador); M. membranacea (Bolivia); M. monopodis (Colombia, Ecuador); M. parameralda (Ecuador); M. pica (Ecuador); M. proakantha (Ecuador); M. prolata (Peru); M. quadrispina (Ecuador); M. rectiflexa (Ecuador); M. ruedae (Bolivia); M. schlingeri (Colombia); M. silacea (Colombia, Ecuador); M. sinuosa (Bolivia, Peru); M. spatulata (Venezuela); M. tanyrhabdos (Venezuela); M. tridens (Peru); M. triramosa (Bolivia); M. tusci (Venezuela); and M. variabilis (Venezuela, Colombia). Species assigned to the subgenus M. (Nanotrichia) which are redescribed and reillustrated include: M. aequalis (Flint, 1963); M. aries (Flint, 1963); M. collegarum (Rueda Martín and Gibon, 2008); M. eduardoi (Rueda Martín and Gibon, 2008); M. macarenica (Flint, 1974); M. pacuara (Flint, 1974); M. usseglioi (Rueda Martín and Gibon, 2008); and M. velasquezi (Flint, 1991). Previously described species of Mortoniella, outside the area of coverage, that are transferred to the subgenus M. (Nanotrichia) include: M. alicula Blahnik and Holzenthal, 2011; M. bocaina Blahnik and Holzenthal, 2011; M. catarinensis (Flint, 1974); M. froehlichi Blahnik and Holzenthal, 2011; M. ormina (Mosely, 1939); M. rodmani Blahnik and Holzenthal, 2008; and M. tripuiensis Blahnik and Holzenthal, 2011. New species in the subgenus M. (Nanotrichia), followed by their respective areas of distribution, include: Mortoniella cognata (Ecuador, Venezuela); M. coheni (Ecuador); M. licina (Ecuador); M. paucispina (Peru); M. quadridactyla (Venezuela); M. simplicis (Venezuela); M. spangleri (Ecuador); M. triangularis (Ecuador); M. venezuelensis (Venezuela); and M. zamora (Ecuador). A key to the males of species from the region is also provided, as well as a key to females for the major subgroups and a species key to females of the velasquezi group. Finally, a partially resolved phylogeny of the species is presented, along with a discussion of evolutionary trends within the genus.
602 [1. Version]
Species of Mortoniella are revised for the northern and Andean part of the South American continent, including the countries of Bolivia, Peru, Ecuador, Colombia, Venezuela, and Guyana. All previously described species from the region are reillustrated and redescribed, except for Mortoniella santiaga Sykora, 1999 and M. quinuas Harper and Turcotte, 1985, whose types could not be located, and M. tranquilla Martynov, 1912, whose type is based on a female specimen and thus is currently unidentifiable. Included in the revision are 35 described species and 59 new species. Mortoniella similis Sykora, 1999 is considered a junior synonym of M. roldani Flint, 1991, and M. macuta (Botosaneanu, 1998) is considered a junior synonym of M. limona (Flint, 1981). A new subgenus, Nanotrichia, is recognized to accommodate species previously referred to as members of the ormina and velasquezi groups. Mexitrichia pacuara Flint, 1974 is designated the type species for the subgenus. Species previously referred to as members of the bilineata and leroda species groups are retained in the nominate subgenus, along with additional taxa not previously placed to species group, and treated within a number of subgroups. Previously described species of M. (Mortoniella) which are redescribed and reillustrated include: M. angulata Flint, 1963; M. apiculata Flint, 1963; M. atenuata (Flint, 1963); M. bifurcata Sykora, 1999; M. bilineata Ulmer, 1906; M. bolivica (Schmid, 1958); M. chicana Sykora, 1999; M. denticulata Sykora, 1999; M. elongata (Flint, 1963); M. enchrysa Flint, 1991; M. flinti Sykora, 1999; M. foersteri (Schmid, 1964); M. hodgesi Flint, 1963; M. iridescens Flint, 1991; M. leei (Flint, 1974); M. limona (Flint, 1981); M. marini (Rueda Martín and Gibon, 2008); M. paralineata Sykora, 1999; M. paraenchrysa Sykora, 1999; M. pocita (Flint, 1983); M. punensis (Flint, 1983); M. roldani Flint, 1991; M. simla (Flint, 1974); M. spinulata (Flint, 1991); M. squamata Sykora, 1999; M. unilineata Sykora, 1999; and M. wygodzinskii (Schmid, 1958). New species described in M. (Mortoniella), followed by their respective areas of distribution, include: M. acutiterga (Ecuador); M. adamsae (Peru); M. akrogeneios (Ecuador); M. applanata (Peru); M. auricularis (Colombia); M. barinasi (Venezuela); M. biramosa (Venezuela); M. bothrops (Peru); M. brevis (Ecuador, Venezuela); M. bulbosa (Peru); M. catherinae (Peru); M. chalalan (Peru); M. cornuta (Peru); M. cressae (Venezuela); M. croca (Peru); M. curtispina (Venezuela); M. curvistylus (Ecuador); M. dentiterga (Ecuador); M. dinotes (Peru); M. draconis (Ecuador); M. emarginata (Ecuador, Colombia); M. esrossi (Colombia); M. flexuosa (Colombia); M. furcula (Ecuador); M. gilli (Ecuador); M. gracilis (Venezuela); M. grandiloba (Venezuela); M. guyanensis (Guyana); M. hamata (Colombia); M. langleyae (Ecuador); M. longiterga (Ecuador); M. membranacea (Bolivia); M. monopodis (Colombia, Ecuador); M. parameralda (Ecuador); M. pica (Ecuador); M. proakantha (Ecuador); M. prolata (Peru); M. quadrispina (Ecuador); M. rectiflexa (Ecuador); M. ruedae (Bolivia); M. schlingeri (Colombia); M. silacea (Colombia, Ecuador); M. sinuosa (Bolivia, Peru); M. spatulata (Venezuela); M. tanyrhabdos (Venezuela); M. tridens (Peru); M. triramosa (Bolivia); M. tusci (Venezuela); and M. variabilis (Venezuela, Colombia). Species assigned to the subgenus M. (Nanotrichia) which are redescribed and reillustrated include: M. aequalis (Flint, 1963); M. aries (Flint, 1963); M. collegarum (Rueda Martín and Gibon, 2008); M. eduardoi (Rueda Martín and Gibon, 2008); M. macarenica (Flint, 1974); M. pacuara (Flint, 1974); M. usseglioi (Rueda Martín and Gibon, 2008); and M. velasquezi (Flint, 1991). Previously described species of Mortoniella, outside the area of coverage, that are transferred to the subgenus M. (Nanotrichia) include: M. alicula Blahnik and Holzenthal, 2011; M. bocaina Blahnik and Holzenthal, 2011; M. catarinensis (Flint, 1974); M. froehlichi Blahnik and Holzenthal, 2011; M. ormina (Mosely, 1939); M. rodmani Blahnik and Holzenthal, 2008; and M. tripuiensis Blahnik and Holzenthal, 2011. New species in the subgenus M. (Nanotrichia), followed by their respective areas of distribution, include: Mortoniella cognata (Ecuador, Venezuela); M. coheni (Ecuador); M. licina (Ecuador); M. paucispina (Peru); M. quadridactyla (Venezuela); M. simplicis (Venezuela); M. spangleri (Ecuador); M. triangularis (Ecuador); M. venezuelensis (Venezuela); and M. zamora (Ecuador). A key to the males of species from the region is also provided, as well as a key to females for the major subgroups and a species key to females of the velasquezi group. Finally, a partially resolved phylogeny of the species is presented, along with a discussion of evolutionary trends within the genus.
