European journal of taxonomy : EJT
Paris : Muséum National d'Histoire Naturelle
ISSN: 2118-9773
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890:136–164
A new species of Liolaemus is described from southwest of the town of Añelo, Neuquén Province, Argentina. Integrative evidence methodology of external morphological characters and molecular phylogenetic analyses of mitochondrial DNA (cyt-b) is used to place the new species to the species group of Liolaemus boulengeri. The new species is phenotypically close to L. mapuche. The new Liolaemus is medium to large in size (males 77.64–83.98 mm, females 72.88–78.58 mm), with evident sexual dichromatism. Genetic distances of the mtDNA (cyt-b) between the new species and its closest relative species are greater than 3% (L. cuyanus 7.48–12.02%; L. josei 7.56–9.60%; L. puelche 8.23–9.93%; L. mapuche 8.51–9.79%). Molecular and morphological phylogenetic results show L. mapuche as the sister species of the new one. The new species is larger than L. mapuche. Dorsal and ventral scales are more numerous in the new species than in L. mapuche, precloacal pores in females are present in L. mapuche and absent in the new species. It has strict psammophilic habits, using sand mounds and sheltering, under Alpataco (Neltuma alpataco) bushes. The L. boulengeri group now contains 75 species distributed in Argentina, Bolivia, Brazil, Chile, Paraguay, Peru and Uruguay.
870:146-166
We describe a new species of Dravidogecko from the poorly-inventoried Periyar Plateau of Southern Western Ghats, Peninsular India. The new species Dravidogecko beddomei sp. nov. is characterized by: precloacal-femoral pores 46‒52; scales across belly 31‒33; supralabials 9–11; infralabials 8–9; lamellae on digit IV of pes 10–11. Additionally, the new species is divergent (>13%) from its related congeners. Like its congeners, the new species is currently known only from its type locality and its vicinity – the Devar Malai, Kakki Reservoir – and Sivagiri Hills in the Periyar Plateau of Southern Western Ghats. This discovery further increases the list of new reptiles described from this under-studied massif in the Southern Western Ghats.
773:169-186
A new species of echinoderid kinorhynchs, Echinoderes xiphophorus sp. nov. collected from oxidized brown silt at the deepest depression in the Sea of Japan, North-West Pacific, is described and illustrated using light and electron microscopy. This new representative of the most speciose kinorhynch genus is characterized by the unique set of spines and tubes and can easily be distinguished from most of its congeners. The second trunk segment bears three pairs of tubes in subdorsal, midlateral and ventrolateral position in both sexes; one pair of tubes on trunk segment 5 in lateroventral position and on trunk segment 8 in sublateral position; aciculate lateroventral spines on trunk segments 6–9; aciculate middorsal spines on trunk segments 4, 6, 8. This species is well recognized by very long tergal extensions of the posteriormost segment, some of the longest within the family Echinoderidae. Males of Echinoderes xiphophorus sp. nov. are well distinguished from all the congeners by extremely long sword-like appendages dorsally to three pairs of penile spines. The species constitutes the first deep-sea representative of the Echinoderidae in the Sea of Japan and the deepest representative of the Kinorhyncha in this sea.
471
During a survey of the fishes in the region of the Wonga-Wongué Presidential Reserve, 14 new populations of the subgenus Chromaphyosemion Myers, 1924 were found. These observations extend the previously known distribution range of the subgenus 120 kilometres southward. None of these populations could be related to any described species. Based on the colouration of the males and females, together with a genetic marker (mitochondrial DNA cytochrome b sequences), the populations studied are grouped into six new species which are described in this article, all close to Aphyosemion alpha Huber, 1998 with which they share the presence of a black alpha-shaped mark on the pre- and post-opercular region. The group composed of A. alpha and the six new species is referred to here as the A. alpha species group. All the new species, A. aurantiacum Chirio, Legros & Agnèse sp. nov., A. barakoniense Chirio, Legros & Agnèse sp. nov., A. flammulatum Chirio, Legros & Agnèse sp. nov., A. flavocyaneum Chirio, Legros & Agnèse sp. nov., A. pusillum Chirio, Legros & Agnèse sp. nov. and A. rubrogaster Chirio, Legros & Agnèse sp. nov., are further unambiguously diagnosed by unique combinations of colour patterns, making it possible to generate an identification key for the A. alpha species group. It is likely that the coastal dunes of Wonga-Wongué that form a sandy relief, could have led to the fragmentation and then isolation of the hydrographical networks that flow into the Atlantic Ocean, making possible a significant number of allopatric speciations.
817: 35–57
The hoverfly genus Eumerus Meigen, 1822 (Eristalinae: Merodontini) comprises 250+ described species, of which 36 are reported from the Iberian Peninsula. The high species diversity linked to the low degree of morphological differentiation between some species, which is even lower in females, leads to a high taxonomic complexity in this genus. The aim of this work is to confirm the morphological and molecular validity of an undescribed species of Eumerus, which is widespread in the Iberian Peninsula. The new species is described and compared with similar species. The genitalia of the new species are similar to those of Eumerus clavatus Becker, 1923 and Eumerus uncipes Rondani, 1850, but also share some features with Eumerus nudus Loew, 1848. The COI-5’ barcode is provided for the new taxon and analysed together with those of other named Eumerus sequences/species publicly available online. In the light of the morphology and barcoding data, the systematic position of the new species is discussed.
852:43-56
Elusive flaws are identified in techniques widely adopted to organize the Material Examined sections in taxonomic publications, mostly regarding the usage of the term ibidem and the nesting of information such as country and states. Logical errors are identified that prevent objective retrieval of the original information and can hinder or block its interpretation, even in case-by-case analyses. It is demonstrated that the free usage of ibidem in the sense of “same as previous except as follows” compromises the interpretation of data, characterizing bad practice. Solutions are proposed for the precise usage of both the term ibidem and the nesting technique. A new technique for organizing, compressing, and presenting information, called grid-setting, is described and evaluated. Its most notable practical effect is that the Material Examined section becomes literally a coded data sheet, which can be accurately converted back to spreadsheet format. In addition, the grid-setting technique was able to generate texts up to 30% shorter than those edited with the best-known traditional techniques. The new ideas and fixes are incorporated into a new software, flexible enough to process varied and unlimited data into largely user-defined texts, which remain nevertheless universal in their format and logical interpretation.
376
Daddy-long-leg giants: revision of the spider genus Artema Walckenaer, 1837 (Araneae, Pholcidae)
(2017)
This is the first revision of Artema Walckenaer, 1837, a genus consisting of large and phylogenetically interesting species. Even though Artema is not species-rich (now eight nominal species), it has suffered from poor descriptions and synonymies. Our main goal was to gather all available material and to clarify species limits. Four species are easily distinguished from other congeners: Artema atlanta Walckenaer, 1837, the type species; A. kochi Kulczyński, 1901 (revalidated); A. bunkpurugu Huber & Kwapong, 2013; and A. nephilit sp. nov. All other species are problematic for varying reasons: species limits are unclear between A. doriae Thorell, 1881 and A. transcaspica Spassky, 1934; A. magna Roewer, 1960 and A. ziaretana (Roewer, 1960) are problematic because they are based on female and juvenile types respectively and little new material is available. The material available to us suggests the existence of a few further species; however, they are not formally described, either because of small sample sizes (Artema sp. a and A. sp. b are represented by only one specimen each) or because of unclear species limits (between Artema sp. c, A. transcaspica and A. doriae).This study is the first serious step towards understanding the genus. Intensive collecting effort is needed in order to fully clarify species limits.
726:24-37
Pholetesor acrocercophagus sp. nov., P. camerariae sp. nov. and P. indicus sp. nov.(Hymenoptera: Braconidae: Microgastrinae) are described as new to science. These three species were reared from Acrocercops sp., Acrocercops phaeospora Meyrick, 1916 and Cameraria virgulata Meyrick, 1914 (Lepidoptera: Gracillariidae), respectively. Characteristics of these new species and their affinities with related taxa are discussed. Data on habitat, host records and host plant species for all the parasitoid species is provided. A key to the Indian species of the genus Pholetesor Mason, 1981 reared from lepidopteran leafminers is also given.