601
Genera of Cephaloscymnini (Coleoptera: Coccinellidae: Coccidulinae) are discussed and a key to all recognized genera and species is provided. Succinctonotum, new genus, is proposed. Scymnus laboulbenii Mulsant and Prodilis maculata Weise are transferred to Neaporia as new combinations. Prodiloides bipunctata Weise, and Neaporia compta are transferred to Prodilis as new combinations. Neaporia cuprea Gorham is considered a junior synonym of Neaporia viridiscens Gorham and Cephaloscymnus bruchi Weise a junior synonym of Prodilis volgus Mulsant. New species described in Cephaloscymnus are C. beulah, C. candice, C. juanita. New species described in Neaporia are N. becky, N. bobbie, N. brandy, N. carole, N. cassandra, N. christy, N. daisy, N. deanna, N. dianne, N. felicia, N. gwendolyn, N. hilda, N. irma, N. jennie, N. jenny, N. kay, N. kayla, N. kristine, N. leah, N. lena, N. leona, N. longifrons, N. mabel, N. mae, N. margie, N. marsha, N. miriam, N. misty, N. myrtle, N. naomi, N. nina, N. nora, N. olga, N. opal, N. patsy, N. penny, N. priscilla, N. shelley, N. sonia, N. tracey, and N. violet. New species described in Succinctonotum is S. frosti. New species described in Prodilis are P. ada, P. alberta, P. alison, P. amelia, P. angie, P. araguaensis, P. bartletti, P. belinda, P. blanche, P. brandi, P. cecilia, P. claire, P. cora, P. dubitalis, P. erika, P. eunice, P. fannie, P. faye, P. flora, P. geneva, P. guadalupe, P. harriet, P. hattie, P. inez, P. iris, P. isabel, P. jan, P. janie, P. joanna, P. jodi, P. katrina, P. kristi, P. kristy, P. lindsey, P. lola, P. lula, P. lynda, P. madeline, P. maggie, P. mamie, P. margarita, P. maryann, P. melody, P. molly, P. monique, P. natasha, P. olivia, P. pecki, P. ramona, P. rosie, P. sabrina, P. sandy, P. shelley, P. sherri, P. sheryl, P. sonya, P. susie, and P. yvette. Ponaria, new genus of Coccidulini is proposed. Neaporia caerulea Gorham is transferred to Ponaria as a new combination. New species described in Ponaria are P. daviesi, P. hurtadoi, and P. paprzyckii. Lectotypes are here designated for Neaporia arrowi, N. coelestis, N. metallica, N. guatemalana, N. indagator, and Prodilis pallidifrons.
600
A new cryptic species, Aleochara (Xenochara) castaneimarmotae Klimaszewski, Webster, and Brunke, new species (Coleoptera: Staphylinidae: Aleocharinae), associated with Marmota monax (Linnaeus) burrows and caves in eastern North America, is described and illustrated. A key to Canadian species of subgenus Xenochara Mulsant and Rey and revised distributions of the taxonomically difficult fumata species group are provided. Aleochara quadrata Sharp is recorded from Washington and Oregon for the first time.
599
El pulgón Sarucallis kahawaluokalani (Kirkaldy) conocido comúnmente como pulgón del árbol de júpiter, se reporta por primera vez para Guatemala. Esta especie de la familia Aphididae (Hemiptera) fue encontrada en Lagerstroemia indica (Lythraceae). También se presenta la información básica de la especie así como una diagnosis para identificarla.
598
Se redescribe el género Ogyges Kaup, 1871 (Coleoptera: Passalidae), incluyendo su historia taxonó- mica. Se presentan las imágenes, los datos registrados en las etiquetas, las coordenadas geográficas, mediciones, comentarios taxonómicos y las instituciones depositarias de 19 tipos primarios que fueron descritos o que están incluidos dentro del género Ogyges Kaup. El nombre infrasubespecífico Ogyges laevior ab. vinculotaenia Kuwert, 1897, está indisponible, de acuerdo al artículo 45.6.2 del ICZN.
597
596
595
Many nomenclatural changes are implemented in the beetle families Georissidae, Histeridae, Hydraenidae, Hydrochidae, Hydrophilidae, Ptiliidae, Leiodidae and especially Staphylinidae, of the beetle series Staphyliniformia (Coleoptera), in preparation for making a world catalog of this group available online. Limited taxonomic changes are also made in the staphylinid subfamilies Osoriinae and Staphylininae.
At the level of family-group taxa, Article 29.4 of the current (1999) Zoological Code is reviewed and the original spellings of two tribal names, Nymphisterini Tishechkin (Histeridae) and Cryptonotopsisini Pace (Staphylinidae), are resurrected. The tribal name Stictocraniini Jakobson (Staphylinidae) is also resurrected as the valid name for its new synonym Fenderiini Scheerpeltz.
Changes at the genus-group level in Histeridae include placing Contipus Marseul as a new synonym of Hister Linnaeus due to the current placement of its validly designated type species C. subquadratus Marseul; proposal of Contipides Newton gen. nov. (type species Contipus digitatus Marseul) for the 10 species that had remained in Contipus of authors; and new designation of Idolia laevigata Lewis as type species of Idolia Lewis. In Ptiliidae, Rodwayia ovata Lea is newly designated as type species of Rodwayia Lea, and Throscidium germainii Matthews is newly designated as type species of Throscidium Matthews. In Staphylinidae, Paramichrotus Naomi is resurrected as a valid subgenus of Hesperosoma Scheerpeltz with Hemihesperosoma Hayashi placed as a new synonym of it; Sonoma corticina Casey is reaffi rmed as the type species of Sonoma Casey in place of Faronus tolulae LeConte; Stanosthetus Dejean is recognized as an available name and junior synonym of Euplectus Kirby; Taplandria Pace (type species T. guyanensis Pace) is recognized as a junior homonym and new synonym of Taplandria Pace (type species T. fl ava Pace); and Termitobiella Wasmann is resurrected as the valid name for the genus Felda Blackwelder. Replacement names for preoccupied generic or subgeneric names include in Histeridae Bellatricides Newton nom. nov. for Pachylister (Bellatrix) Mazur, junior homonym of Bellatrix Boie; and in Staphylinidae Foxiides Newton nom. nov. for Foxia Pace, junior homonym of Foxia Ashmead, and Xenasterides Newton nom. nov. for Xenaster Bierig, junior homonym of Xenaster Simonwitsch. Taxonomic changes at the generic level in Staphylinidae include proposal of Prolibia Newton gen. nov. (type species Lispinus californicus LeConte) for four Nearctic species recently placed in Clavilispinus Bernhauer; placement of Heterotrochinus Coiffait and its synonym Heterotrochus Coiffait as new synonyms of Eulibia Cameron; placement of the generic or subgeneric names Chapmaniella Bernhauer, Glenothorax Bierig, Euryolinus Bernhauer and Plesiolinus Bernhauer as new synonyms of Platydracus Thomson; and transfer of the subgenus Poikilodracus Scheerpeltz from Staphylinus Linnaeus to Platydracus. First reviser actions are used to select Georissites Ponomarenko (Georissidae) as the correct original spelling over the alternate original spelling Georyssites, and Kyrtusa Pace (Staphylinidae) as correct original spelling over Kirtusa.
Several hundred nomenclatural and taxonomic changes at the species group level are briefl y summarized here but are too numerous to list completely. Replacement names for preoccupied species or subspecies names in current use are proposed in Histeridae (3), Hydrochidae (1), Hydrophilidae (1), Leiodidae (2), Ptiliidae (3) and Staphylinidae (180); an additional staphylinid replacement name, Phloeopora nilgiriensis, is newly proposed by G. Paśnik. New or resurrected combinations are proposed for either nomenclatural or taxonomic reasons in the following genera (with indication of how many names in each genus): in Histeridae, Contipides Newton (10); in Staphylinidae, Abemus Mulsant and Rey (4), Allotrochus Fagel (6), Atheta Thomson (1), Cheilocolpus Solier (4), Eulibia Cameron (4), Foxiides Newton (1), Lispinus Erichson (3), Loncovilius Germain (2), Nacaeus Blackwelder (119), Naddia Fauvel (1), Neohypnus Coiffait and Sáiz (8), Neolosus Blackwelder (1), Ocypus Leach (2), Ontholestes Ganglbauer (1), Platydracus Thomson (59), Prolibia Newton (4) Termitobiella Wasmann (10), Thyreocephalus Guérin-Méneville (4), Xenasterides Newton (1), and Zeoleusis Steel (3). First reviser actions are used to resolve the correct original spellings (of two or more original spellings) of two species of Hydraena Kugelann (Hydraenidae) and 21 species of Staphylinidae. Changes in priority or availability of names are cited to establish the following names as valid over one or more new synonyms each: Acrotrichis rotundata (Haldeman) and Acrotrichis glabricollides Newton sp. nov. in Ptiliidae, Nemadiopsis franki Perreau in Leiodidae, and Gyrophaena nigra Kraatz, Heterothops fumigatus LeConte, Loncovilius germaini (Scheerpeltz), Philonthus upotovus Newton, sp. nov., Stenus fulviventris Rougemont, and nine species of Homalota Mannerheim in Staphylinidae. Finally, the species Eleusis lata Coiffait and Eleusis microlestiformis Coiffait are noted as not belonging to the genus Eleusis Laporte de Castelnau or to Staphylinidae, and are transferred without generic assignment to the subfamily Inopeplinae of the family Salpingidae.
594
The tribe Zolini (Carabidae: Trechinae) is revised for New Zealand. Two subtribes, fi ve genera, and fortyeight species are recognized. The presence of the subtribe Merizodina is confi rmed for New Zealand; it includes three genera (Maungazolus n. gen., Pseudoopterus Csiki, 1928, and Synteratus Broun, 1909). The subtribe Zolina contains two genera (Oopterus Guérin-Méneville, 1841 and Zolus Sharp, 1886).