702
This paper reports on the genus Cobbionema Filipjev, 1922 in Sweden with the description of four species and a revision of the genus. Cobbionema acrocerca Filipjev, 1922 is relatively small in size, with a tail that has a conical proximal and a digitate distal section. Cobbionema cylindrolaimoides Schuurmans Stekhoven, 1950 is similar to C. acrocerca in most characters except having a larger body size and heavily cuticularized mandibles. Cobbionema brevispicula sp. nov. is characterised by short spicules and a conoid tail. Cobbionema acuminata sp. nov. is characterised by a long two-part spicule, a conical tail and three (one mid dorsal and two ventrosublateral) sharply pointed tines in the anterior chamber of the stoma that are located more anterior than in all the other species. We also present a molecular phylogeny of the family based on the nearly full-length 18S and the D2-D3 expansion segment of the 28S rRNA genes. Maximum Likelihood and Bayesian trees inferred from both genes strongly support a clade that included Cobbionema, Demonema Cobb, 1894 and Halichoanolaimus de Man, 1888 and another clade with Gammanema Cobb, 1920 and Latronema Wieser, 1954 nested together. None of the trees supported the monophyly of the subfamilies Choniolaiminae and Selachinematinae.
888:97-110
Here, I describe four new species of the Neoserica vulpes species group: Neoserica daxue sp. nov., N. mianningana sp. nov., N. myanmarensis sp. nov., and N. yanyuan sp. nov. Genitalia and habitus of the new species are illustrated. Additional records of species from the group are given and the key to species of the Neoserica vulpes species group is updated.
921:251-275
The Cladonematidae are a family of hydrozoans with a worldwide distribution and morphological adaptations for a benthic mode of life. Species of this family are characterized by high morphological variability, which has caused many taxonomical debates, mainly for the species of the genera Eleutheria Quatrefages, 1842 and Staurocladia Hartlaub, 1917. Herein, we describe Staurocladia dzilamensis sp. nov., a new species of crawling hydromedusa from the southern Gulf of Mexico. This finding also constitutes the first record of the genus Staurocladia for the Gulf of Mexico. The presence of additional nematocyst clusters, supplementing the apical one on the upper branch of the tentacles, places it within Staurocladia. The presence of exumbrellar buds, a conspicuous marginal ring of nematocysts, 6–11 bifid tentacles with lower branches longer than their upper counterpart, the cnidome with stenoteles of two size classes, and two nematocyst clusters on the upper branch supplementing the apical one, opposite placed alternately on its aboral and oral sides permits to differentiate S. dzilamensis from its congeners. A taxonomic key for the species of Staurocladia is provided.
717:108–128
This paper treats the planthopper genus, Atracis Stål, 1866 (Flatidae: Flatoidinae) in China. Five new species, A. ocularia sp. nov., A. patefacta sp. nov., A. punctulata sp. nov., A. sphaerica sp. nov. and A. ungulata sp. nov., are described and a checklist of all its known species from China is provided.
193
The variability of the reticulum pattern, ornamentation and outline of the Urocythereis populations of the La Strea Bay is analysed. The results show that the shell features of the form U. distinguenda (Neviani, 1928) (= U. oblonga Brady, 1866) have to be included within the high variability range of U. margaritifera (G.W. Müller, 1894). Consequently, it is suggested that in the upper infralittoral waters of the inlet two (and not three, as stated in previous investigations) species of the genus Urocythereis presently live. A second form, displaying a relatively low variability, is described as a new species, U. ilariae sp. nov.
908:77-107
The ant genus Vollenhovia Mayr, 1865 (Myrmicinae, Crematogastrini) mostly occurs in the Australasian and Oriental regions. We revised its diversity in India and Sri Lanka based on qualitative and quantitative morphology, recognizing eleven taxa, including a new species which is described herewith: V. escherichi Forel, 1911, V. gastropunctata Bharti & Kumar, 2013, V. karimalaensis Dhadwal et al., 2023, V. keralensis Kripakaran & Sadasivan, 2022, V. mawrapensis Dhadwal et al., 2023, V. oblonga laevithorax Emery, 1889, V. penetrans (Smith, 1857), V. pfeifferi Bharti et al., 2023, V. taylori Rilta et al., 2023, V. terayamai Rilta et al., 2023, and V. yasmeenae sp. nov. The subspecies status of V. oblonga laevithorax and its relationship with V. penetrans, whose type series does not contain workers, still requires to be assessed in the context of a broader revision including the whole Oriental region. The known distribution of the genus in the Indian subcontinent appears to be fragmentary, still requiring extensive sampling efforts. Four species are from the Western Ghats biodiversity hotspot in the southern Indian state of Kerala, one is endemic to the biogeographically related Sri Lanka, three are known from Eastern India near the border with Bangladesh, two are reported from the Andaman and Nicobar Islands, and one is restricted to the Himachal Pradesh in northern India. A comprehensive key of the known Vollenhovia species from India and Sri Lanka is provided.
575
We revise the genus Attemsostreptus Verhoeff, 1941 based on type material of the type species, A. costatus Verhoeff, 1941, synonymise A. orobius (Kraus 1958) with A. costatus and describe a second species of the genus, A. reflexus sp. nov., collected from Kimboza Forest Reserve in Tanzania, and discuss the dubious tribe Trachystreptini.
295
In this work we present a revision of the genus Ommatoiulus Latzel, 1884 in Portugal. Based on recently collected material and older museum samples, including type specimens, we describe six new species to science, viz. Ommatoiulus alacygni sp. nov., O. camurus sp. nov., O. denticulatus sp. nov., O. litoralis sp. nov., O. staglae sp. nov. and O. stellaris sp. nov. The species O. alacygni sp. nov., O. denticulatus sp. nov. and O. staglae sp. nov. described from the Algarve are outstanding by their extremely reduced mesomerital process. The species O. porathi (Verhoeff, 1893) and O. andalusius (Attems, 1927) are recorded and redescribed for the first time after their original description. The finding of O. andalusius – originally described from Andalusia in Spain – constitutes a new record for Portugal together with two species, viz. O. fuentei (Brolemann, 1920) and O. martensi Mauriès, 1969. The taxonomic status of several species is revised. Thus Archiulus (Schistocoxitus) cingulatus Attems, 1927 is here considered as a junior synonym of Ommatoiulus lusitanus (Verhoeff, 1895) while Schizophyllum cervinum Verhoeff, 1910 is synonymized with Ommatoiulus moreleti (Lucas, 1860). An identification key to all hitherto known Portuguese species of Ommatoiulus is presented as well as a distribution map illustrating the various species occurrences in the country.
497
This paper is part of a project of studying benthic diatom biodiversity on marine coastal regions of Sweden with focus on rare and less known species. Two new species of Cocconeis Ehrenb. are described from Vrångö, a small island in the west coast of Sweden. Both species were found as epiphytic on the green alga Ulva intestinalis L. Cocconeis magnoareolata Al-Handal, Riaux-Gob., R.Jahn & A.K.Wulff sp. nov. is a small species not exceeding 9 μm in length and characterized by having large subquadrangular areolae on the sternum valve. Cocconeis vrangoensis Al-Handal & Riaux-Gob. sp. nov. appears similar to some taxa of the 'Cocconeis scutellum complex', but differs by its stria density on both valves and variable features of the areola and valvocopula ultrastructure. Detailed descriptions based on light and electron microscopy examination, a comparison with closely related taxa, as well as a description of the habitat of both species are here presented.
145
Keys and diagnoses of North European aphids (Hemiptera, Aphidoidea) associated with mosses, horsetails and ferns are given, based on fresh and freeze-dried material. Numerous externally visible and thus informative characters, that are absent in cleared, slide-mounted specimens, such as body shape colours, wax coating and pattern etc., are utilized. Most of the species are illustrated by photographs of live specimens and drawings. Root-feeding species living in the moss layer or otherwise often present in moss samples are also included, even if their hosts were spermatophytes. The combination of colour images and diagnoses, utilizing easily observed characters, allows the identification of a large number of species already in the field, and many more at home with the aid of a stereo microscope. Host plant relationships and association with ants are summarised, including new records. Brief accounts on aphid life cycles, freeze-drying preparation techniques, etc. are also given to support the use of the keys.
338
Keys and diagnoses of North European aphids (Hemiptera, Aphidoidea) feeding on conifers are given, including species from nearby areas of Central and Western Europe, based on live and freeze-dried material. Externally visible informative characters, such as body shape, colours, wax coating, and pigmentation pattern are utilized, in addition to characters traditionally used in the literature. Rich illustrations with photographs of live colonies and freeze-dried specimens, supported by drawings where needed, are presented. The combination of colour images and diagnoses, utilizing easily observed characters, allows the identification of a large number of species already in the field, and many more at home with the aid of a stereo microscope. Host plant relationships and aphid-ant associations are presented.