One genus and twenty-fi ve species are described as new: Maungazolus n. gen.; Maungazolus acutus n. sp., Maungazolus priestleyensisn. sp., Maungazolus ranatungaen. sp., Maungazolus septempunctatus n. sp., Maungazolus tararuaensisn. sp., Oopterus anglemensisn. sp., Oopterus arthurensisn. sp., Oopterus corvinki n. sp., Oopterus discoideus n. sp., Oopterus garnerae n. sp., Oopterus marrisin. sp., Oopterus mohiensis n. sp., Oopterus monticola n. sp., Oopterus nanus n. sp., Oopterus nunni n. sp., Oopterus palmai n. sp., Oopterus punctatus n. sp., Oopterus quadripunctatus n. sp., Oopterus taieriensis n. sp., Oopterus trechoides n. sp., Pseudoopterus fi ordlandensis n. sp., Pseudoopterus otiraensis n. sp., Zolus kauriensis n. sp., Zolus unisetosus n. sp., Zolus wongi n. sp.
Lectotypes are designated for Zolus helmsi Sharp, 1886 and Oopterus parvulus Broun, 1903.
Seven taxa are reinstated as valid genera and previous combinations: Zolus Sharp, 1886 and Pseudoopterus Csiki, 1928 (previously synonymized with Oopterus Guérin-Méneville,1841); Oopterus carinatus Broun, 1882 = Zolus carinatus (Broun, 1882); Oopterus helmsi (Sharp, 1886) = Zolus helmsi Sharp, 1886; Oopterus latipennis Broun, 1903 = Pseudoopterus latipennis (Broun, 1903); Oopterus plicaticollis Blanchard, 1843 = Pseudoopterus plicaticollis (Blanchard, 1843); Oopterus subopacus (Broun, 1915) = Zolus subopacus Broun, 1915.
Three new combinations are established: Oopterus pallidipes Broun, 1893 = Maungazolus pallidipes (Broun, 1893); Oopterus parvulus Broun, 1903 = Maungazolus parvulus (Broun, 1903); Oopterus pygmeatus Broun, 1907 = Maungazolus pygmeatus (Broun, 1907).
Eleven new synonymies are established: Oopterus basalis Broun, 1915 and Oopterus minor Broun, 1917 = Maungazolus pygmeatus (Broun, 1907); Oopterus nigritulus Broun, 1908 and Oopterus laevigatus Broun, 1912 = Oopterus sobrinus Broun, 1886; Oopterus sculpturatus ovinotatus Broun, 1908 = Oopterus sculpturatus Broun, 1908; Oopterus probus Broun, 1903 and Oopterus latifossus Broun, 1917 = Pseudoopterus latipennis (Broun, 1903); Oopterus puncticeps Broun, 1893 = Oopterus laevicollis Bates, 1871; Zolus atratus Broun, 1893 and Zolus femoralis Broun, 1894 = Zolus carinatus (Broun, 1882); Zolus labralis Broun, 1921 = Zolus helmsi Sharp, 1886.
A revision of all taxa is provided. Descriptions, identifi cation keys, illustrations of male genitalia, habitus photos, distributional data and maps are given. Extensive information on ecology, biology, dispersal power, and collecting techniques is included for each species.
593
Species of Hydrosmecta C. G. Thomson, occurring in eastern Canada, are studied and reviewed. Six species are recognized, and two of them are described as new to science. The new species are: Hydrosmecta canadensis Webster and Klimaszewski, new species, and Hydrosmecta minutissimoides Webster and Klimaszewski, new species. Two species described in the USA are recorded from Canada and New Brunswick for the first time: Hydrosmecta caduca Casey and Hydrosmecta dulcis Casey. New distribution and collection data, diagnoses, a key for species identification, and images of habitus and genital structures are provided. Hydrosmecta newfoundlandica Klimaszewski and Langor, 2011, is transferred to the genus Atheta Thomson based on morphology of genital structures.
592
This paper revises the genus Ganelius Benesh, which is endemic to Madagascar, in the stag beetle tribe Figulini Burmeister (Coleoptera: Lucanidae: Lucaninae). Species in the genus are striking in their highly sexually dimorphic mandibles, a character that rarely occurs in the tribe. The genus was previously comprised of four species, but three of these names were problematic or questionably applied. A lectotype is designated for G. madagascariensis Laporte, a neotype is designated for Ganelius oberndorferi Nonfried, and Nigidius passaliformis Benesh is newly synonymized with G. oberndorferi. Two new species, G. gnamptus Paulsen and G. zombi Paulsen, are described from western Madagascar. The identity of the overlooked Ganelius nageli (Kriesche) is fixed through a neotype designation, and the species is moved to the new genus Agnelius, which is distinguished from Ganelius by a lack of sexual dimorphism, serrate protibiae, and a more flattened body.
591
New genera and species of mimetic Cleridae from Mexico and Central America (Coleoptera: Cleroidea)
(2017)
New genera and species of putatively mimetic Cleridae (Coleoptera: Cleroidea) are described from Mexico and Central America: Namba testacea, new genus and species from Nuevo León, Mexico; Aulicoides leavengoodi, new genus and species from Jalisco, Mexico; and Quadrophenia, new genus, which includes Q. townshendi, new species, from Mexico, Nicaragua, Costa Rica and Panama, Q. mooni, new species, from Costa Rica, Q. entwistlei, new species, from Guatemala and Honduras, Q. daltreyi, new species, from Costa Rica, and Q. sumidero new species, from Chiapas, Mexico. A key to the species of Quadrophenia is provided. All of the newly described species are hypothesized to be Batesian mimics of aposematically colored coleopteran models, some of which are known to be chemically protected.
590
A systematic redefinition of the species belonging to the genus Geomyphilus Gordon and Skelley, 2007 (Coleoptera: Scarabaeidae: Aphodiinae) of Mexico and neighboring countries is presented. The new species G. tuzincola of Mexico is described and figured. The new combination Coelotrachelus macgregori (Islas, 1955) is proposed.
589
Distributional records of the 177 tiger beetle species and subspecies known for Vietnam are given together with a key and habitus photos (Coleoptera: Carabidae: Cicindelinae). Tricondyla (Tricondyla) macrodera abruptesculpta Horn, 1925, Protocollyris festiva Naviaux, 2008, Neocollyris (Brachycollyris) purpureomaculata borea Naviaux, 1994, Neocollyris (Brachycollyris) torosa Naviaux, 2010, Neocollyris (Leptocollyris) rogeri Shook and Wu, 2006, Neocollyris (Leptocollyris) laosensis Naviaux, 1999, Neocollyris (Stenocollyris) rubens (Bates, 1878), Neocollyris (Pachycollyris) tricolor Naviaux, 1991, Neocollyris (Pachycollyris) panfi lovi Naviaux and Matalin, 2002, Therates rugulosus Horn, 1900, Therates bannapecolus Sawada and Wiesner, 1999, Calochroa interruptofasciata fl avolineata (Chaudoir, 1865), Cylindera (Ifasina) cyclobregma Acciavatti and Pearson, 1989, Cylindera (Ifasina) decolorata (Horn, 1907), Cylindera (Ifasina) humillima (Gestro, 1893) and Cylindera (Eugrapha) elisae elisae (Motschulsky, 1859) are reported for the fi rst time in the fauna of Vietnam. Neocollyris (Isocollyris) septentrionalis Naviaux, 1999, Neocollyris (Leptocollyris) linearis tenuicornis (Chaudoir, 1864), Neocollyris (Pachycollyris) aptera apicalis (Chaudoir, 1864), Neocollyris (Pachycollyris) contracta (Horn, 1905), Therates dimidiatus wallacei Thomson, 1857, Cylindera (Cylindera) pseudocylindriformis Horn, 1913, Cylindera (Ifasina) viridilabris (Chaudoir, 1852), Cylindera (Ifasina) psilica psilica (Bates, 1866), Cylindera (Eugrapha) venosa (Kollar, 1836) and Myriochila (Myriochila) undulata (Dejean, 1825) are excluded from the fauna of Vietnam
588
587
Adesmus martinsi (Coleoptera, Cerambycidae, Lamiinae, Hemilophini), a new species from Bolivia, is described, illustrated, and included in a previous key. The new species displays gender dimorphism in the pubescent pattern and in anatomical structure. Thus, Adesmus becomes the second genus recorded in the Hemilophini to have visual chromatic dimorphism.
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A preliminary catalogue of the moths (Lepidoptera except Papilionoidea) of Tobago, West Indies
(2017)
This catalogue comprises records of 355 species of moths (non-papilionoid Lepidoptera) from Tobago, of which 15 are partially identified. Of this total, all except 17 (5%) are known from Trinidad, although not all these records from Trinidad are published yet. Of these 17, eleven are expected to occur in Trinidad as they also occur on the mainland of South America and two are only known from Tobago but will probably also occur in Trinidad. This leaves just four species (1% of the total) that are known from the Lesser Antilles and are currently not known from further south than Tobago. The families represented by the most species are Erebidae, Crambidae, Geometridae, Noctuidae and Sphingidae, which between them account for 73% of records. Taxonomic changes are made as follows. Podalia farmbri (Kaye, 1925) sp. rev. (Megalopygidae) is removed from the synonymy of P. nigrescens Schaus, 1905. Podalia walkeri Hopp, 1935 and P. dimidiata (Walker, 1865) are syn. nov. of P. farmbri Kaye, 1925. Renia bipunctata (Kaye, 1901) (Erebidae) is a comb. nov. for Zanclognatha bipunctata. Aristaria trinitalis Schaus, 1906 (Erebidae) is a syn. nov. of Renia bipunctata Kaye, 1901. Aglaonice deldonalis Walker, 1859 sp. rev. (Erebidae) is removed from the synonymy of A. hirtipalpis Walker, [1859]. Plusiodonta cupristria Kaye, 1923 (Erebidae) is a syn. nov. of Oraesia excitans Walker [1858]. Oroscopa abluta (Schaus, 1912) (Erebidae) is a comb. nov. for Freilla abluta Schaus, 1912, which is a new combination in common use, but not previously published. Ptichodis dorsalis (Fabricius, 1797) (Erebidae) is a comb. nov. for Noctua auct. dorsalis Fabricius, a new combination already in use, but not formally published. I endorse the unpublished conclusion of I.W.B. Nye that Ptichodis basilans (Guenée, 1852) is a syn. nov. of Ptichodis dorsalis (Fabricius, 1797). Ptichodis agrapta Hampson, 1913 is also a syn. nov. of Ptichodis dorsalis (Fabricius, 1797).
584
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Se describe con medidas, dibujos e imágenes obtenidas con microscopio compuesto, una nueva especie de mosca blanca, Tetraleurodes dorsibandas Sánchez y Carapia sp. nov. (Hemiptera: Aleyrodidae) que se encontró alimentándose de Casimiroa edulis (Rutaceae) y Ehretia tinifolia (Boraginaceae) en Morelos y Oaxaca (México), respectivamente. La especie se diferencia de Tetraleurodes perileuca (Cockerell) por tener el raquis en forma de bandas, presentar estructuras en forma de mosaico irregular en la parte media de los segmentos abdominales y la presencia de setas cefálicas. Se proporciona una clave de las especies de México y se enlistan las especies del nuevo mundo.
582
New state records for 49 species of Eucnemidae (Coleoptera) are reported throughout the United States and a single species is newly reported from Utah. Diagnostic remarks are offered for Xylophilus crassicornis Muona and distributional observations are discussed for Dirrhagofarsus lewisi. The importance of retaining by-catch from statewide, regional, and national surveys for future studies is also discussed.
581
Thirty-one species of mealybugs (Hemiptera: Pseudococcidae) are reported from South Korea based upon monitoring surveys conducted from 2010 to 2016, along with a dichotomous key to separate them. Six species, Nipaecoccus nipae (Maskell), Palmicultor lumpurensis (Takahashi), Planococcus citri (Risso), Pseudococcus dendrobiorum Williams, Pseudococcus longispinus (Targioni-Tozzetti), and Pseudococcus orchidicola Takahashi, known only to occur in greenhouses, are considered as adventive mealybug species to South Korea; of these, P. citri has already established in the exterior environment in South Korea.
580
579
A statewide survey of Wisconsin’s Nemonychidae and Anthribidae (Coleoptera: Curculionoidea) was conducted over one full (2012) and two partial (2011, 2013) fi eld seasons. Specimens were collected using a variety of techniques. Two species of the nemonychid genus Cimberis and 22 species of Anthribidae in 13 genera are now confi rmed. Generic and species-level keys for Wisconsin species are provided along with generic and species diagnoses. Species treatments also include a synonymy, a description, and information on natural history, phenology, distribution and collecting methods. Dorsal and lateral habitus images are also provided for most of the species.
578
Tragosoma soror (Coleoptera: Cerambycidae: Prioninae), a new Nearctic species, is described and compared to its North American congeners. Tragosoma harrisii LeConte, revised status, and Tragosoma spiculum Casey, revised status are both resurrected from synonymy and considered valid species distinct from T. depsarium (Linnaeus). The Mexican T. nigripenne Bates is confi rmed as a valid species. Tragosoma chiricahuae Linsley, new synonym, is placed in synonymy under T. spiculum Casey, and T. sodale Casey, new synonym, T. parvicolle Casey, new synonym, and T. repens Casey new synonym are placed in synonymy under T. harrisii. Tragosoma repens is a junior objective synonym of T. parvicolle. A lectotype is designated for T. sodale. An updated synonymical and distributional checklist of Tragosoma of the world is provided.
577
Two new genera and species of tiger beetles from Baltic amber (Coleoptera: Carabidae: Cicindelinae)
(2017)
Two fossil tiger beetle species (Coleoptera, Carabidae, Cicindelinae) are described from Eocene Baltic amber using light microscopic and X-ray microscopic techniques. Both species are considered representatives of the subtribe Iresina Rivalier, 1971 due to the shared combination of character states: glabrous head, six labral and four suborbital setae, and glabrous pronotum. Palaeopronyssiformia groehni Wiesner, Will, and Schmidt, new genus, new species, is characterized by a glabrous and furrowed head with six labral setae, large eyes, presence of two supraorbital setae on each side, mandibles with two teeth of the incisor region, and a glabrous and furrowed pronotum. Palaeoiresina cassolai Wiesner, Will, and Schmidt, new genus, new species, is characterized by a unicolored, undentated labrum, mandibles with two teeth of the incisor region, glabrous head with six labral setae, two clypeal setae, two supraorbital setae on each side, and a glabrous pronotum, mesepisternum, mesepimeron, and metepisternum. The species described here represent the only known tiger beetle fossils preserved in Baltic amber.
576
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Redescription of adult and observations of the weevil Revena plaumanni Bondar (Coleoptera: Curculionidae), preying on seeds of the palm tree Butia eriospatha (Mart. ex Drude) Becc. (Arecaceae), as well as oviposition and larval behavior, pupation site, and adult emergence are reported for the fi rst time in Brazil. Photos of host plant, immature forms (egg, larva and pupa) and adult are given. Illustrations of male and female genitalia are also provided.
574
The New World genus Chariessa Forster (Coleoptera: Cleroidea: Cleridae) is revised and includes C. catalina Opitz, new species, C. elegans Horn, C. dichroa (LeConte), C. floridana Schaeffer, C. pilosa (Forster), C. texana Wolcott, C. ramicornis Perty, C. vestita (Chevrolat), and C. duponti (Spinola). Enoplium pilosa var. marginata Say is synonymized with Chariessa pilosa Forster. Lectotypes are designated for C. pilosa (Forster), C. ramicornis Perty, and C. vestita (Chevrolat). Available information indicates that Chariessa adult and immature individuals are predatory on lignicolous insects with a particular affinity for cerambycids and buprestids that infest species of oak. It is postulated that Pleistocene speciation generated the North American components of Chariessa with more ancient southern species generated during the Middle Tertiary; after closures of the Middle American portals and orogeny of the South American Andes. Included in this treatise is a discussion of natural history, key to species, narratives of zoogeography and phylogeny, one diagram of a phylogenetic tree, 35 line drawings, eight SEM micrographs, twelve habitus photographs, nine photographs of male genitalia, and five distributional maps.
573
With an incident in Palo Duro Canyon, Texas, USA, Scolopendra heros Girard (Chilopoda: Scolopendromorpha: Scolopendridae) becomes the third centipede species known to prey on bats; S. gigantea Linnaeus and S. viridicornis Newport have been so documented in Venezuela and Brazil, respectively. The Texas predation was interrupted by the predator/prey pair’s falling around 15–20 m from the canyon wall and, perhaps also, by human presence where they landed. The centipede uncoiled and retreated to shelter under a nearby rock and, after initial immobilization, so did the bat.