4
Review of Neurepyris Kieffer, 1905 (Hymenoptera, Bethylidae), a new synonym of Pristocera Klug, 1808
(2011)
The genus Neurepyris Kieffer, 1905 is revised. The adult male holotypes N. rufiventer Kieffer, 1913 from Eritrea and N. tagala (Ashmead, 1905) from the Philippines are redescribed and illustrated. Both species are transferred from the subfamily Epyrinae to Pristocerinae because they have the metanotum well developed medially. Neurepyris rufi venter is transferred to Pristocera Klug, 1808 because the hypopygium is deeply divided into two apical lobes, the subdiscoidal and cubital veins do not reach the border of the forewing, the pronotal disc has the anterior region slightly elevate medially, and the stigma is elongate. Neurepyris tagala is transferred to Apenesia Westwood, 1874 because the basal tooth of mandible is not curved inward, the median lobe of clypeus is not depressed near the antennal insertions, and the aedeagus consists of one lamina. The genus Neurepyris is considered a junior synonym of Pristocera because its type species is N. rufiventer.
885:1-20
In this study, we revise the taxonomy of the endemic Mexican genus Ceratotrupes Jekel, 1865. It comprises three species, distributed in the Sierra Madre Occidental and Trans-Mexican Volcanic Belt (Ceratotrupes fronticornis (Erichson, 1847) and Ceratotrupes bolivari Halffter & Martínez, 1962), and the Sierra Madre del Sur (Ceratotrupes sturmi (Jekel, 1865)). We confirm C. mniszechi (Jekel, 1865) as a junior subjective synonym of C. sturmi. The examination of the holotype of C. sturmi and specimens recently collected in the Sierra Sur de Oaxaca (Oaxaca) and Omiltemi (Guerrero) allowed us to identify and describe a new species, Ceratotrupes gonzaloi sp. nov. from Omiltemi, Guerrero. We redescribe the morphology of C. sturmi and provide a diagnostic key to the four members of the genus Ceratotrupes. Photographs of the habitus and male genitalia of the four species, a distribution map, and observations on their ecology and biogeography are provided.
683
Two new species of Grandidierella Coutière, 1904 from Singapore waters are described based on specimens collected during the Comprehensive Marine Biodiversity Survey (CMBS) between 2010–2015. Grandidierella pawaiensis sp. nov. differs from the rest in the genus by a combination of characters; (1) gnathopod 1 carpus proximoventral corner with apically acute process, (2) inner face of carpus of gnathopod 1 proximoventral corner with curved process, (3) proximal margin of dactylus of gnathopod 1 male with excavation, and (4) gnathopod 2 basis anterior margin crenulate. Grandidierella sungeicina sp. nov. on the other hand, is distinguishable from other related species of Grandidierella found in the tropical region by its paired mid-dorsal projections on pereonites 2 to 5, 1 ventrodistal triangular projection on pereonite 4, and gnathopod 2 ischium with 1 large trapezoid projection.
746:1-25
Field studies of the inselberg flora of north-eastern Brazil have resulted in the discovery of much greater morphological diversity of the genus Ameroglossum, than previously envisaged. These enigmatic plants are endemic to isolated rock outcrops, which have long been surrounded by unsuitable habitat. The morphological forms are geographically isolated and stable in cultivation when grown under similar conditions. We here provide detailed descriptions of the two previously known species and propose an additional seven new species in the genus. A taxonomic revision of the genus, including an identification key is provided. We hypothesize that the evolution of this genus is driven by the diversity of their hummingbird pollinators. Due to the threatened nature of inselberg habitats and the small populations found for most taxa, the species described here are likely to be under threat and in need for formal redlisting.
778:148-166
Three species of Corethrella Coquillett, 1902 from the state of Amazonas, Brazil are described as new to science based on female adult specimens. Corethrella cabocla Feijó, Belchior, Marialva & Pessoa sp. nov. possesses four large setae on the frons between the ventromedial area of ommatidia, a wide clypeus with 1–4 setae, a wing with the apex of R2 basal to the apex of M2 and with a midlength band, and with the abdomen entirely dark brown. Corethrella ielemdei Feijó, Ramires, Lima & Pessoa sp. nov. possesses an elongated coronal suture, four large setae on the frons between the ventromedial area of ommatidia, a clypeus squarish with 42–43 setae, a wing with the apex of R2 basal to the apex of M1 and with a midlength band and dark scales on the basal and subbasal areas of the anterior margin, legs with dark scales, and with the abdomen entirely dark brown. Corethrella menini Feijó, Picelli, Ríos-Velásquez & Pessoa sp. nov. possesses wings with the apex of R2 basal to the apex of M2 and a midlength band, with darker basal scales along all veins, basal band dark scales on C, Sc, R, M, and Cu and the abdomen entirely dark brown. With the addition of the new species, the numbers of frog-biting midges described in the Amazon basin, Brazil and in Neotropical region are now 31, 49 and 80 species, respectively.