572
Abstract. Six new species of Coleoxestia Aurivillius, 1912 (Cerambycidae, Cerambycinae, Cerambycini) are described: C. clarkei from Bolivia; C. chemsaki and C. eyai from Peru; C. fragosoi and C. rafaeli from Brazil; and C. hovorei from Ecuador. Coleoxestia rachelae Eya and Chemsak is also newly recorded from Peru (new country record). Dorsal, ventral and lateral illustrations along with illustrations of various other structures are provided for each of the new species. In some cases, existing key couplets by Eya and Chemsak (2005) or Martins and Monné (2005) are modifi ed to help separate a new species from similar existing species. A reproduction of the general descriptive terminology for the head, prothorax and antennomeres of Coleoxestia, from Fragoso (1993), is also included in the illustrations.
571
A trimaculate male of the diplopod genus Apheloria Chamberlin (Polydesmida: Xystodesmidae/-inae: Apheloriini) from 1.3 km (0.8 mi) west of McKenney, Dinwiddie County (Co.), Virginia, is designated the Neotype of Julus virginiensis Drury 1770, thereby stabilizing the earliest name for a North American milliped and authenticating its prior assignment to this taxon. The existing concept of Apheloria is accepted in the absence of a revisionary treatment, and a modern description of A. v. virginiensis with gonopod drawings and color photos is provided. Drury’s original account and his letter to the Virginian who sent him the original specimens are quoted verbatim to eliminate future library searches. The specific name has been associated with at least three genera, and its confusing history is clarified by summarizing works in each. Authentic localities, mapped to the extent now possible, reveal a distribution south of the James River in piedmont and coastal Virginia that extends southwestward to the Blue Ridge foothills and at least as far south in North Carolina (NC) as Greensboro, the “Triangle” (Raleigh/Durham/Chapel Hill region), and Albemarle Sound in the east. Based on the holotypes, A. aspila and A. tigana, both by Chamberlin, are placed in synonymy under A. v. virginiensis (syns. nov.), and although its status is still under review, A. waccamana Chamberlin, whose type locality is Lake Waccamaw, Columbus Co., in southeastern NC, may be the correct name for today’s A. tigana. All samples so labeled must be reexamined for misidentifications of A. v. virginiensis.
570
A detailed study of the holotype of Sphecomyrma canadensis Wilson, 1985 (Hymenoptera: Formicidae) from Canadian amber has led to the conclusion that the specimen belongs to a new genus, here named Boltonimecia gen.n. Since the taxonomy of stem-group ants is not well understood, in order to find the taxonomic position of this genus, it is necessary to review the classifi cation of stem-group ants in a study of their relation to crown-group ants. In the absence of data for traditional taxonomic approaches, a statistical study was done based on a morphometric analysis of antennae. Scape elongation is believed to play an important role in the evolution of eusociality in ants; however, this hypothesis has never been confirmed statistically. The statistical analysis presented herein lends support to the view that antennal morphology reliably distinguishes stem-group ants from crown-group ants, to determine whether a species belongs to one or the other group. This, in turn, may indicate a relationship exists between eusociality and scape elongation. A review of Cretaceous records of ants is made and the higher classification of Formicidae with definitions of stem and crown groups is proposed. Newly obtained data are discussed focusing particularly on the origin, evolution and diversity of ants.
569
A primitive subfamily of false click beetles (Coleoptera: Eucnemidae: Phlegoninae) distributed primarily in the Neotropical region is revised. Euryphlegon gen. nov. is described from Belize in Central America. New species include: Phlegon chiriquiensis sp. nov. (Panama), Phlegon panamensis sp. nov. (Panama), Euryphlegon jacqueschassaini sp. nov. (Panama) and Euryphlegon parallelus sp. nov. (Belize). Phlegon herculeanus (Lacordaire) stat. res. is resurrected from synonymy with Phlegon buqueti Laporte. One new combination is proposed: Euryphlegon degallieri (Muona) (Phlegon). Based on a number of observed external character traits, Euryphlegon is placed in Orodotini Muona, 1993 within Macraulacinae Fleutiaux, 1922. Identifi cation keys are provided for species of Phlegon and Euryphlegon in the Neotropical region. The relationships among Phlegon, Euryphlegon, Euryptychus LeConte and other groups within Echthrogasterini Cobos, 1964 and Orodotini are discussed.
568
New Bolivian Rhinotragini (Coleoptera, Cerambycidae, Cerambycinae) are described: three species of Phygopoda Thomson, 1864 (P. longiscopifera sp. nov., P. boliviensis sp. nov. and P. chaquensis sp. nov.); and one species of Phygopoides Peñaherrera-Leiva and Tavakilian, 2007 (P. maxwelli sp. nov.). Two Brazilian species of Neophygopoda Melzer, 1933 are transferred to the genus Phygopoda: P. exilis (Melzer, 1933) comb. nov. and P. agdae (Martins, Galileo and Santos-Silva, 2015) comb. nov. All the species are illustrated, and a key to the Bolivian species of Phygopoda and host flower records are provided.
567
The taxonomic history of the rhinotragine genera Phygopoda Thomson, 1864 and Pseudophygopoda Tavakilian and Peñaherrera-Leiva, 2007 (Coleoptera: Cerambycidae: Cerambycinae) are discussed, and evidence is presented to suggest that some recent taxonomic changes made by Carelli and Monné (2015) were unjustified. Consequently, Phygopoda nigritarsis Gounelle, 1911 is moved to the genus Neophygopoda Melzer, 1933, creating the new combination Neophygopoda nigritarsis, the genera Panamapoda Clarke, 2014 and Paraphygopoda Clarke, 2014 are revalidated, and the species Paraphygopoda viridimicans (Fisher, 1952) and Paraphygopoda nappae Clarke, 2014 are also revalidated.
566
Dipropus tequesta Johnson new species (Coleoptera: Elateridae) is described from southern Florida. Dipropus fuscus (LeConte) is a new synonym of D. soleatus (Say), and D. granosus (Fall) is a new synonym of D. asper (LeConte). The fl ightless female of D. asper is described and provides the fi rst report of brachyptery and endogean habits in Dipropus. A key to the species of the eastern United States and a new checklist of species for the country are provided.
565
Una nueva especie del género Eugnosta Hübner, E. lankinae n.sp. (Lepidoptera: Tortricidae), es descrita para la Provincia de Elqui, Chile. Se entregan los caracteres morfológicos del adulto y las fi guras de la genitalia masculina. Además se discute su posición sistemática con dos especies congéneres Neotropicales.
564
563
562
The primary types of Onciderini Thomson, 1860 (Coleoptera: Cerambycidae: Lamiinae) of the Museum of Comparative Zoology (MCZ), Harvard University, are catalogued and illustrated. Data on the original combination, current name, gender, and type locality are verifi ed and presented. There are 19 primary types of Onciderini including seven in Oncideres Lacordaire, 1830; two in Hesychotypa Thomson, 1868; and two in Hypsioma Audinet-Serville, 1835. Of the 19 primary types, 18 were described by L. S. Dillon and E. S. Dillon. A brief history of the Coleoptera collection at the MCZ is also presented.
561
A number of nomenclatural issues regarding Neotropical Scarabaeoidea are addressed. Athyreus soesilae Makhan, 2008 is synonymized with Neoathyreus (Neoathyreus) excavatus (Laporte, 1840) (Geotrupidae: Bolboceratinae: Athyreini). Scalmus Zang, 1905 is replaced with its senior synonym Neleuops Kuwert, 1891, nec Neleurops Kuwert, 1891, and the four species it presently contains are thus transferred to Neleuops (Passalidae: Passalinae: Passalini). The spelling of the name Oxysternon oberthueri Olsoufi eff, 1924, a junior synonym of Oxysternon (Oxysternon) conspicillatum (Weber, 1801), is fi xed (Scarabaeidae: Scarabaeinae: Phanaeini). The year of description of Oxysternon (Oxysternon) festivum (Linnaeus, 1758) is corrected, its existing lectotype invalidated, and a new lectotype designated (Scarabaeidae: Scarabaeinae: Phanaeini). The correct spelling of Phanaeus (Phanaeus) prasinus trinidadens Arnaud, 2001 and Phanaeus (Phanaeus) prasinus joliyi Arnaud, 2001 is discussed (Scarabaeidae: Scarabaeinae: Phanaeini). The seniority of Chlorota aulica Burmeister, 1844 over Chlorota metallica Burmeister, 1844 is restored (Scarabaeidae: Rutelinae: Rutelini). The year of description of Rutela lineola (Linnaeus, 1758) is corrected and a lectotype is designated (Scarabaeidae: Rutelinae: Rutelini). Cyclocephala aulustjaorum nomen novum is proposed as a replacement name for the preoccupied name Cyclocephala brevis Höhne, 1923 (Scarabaeidae: Dynastinae: Cyclocephalini). The spelling of the generic name Harposceles Burmeister, 1847 is fi xed (Scarabaeidae: Dynastinae: Cyclocephalini). The incorrect subsequent spelling Stenocrates haackae Ratcliffe, 1977 is preserved (Scarabaeidae: Dynastinae: Cyclocephalini). The name Scarabaeus simson Linnaeus, 1767 is invalidated as a synonym of Megasoma actaeon (Linnaeus, 1758) (Scarabaeidae: Dynastinae: Dynastini). The spelling of the generic name Coelosis Hope, 1837 is found to be correct (Scarabaeidae: Dynastinae: Oryctini). The validity of Steyskal’s emendation of the name Megaceropsis quadridentata Dechambre, 1976 is confi rmed (Scarabaeidae: Dynastinae: Oryctini). The lectotype of Podischnus agenor (Olivier, 1789) is invalidated and a new lectotype is designated (Scarabaeidae: Dynastinae: Oryctini).
560
Chiquitano gen. nov. Chiquitano volcanesensis sp. nov., Compsibidion achiraensis sp. nov. and Compsibidion amboroensis sp. nov. (Coleoptera: Cerambycidae: Cerambycinae: Neoibidionini) are described from Bolivia. Notes on Rhysium Pascoe, 1866 and Rhysium bimaculatum Pascoe, 1866 are provided, and Brechmoidion separatum Martins and Galileo, 2007 is transferred to Rhysium. Keys to species of Compsibidion Thomson, 1864, Brechmoidion Martins, 1969 and Rhysium Pascoe, 1866 are also provided.
559
The Antillean fauna of Mantispidae (Neuroptera) consists of six species in four genera, all in the subfamily Mantispinae. Two new species are here described: Leptomantispa antillesensis Hoffman, n. sp., is described from Puerto Rico, Anguilla, British Virgin Islands (Grand Camanoe, Guana, Tortola), United States Virgin Islands (St. Croix, St. John, St. Thomas), Dominica, and Guadeloupe, and Leptomantispa hispaniolaensis Hoffman, n. sp. is described from Hispaniola (Dominican Republic). Mantispilla zayasi Alayo is synonymized under Leptomantispa pulchella (Banks), new synonomy, and Mantispilla taina Alayo is synonymized under Zeugomantispa minuta (Fabricius), new synonomy. Climaciella cubana Enderlein is recorded for the fi rst time from Hispaniola (Dominican Republic). Dicromantispa sayi (Banks) is newly reported from the Cayman Islands (Cayman Brac and Little Cayman), and Hispaniola (Dominican Republic). Zeugomantispa minuta (Fabricius) is now recorded from the Bahamas (Andros) and Hispaniola (Dominican Republic).
558
Six new species of false click beetle (Coleoptera: Eucnemidae) are described. These new species are: Fornax dixiensis sp. nov. (Florida, USA), Dromaeolus comayaguiensis sp. nov. (Honduras), Asiocnemis bicolor sp. nov. (Ecuador), Miruantennus chinensis sp. nov. (Yunnan prov., China), Miruantennus cuneiformis sp. nov. (Malaysia) and Nematodes africanus sp. nov. (Côte d’Ivoire). Identifi cation keys are provided for species of Fornax in the Nearctic ecozone and Miruantennus in the Palearctic and Indo-Malayan ecozones. A new key to include all the genera within the tribe Nematodini Leiler is also provided.
557
Conognatha iris iris Olivier (Coleoptera: Buprestidae) is recorded for the fi rst time for Venezuela. The record is based on a single female specimen from Caicet hill, Amazonas state. A map of the known distribution of the taxon is provided
556
Se presenta una sinopsis de los Phycitinae (Lepidoptera: Pyralidae) de Chile, incluyendo listado sistemático con su distribución geográfi ca. Se registra la presencia de Inverina suizensis Neunzig y Goodson, 1992, por primera vez en Chile y se describe una nueva especie del género Homoeographa Ragonot, 1888.
555
The monotypic clerid genus Ohanlonella Rifkind (Coleoptera: Cleridae: Clerinae), described from Oaxaca, is reported for the first time from the state of Veracruz. The recently discovered population shows distinctly different coloration from the type series of O. esperanzae Rifkind, but appears to be otherwise identical. The Veracruz morph is briefly characterized and illustrated, and its significance is discussed.
554
A new species of Ctilodes Murray, 1864 (Coleoptera: Nitidulidae) was recovered in material from Vietnam during a large study of Carpophilinae. A description and detailed diagnosis of Ctilodes clinei Powell and Duffy new species is presented here along with a key to identify all currently known species of Ctilodes.
553
Coprophagy and probable saprophagy are reported for larvae of two species of chrysaugine moths (Lepidoptera: Pyralidae). Larvae of Parachma ochracealis Walker are found in rotten pine wood and mulch in North-Central Florida. Larvae of Basacallis tarachodes (Dyar) inhabit feces of an unidentified small mammal in a cave in Central Florida and seem to be troglophilic. These behaviors are compared to similar ones in Neotropical chrysaugines such as Humiphila Becker and Cryptoses Dyar. Saprophagy and coprophagy are predicted to be more general habits among Chrysauginae.
552
Distributional records of the 30 tiger beetle species and subspecies (Coleoptera: Carabidae: Cicindelinae) known for Brunei Darussalam are given together with habitus photos for 21 species. Neocollyris (Neocollyris) labiomaculata (Horn, 1892), Neocollyris (Neocollyris) emarginata (Dejean, 1825), Therates spectabilis fl avissimus Brouerius van Nidek, 1957, Heptodonta analis s. str. (Fabricius, 1801), Cosmodela velata (Bates, 1872), Lophyra (s. str.) fuliginosa (Dejean, 1826), Cylindera (Leptinomera) fi ligera (Bates, 1878), Myriochila (s. str.) specularis brevipennis (Horn, 1897), Abroscelis tenuipes araneipes (Schaum, 1863) and Callytron doriai (Horn, 1897) are reported for the fi rst time for the Sultanate.
551
Scale insects (Hemiptera: Coccoidea) were collected on oak trees (Quercus spp.; Fagaceae) in Korea and identifi ed as Asterodiaspis luteola (Russell) (Asterolecaniidae) and Nidularia japonica (Kuwana) (Kermesidae), which are newly documented in the Korean fauna of scale insects. Characteristics of these species are briefl y given and illustrative photographs and information on their distribution and hosts are provided.
550
Cixidia fusca and Synecdoche impunctata (Hemiptera: Achilidae) are reported from Missouri for the first time, new state records. Ecological and trapping information is also provided.
549
548
We describe a new butterfl y species, Anaeomorpha mirifi ca Simon and Willmott, n. sp. (Lepidoptera, Nymphalidae, Charaxinae), from premontane rain forest of the Chocó region of northwestern Ecuador. This represents the second known species and the fi rst record outside of the Amazon basin for this taxonomically and biologically enigmatic genus. Although the two species are not known to be sympatric, we identifi ed 21 characters in the external color pattern, wing shape and male genitalia that together support distinct species status. Most notably, the new species possesses an ocellus in the ventral hind wing tornus, a character which occurs in the Neotropical Charaxinae only in the genus Prepona Boisduval, 1836. A mean divergence of 6.8% in the COI ‘barcodes’ between the two species underlines their taxonomic distinctness.
547
The eight species in the genus Tomarus Erichson (Coleoptera: Scarabaeidae) in Argentina, Chile, and Uruguay are reviewed. Tomarus roigjunenti new species and Tomarus spinipenis new species are described from Argentina. We include a key to species, representative habitus illustrations for all species, character illustrations, and distribution maps for each, as well as commentary about the natural history and distributions for each species. Diagnostic characters are discussed for each species, and species relationships are hypothesized based on the analysis of internal and external morphological characters. The male of T. bidentulus (Fairmaire) is described for fi rst time. The following taxonomic changes are made: Tomarus guianucai Dechambre and Lumaret, 1985 is a new junior synonym of Tomarus rubripes (Boheman, 1858), which was formerly and incorrectly cited as occurring in Argentina.
546
545
Descriptions and illustrations of the triungulin, 5th instar, and prepupal larval forms for Nematodes penetrans (LeConte, 1852) (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) are given. The descriptions are based on triungulins collected in a plastic vial as well as 5th instars and prepupal larval stages collected from hard sections of sugar maple (Acer saccharum Marshall; Aceraceae) logs and limbs. Adults were reared from those pieces placed in plastic bags. Biological information is provided, based on literature search and personal observations.
Comparative studies between larvae of N. penetrans and the European species, Nematodes fi lum (Fabricius, 1801) along with other Nearctic larval forms are briefl y discussed. This discovery represents the fi rst larval description for the genus in the Nearctic region.