762:1-48
We describe one new tribe, two new genera and three new species of the subfamily Cypridopsinae Kaufmann, 1900 from Brazilian floodplains. Brasilodopsis gen. nov. belongs in the nominal tribe Cypridopsini, and both new species in this new genus were found in both sexual and asexual populations. Brasilodopsis baiabonita gen. et sp. nov. has a wide distribution and was found in three of the four major Brazilian floodplains. Brasilodopsis amazonica gen. et sp. nov. was recorded only from the Amazon floodplain. Brasilodopsis baiabonita gen. et sp. nov. has a subtriangular shape in lateral view, whereas Brasilodopsis amazonica gen. et sp. nov. is more elongated and has more rounded dorsal margins in both valves, as well as more pronounced external valve ornamentation, consisting of rimmed pores in shallow pits. Paranadopsis reducta gen. et sp. nov. was found in asexual populations in the Upper Paraná River floodplain only and differs from other Cypridopsinae in the more elongated carapace, an A1 with strongly reduced chaetotaxy (hence the specific name) and the total absence of caudal rami in females. Because of these strong reductions in valve and limb morphology, Paranadopsini trib. nov. is created within the Cypridopsinae for this intriguing new genus and species.
839:149-175
The genus Seticornuta Morley, 1913 currently comprises nine described species; here, nine new species are described: S. anchanchu sp. nov., S. carinata sp. nov., S. cuckoo sp. nov., S. curupira sp. nov., S. flava sp. nov., S. muqui sp. nov., S. nigroflava sp. nov., S. quilmes sp. nov., and S. rufa sp. nov. The genus is redescribed to encompass the features found in Neotropical species and distinctions between the species in the New World and Old World are presented. The genus is recorded for the first time for Argentina, Bolivia, Ecuador, Guatemala and Peru.
885:99-133
Eight new species of Itauara Müller, 1888 are described based on specimens collected in southeastern Brazil: I. bispinata sp. nov., I. caparao sp. nov., I. cipoensis sp. nov., I. cristata sp. nov., I. holzenthali sp. nov., I. mangaratiba sp. nov., I. robertsonae sp. nov., and I. rupicola sp. nov. In addition, we provided an updated distributional list of species of Itauara, with new records of I. jamesii Robertson & Holzenthal, 2011 and I. lucinda Robertson & Holzenthal, 2011 for Espírito Santo State (Brazil), I. plaumanni (Flint, 1974) for Paraná State (Brazil), and I. tusci Robertson & Holzenthal, 2011 for Minas Gerais State (Brazil). We performed an equal weighted parsimony analysis adding the new species to the dataset provided by Robertson & Holzenthal (2013), with modification of the interpretation of some morphological characters. The genus was recovered as monophyletic, but overall statistic support for clades was weak. Itauara is restricted to South America and has a disjunct distribution, with some species occurring in northern South America and others in southeastern South America. Probably, the diversification of Itauara in South America is related to the connections between the Amazon Forest and the Atlantic Forest through historical events at different times.
318
The Middle East biting midges of the tribes Palpomyiini (20 species in three genera) and Sphaeromiini s. lat. (six species in five genera) are reviewed. Three new species are described and illustrated: Bezzia libanensis Alwin & Szadziewski sp. nov., B. sharjahi Alwin & Szadziewski sp. nov. and Palpomyia freidbergi Alwin & Szadziewski sp. nov. Bezzia aegyptia Kieffer, 1925 is recognized as a new junior synonym of B. albicornis (Meigen, 1818) (syn. nov.) and B. omanensis Boorman & van Harten, 2002, is recognized as a junior synonym of B. (Sivabezzia) pachypyga Remm, 1974 (syn. nov.). Keys to the genera and species of the tribes Palpomyiini and Sphaeromiini of the Middle East are also provided.