544
543
Four new species of Chrysina Kirby (Coleoptera: Scarabaeidae: Rutelinae) from Guatemala and Honduras
(2017)
Four new species of the genus Chrysina Kirby (Coleoptera: Scarabaeidae: Rutelinae) are described from cloud forests of Guatemala and Honduras. The Guatemalan species are C. alexae n. sp. from Purulhá, Baja Verapáz in the central mountains and C. woodruffi n. sp. from Bulej, Huehuetenango in Los Cuchumatanes Mountains at elevations from 1500 to 2000 meters. The species from Honduras are C. antonkozlovi n. sp. from Celaque, Lempira and C. maishei n. sp. from El Güisayote, Ocotepeque, at elevations between 2000 and 2600 meters above sea level.
542
Species descriptions, keys to genera and species, and geographical distributions are presented for 43 species of the family Bruchidae (Coleoptera: Chrysomeloidea) for Chile. Of these species, seven are described as new:
Acanthoscelides aricae sp. nov., Lithraeus chillan sp. nov., L. comptus sp. nov., L. elguetai sp. nov., L. limari sp. nov., L. lonquimay sp. nov., and L. penai sp. nov. Eight species are endemic to Chile. A list of true host plants and floral records for those with known host associations is presented. Habitus photographs and drawings of pertinent body parts, including male genitalia, are provided. References pertaining to the previously described species are listed.
541
The genus Paraphloeolaemus Thomas (Coleoptera: Cucujoidea: Laemophloeidae) is described for two new Neotropical species, P. vorticosus Thomas, new species, and P. pterosiagon Thomas, new species. Diagnoses and illustrations are provided.
The following 16 species are transferred from Laemophloeus Dejean (s. l.) to Phloeolaemus Casey: Phloeolaemus anticus (Sharp, 1899: 518) [= Laemophloeus anticus Sharp, 1899], new combination; Phloeolaemus boops (Sharp, 1899: 517) [= Laemophloeus boops Sharp, 1899], new combination; Phloeolaemus castaneipennis (Grouvelle, 1876: 494) [= Laemophloeus castaneipennis Grouvelle, 1876: 494], new combination; Phloeolaemus championi (Sharp, 1899: 516) [= Laemophloeus championi Sharp, 1899], new combination; Phloeolaemus curtus (Grouvelle, 1876: xxxiii) [= Laemophloeus curtus Grouvelle, 1876], new combination; Phloeolaemus endomychus (Sharp, 1899: 519) [= Laemophloeus endomychus Sharp, 1899], new combination; Phloeolaemus hoplites (Sharp, 1899: 517) [= Laemophloeus hoplites Sharp, 1899], new combination; Phloeolaemus ignobilis (Sharp, 1899: 518) [= Laemophloeus ignobilis Sharp, 1899], new combination; Phloeolaemus impressus (Grouvelle, 1876: xxxiii) [= Laemophloeus impressus Grouvelle, 1876], new combination; Phloeolaemus lacerdae (Grouvelle, 1877: 211) [= Laemophloeus lacerdae Grouvelle, 1877], new combination; Phloeolaemus macrocephalus (Schaeffer, 1910: 214) [= Laemophloeus macrocephalus Schaeffer, 1910], new combination; Phloeolaemus punctulaticollis (Hetschko, 1929: 94) [= Laemophloeus punctulaticollis Hetschko, 1929], new combination; Phloeolaemus reitteri (Grouvelle, 1877: 210) [= Laemophloeus reitteri Grouvelle, 1877], new combination; Phloeolaemus semiflavus (Grouvelle, 1876: 497) [= Laemophloeus semiflavus Grouvelle, 1876], new combination; Phloeolaemus sharpi (Hetschko, 1929: 41) [= Laemophloeus sharpi Hetschko, 1929], new combination; Phloeolaemus straminipennis (Reitter, 1876: 47) [= Laemophloeus straminipennis Reitter, 1876], new combination; Phloeolaemus teapensis (Grouvelle, 1876: 494) [= Laemophloeus teapensis Grouvelle, 1876], new combination.
540
A new genus and two new species of Argentine Rhinotragini
(Coleoptera, Cerambycidae, Cerambycinae)
(2017)
Rhopalessa irwini sp.nov. and Rhinion parkeri gen. nov., sp. nov. (Coleoptera, Cerambycidae, Cerambycinae, Rhinotragini) are described from Argentina. A key to species of Rhopalessa is provided.
539
Six annotated lists are presented: A, a checklist of the butterflies (Lepidoptera, Papilionoidea) of Tobago (150 species); B, species for which there are no records in the last 80 years (49 species); C, species needing confi rmation from Tobago (5 species); D, species not accepted from Tobago (12 species); E, species which are likely to occur in Tobago, but have not been recorded (6 species); and F, species and subspecies recorded from Tobago, but not from Trinidad (2 species and 2 subspecies). Remarkably, 33% of the 150 recorded species have not been reported
in the last 80 years. While it is possible that some of these are not resident or have become extinct, it seems more likely that most have simply not been found in the last 80 years. The butterfly fauna of Tobago merits further study; year-round collecting in different habitats and areas, using a variety of techniques, will surely fi ll in many of the apparent gaps in our knowledge. Ouleus fridericus sheldoni ssp. nov. (Hesperiidae, Pyrginae) is described from Tobago, with illustrations of adults and male genitalia, and is compared to O. fridericus sinepunctis (Kaye) from Trinidad. Danaus plexippus tobagi A.H. Clark, 1941 is a syn. nov. of D. plexippus nigrippus (Haensch, 1909) (Nymphalidae, Danainae).
538
The female of Callichroma magnifi cum Napp and Martins, 2009 (Coleoptera: Cerambycidae: Cerambycinae: Callichromatini) is described and illustrated for the fi rst time. The geographical distribution of the species is expanded to the Colombian Caribbean and information on the collection site is provided.
537
The Republic of Panama currently has 21 recorded species of stoneflies, all in the genus Anacroneuria (Plecoptera: Perlidae). Herein, we record five species of this genus from the Mount Totumas Cloud Forest and Biological Reserve, in the upper reaches of the Río Chiriquí Viejo watershed. One of these species, A. plutonis (Banks), represents a new country record for Panama. These results are part of an ongoing effort to characterize the aquatic insect fauna of Panama, and to evaluate that country’s major watersheds.
536
A new genus, Dejunaleon Miller and Stange, is described with two new species: Dejunaleon loja Miller and Dejunaleon maculosus Stange (Neuroptera: Myrmeleontidae: Brachynemurini). A key to species is provided as well as distributional data. A description of the larva of Dejunaleon loja is given. A key to genera of the tribe Brachynemurini is provided based on larvae and adults.
535
Aphis elena Lagos-Kutz and Voegtlin, sp. nov. (Hemiptera: Aphididae), is described from specimens collected in Illinois, USA, on the North American native plant, Pycnanthemum virginianum (L.) T. Dur. & B.D. Jacks. ex B.L. Rob. & Fernald (Family: Lamiaceae). Both apterous and alate viviparae are described and illustrated, and a dichotomous key is provided to apterous and alate viviparae of the six species of Aphis that have been recorded on plants in the family Lamiaceae in North America. Neighbor-joining analysis of cytochrome oxidase 1 (Cox1) indicated a close relationship of the new species with Aphis monardae Oestlund, which also feeds on a member of Lamiaceae. The range of pair-wise distances for DNA barcoding of these species is 0.17–0.33%. The newly described Aphis elena is morphologically more similar to Aphis gossypii Glover than it is to Aphis monardae.
534
533
Females of Traumatomutilla André (Hymenoptera: Mutillidae) are organized into 14 species groups; 136 of the 138 Traumatomutilla taxa known from females were studied. Of these species groups, only the inermis species group was adequately established and diagnosed in the literature (Casal 1969). The remaining newly established species groups are as follows: americana, auriculata, bellica, bifurca, diabolica, gemella, indica, integella, juvenilis, quadrinotata, tabapua, trochanterata, and vitelligera. Diagnoses and lists of included species are provided for each species group. Eighty-three new country records are provided for 49 species.
532
Effective island conservation depends on thorough biodiversity surveys and species assessment. The U.S. Fish and Wildlife Service in coordination with Point Blue Conservation Science undertook a two-year insect survey of the Farallon Islands, California, in order to catalog current insect and spider species on the island and to gather information that will be used to examine prey dynamics to aid in the conservation of the endemic salamander Aneides lugubris (Hallowell). The report lists 11 insect orders representing 60 families, 107 genera and 112 insect species on Southeast Farallon Island. Holometabolous orders were the most represented on the island with Coleoptera and Diptera being the most abundant, followed by Lepidoptera and Diptera. One spider order was identified, representing six genera and six species.
531
530
Nine new species of Rhinotragini (Coleoptera: Cerambycidae: Cerambycinae) are described from the Neotropical region: Odontocera galileoae (Mexico); Odontocera mthomasi (Guatemala); Ecliptoides vandenberghei (Nicaragua); Eclipta ricei and Odontocera skelleyi (Costa Rica); Eclipta nearnsi (Panama); Odontocera stangei (Venezuela); Chariergodes lingafelteri and Ischasia martinsi (Costa Rica and Panama). Ischasia rufi na Thomson, 1864 is formally excluded from the fauna of Costa Rica and Panama, and Nicaragua is added as a new country record for Eclipta fi cta Bezark, Martins and Santos-Silva, 2013. Notes on Ecliptoides monostigma (Bates, 1869)
are provided in the “Diagnosis” of E. vandenberghei.
529
The genus Speorthus Chamberlin is revalidated for S. tuganbius Chamberlin and S. reyesi, new species (Diplopoda, Polydesmida, Macrosternodesmidae). The families Polydesmidae and Macrosternodesmidae (as they occur in North America) are briefly discussed, and their component North American genera listed. The validity of Macrosternodesmidae is reaffi rmed; the family is predominantly North American. Nearctodesminae, new status, is formally designated as a subfamily of Macrosternodesmidae. The superfamily Trichopolydesmoidea is synonymized with the superfamily Polydesmoidea, new combination.
528
This revision concerns a small group of Western Palaearctic Copris species (Coleoptera: Scarabaeoidea:
Scarabaeidae) distinguished by having three lateral teeth on the foretibae. According to the literature, this group consists of four taxa: Copris armeniacus Faldermann, 1835, C. felschei Reitter, 1892, C. pueli Mollandin de Boissy, 1905 and C. umbilicatus Abeille de Perrin, 1901. Copris armeniacus is herein deemed a species inquirenda, and C. felschei, for which a neotype is designated, is deemed valid. Copris umbilicatus is recorded for the fi rst time from Turkey. A key to all species of the genus Copris known from the Western Palaearctic is provided. Variability of the cephalic and pronotal armature, and morphology of the parameres are illustrated.
527
The melon fly, Bactrocera cucurbitae (Coquillett) (Diptera: Tephritidae), is a widespread, economically important tephritid fruit fly species. Bactrocera cucurbitae infests fruits and vegetables of a number of different plant species, with many host plants in the plant family Cucurbitaceae, but with additional hosts scattered across many other plant families. Although thought to be native to India, its distribution has spread throughout many countries in Oriental Asia, into a number of Pacifi c Island nations, and into Africa. The documented introductions into countries outside its native distribution show that this species could establish in other countries where it does not presently occur, particularly through the movement of infested fruit. As with other tephritid fruit fly species, establishment of B. cucurbitae can have signifi cant economic consequences, including damage and loss of food production, as well as requirements for implementation of costly quarantine treatments to permit export of commodities susceptible to infestation by B. cucurbitae and inspection of susceptible imported commodities. In order to avoid these adverse economic consequences, one needs to prevent the entry, establishment and spread of B. cucurbitae into a new habitat. To successfully achieve this, an accurate knowledge of the fly’s host plants is essential. Cognizant of this need, we prepared, and present here, a worldwide list of host plants for B. cucurbitae, with annotations on reported laboratory and fi eld infestation data. Overall, 136 plant taxa from 62 plant genera and 30 plant families are identifi ed as hosts of B. cucurbitae, based on reported fi eld infestation data. The predominant family, as expected, is Cucurbitaceae, with 56 plant taxa (41.2% of all host plant taxa) in which fi eld infestation by B. cucurbitae has been documented. The family with the 2nd highest number of documented infested plant taxa is Solanaceae, for which there are published fi eld infestation data for 20 plant taxa (14.7% of plant taxa for which there is documented fi eld infestation). Papers that list plants as hosts of B. cucurbitae based only on laboratory data, those that list plants as a host but do not report any fi eld infestation data, and those that report interception data add an additional 137 host plant taxa, representing a total of 80 genera and 39 plant families, 20 of which are additional plant families beyond those for which there is fi eld infestation data. These additional species must be considered “undetermined” hosts for which additional data are needed to document actual host status. This paper is a comprehensive documentation of host plants of the melon fly based on recorded infestations in laboratory and/or fi eld, interceptions at ports of entry, or “listing only” associations. Host records presented here will be used in vetting and developing the offi cial USDA list of host plants of the melon fly, which will be published by APHIS as a federal order.
526
525
The first myrmecophilous fl ea beetle genus (Myrmeconycha Konstantinov and Tishechkin, new genus) with four new species (M. erwini Konstantinov and Tishechkin, new species – Ecuador, M. gordoni Konstantinov and Tishechkin, new species – Brazil, M. pakaluki Konstantinov and Tishechkin, new species – Panama, and M. pheidole Konstantinov and Tishechkin, new species – Costa Rica) is described and illustrated. It is compared with fl ea beetles of the subtribe Disonychina (Coleoptera: Chrysomelidae: Galerucinae: Alticini) and may be easily differentiated based on the external and internal features, which include the waxy surface of the head and pronotum, reticulated surface of the pronotum, and four longitudinal ridges on each elytron.
524
A new species of Autostichidae, Eraina beechei nov. sp., is described from specimens collected in the Andean mountains of Ñuble, Biobío. Diagnostic characters for the species are presented; photographs of adults, illustrations of male and female genitalia, and a key for identifi cation of Eraina species are provided.
523
The species of Dipropus Germar (Coleoptera: Elateridae) of the west-central region of North America are taxonomically reviewed. Historical records of D. approximatus (Candèze), D. ferreus (LeConte), D. simplex (LeConte) and D. soleatus (Say) in the region are based on misidentifi cations. Dipropus pericu new species is described from Baja California Sur. Dipropus reinae new species, D. sonora new species and D. yaqui new species are described from southern Arizona, New Mexico and Sonora. Dipropus warneri new species is described from Arizona, New Mexico and Texas. Dipropus sus (Candèze) is reported from Morelos, Sonora and Sinaloa as new state records. Keys to the species of the region are provided. The historical value of a specimen of Elater soleatus Say in the Dejean collection is discussed and its lectotype designated. Ischiodontus oblitus Candèze is treated as an objective synonym of Dipropus soleatus.
522
Ptyoiulus Cook 1895, the dominant parajulid diplopod genus in the eastern United States (US), comprises two species – P. impressus (Say 1821), with a slanted, fl ared, circumferentially entire, and marginally serrate apical calyx on the anterior gonopod coxal process, and P. montanus (Cope 1869), n. comb., with a smooth, upright, cupulate calyx that is open caudad and coaxial with the process’ stem. The genus occupies a broad area between the Mississippi River and Atlantic Ocean extending from southern New England, Ontario, and Michigan to the Florida Panhandle and four small disjunct ones – from Montreal, Québec, to northern Vermont, along southwestern Lake Michigan in Wisconsin and Illinois; northeastern/eastcentral Arkansas, primarily in Crowley’s Ridge physiographic feature and beside the “bootheel” of Missouri; and a point locality in northeastern Louisiana just south of the Arkansas line. A male from Chester County (Co.), Pennsylvania, is designated as the neotype of Julus impressus, as is one from Durham Co., North Carolina, for J. montanus. As both species inhabit Montgomery Co., Virginia, the type locality of J. montanus, we exercise the right of first reviser, conserve the latter name, and assign it to the species with the smooth, cupulate, and coaxial calyx. We also exercise first reviser rights and assign Parajulus ectenes Bollman 1887 to this form, thereby relegating it to synonymy under Ptyoiulus montanus. Other new synonymies include Ptyoiulus georgiensis Chamberlin 1943 under P. impressus and P. coveanus Chamberlin 1943 under P. montanus. Both Ptyoiulus and P. impressus are projected for Delaware and Rhode Island and newly reported from Québec, Connecticut, District of Columbia, Maryland, Mississippi, South Carolina, Vermont, West Virginia, and Wisconsin, and the genus and species, respectively, are newly documented from Louisiana and Arkansas; P. montanus is newly cited from Alabama, Arkansas, Georgia, Mississippi, and South Carolina. Ptyoiulus impressus occupies every state except perhaps Louisiana and is the only species in areas that were inundated during the Cretaceous and glaciated during the Pleistocene; by contrast, P. montanus inhabits a relatively narrow east/west transect through the center of the generic range. Their distribution patterns suggest an old species, montanus, being actively displaced by the younger and more successful impressus. The decurvature of the epiproct in uroblaniulinines appears to increase with age and developmental stage. A key is presented to parajulid familygroup taxa in the US and Canada east of the Rocky Mountains.