874
Family Corethrellidae has a worldwide distribution with most known extant species occurring in the Neotropics, many with only the adult stages described. Despite previous efforts, one remaining problem in the taxonomy of Corethrellidae is the challenge of delimiting species due to the current range of morphological characters included in descriptions to date. This paper discusses additional morphological evidence to distinguish species and their phylogenetic relationships. Together with fresh samples collected in Bahia, the only state in the Northeast Region with records of corethrellids, and slide-mounted specimens from other regions, five new species are described: Corethrella pindorama sp. nov., C. patasho sp. nov., C. fuscifimbria sp. nov. and C. unifasciata sp. nov. based on female adults; and C. bifida sp. nov. based on male and female adults, and larval and pupal exuviae. We also provide complementary descriptions comprising the novel characters for 33 previously described species, including the immature stages of nine. Fifteen species are newly recorded for Bahia and the Northeast Region, of which two are new records for Brazil. Additionally, we provide remarks about the biology of species, an updated list of species recorded in Brazil and an updated taxonomic key of New World species, modified from Borkent (2008).
821:102-149
The pseudoscorpion (Arachnida: Pseudoscorpiones) fauna of mainland Ecuador, excluding the Galápagos Islands, is poorly known, with only 41 described species in 9 families. The family Syarinidae has a pantropical distribution and presently comprises ca 120 species in 17 valid genera that are found in leaf litter and subterranean habitats, mostly in tropical and subtropical climates. Four syarinid species have been recorded from Ecuador, including the Galápagos, in two widespread genera, Ideobisium and Ideoblothrus, but field collections suggest that these pseudoscorpions are common and abundant in all forest systems across the country. Here, we review field collections of syarinids from mainland Ecuador and describe five new species in these genera: Ideobisium kichwa sp. nov. (Napo Province, Colonso Chalupas Natural Reserve), I. sonqo sp. nov. (Napo Province, Colonso Chalupas Natural Reserve), I. susanae sp. nov. (Napo Province, Jatun Sacha Natural Reserve), Ideoblothrus nadineae sp. nov. (Napo Province, Colonso Chalupas Natural Reserve) and I. safinai sp. nov. (Pichincha Province, Otongachi Natural Reserve) based on morphology. These species seem to have narrow distributions and we therefore restrict the ranges of two species previously recorded from Ecuador (Ideobisium crassimanum Balzan, 1892 and Ideoblothrus costaricensis (Beier, 1931)) to their countries of origin, which is Costa Rica and Venezuela, respectively.
723:108-134
The genus of Neotropical frogs Pithecopus includes 11 species occurring east of the Andes from southern Venezuela to northern Argentina. Recent genetic approaches pointed out an unusual genetic diversity among populations from localities in north-eastern Brazil recognized as P. nordestinus. In fact, one of these studies confirmed the hypothesis that the São Francisco River acted as an effective geographical barrier during vicariant events in the evolutionary history of P. nordestinus, resulting in two principal, highly divergent clades. Herein we formally describe this divergent clade as a new cryptic species of Pithecopus from north-eastern Brazil, the sister clade of P. nordestinus. It differs from other species of Pithecopus, except for P. azureus and P. nordestinus, by its small body size, lack of the reticulate pattern on flanks, smaller head width, and advertisement calls generally composed of a three-pulsed core.
679
The Neotropical frog genus Pseudopaludicola includes 25 species distributed throughout South America. Herein we review the taxonomic status of P. parnaiba relative to P. canga and the specific identity of the population treated in previous studies as Pseudopaludicola sp. 3 from Barreirinhas in the Brazilian state of Maranhão. The lack of differentiation in advertisement call, morphology, and mitochondrial markers from topotypes and different populations rejects the status of P. parnaiba and Pseudopaludicola sp. 3 from Barreirinhas as distinct species. For these reasons, we suggest to formally consider P. parnaiba as a junior synonym of P. canga. We also found that a population previously reported as P. facureae from central Brazil (Palmeiras de Goiás, Goiás) corresponds to a cryptic species that we describe here as a new species. Lastly, we provide for the first time the phylogenetic positions of P. giarettai, P. llanera and P. pusilla.
480
The genus of Neotropical frogs Pseudopaludicola Miranda-Ribeiro, 1926 includes 23 species, which occur throughout South America. Herein we describe a new species of Pseudopaludicola from the central region of the state of Minas Gerais (southeastern Brazil). This new species is distinguished by the adult morphology, the advertisement call, and molecular data. It is diagnosed mainly by its small size, terminal phalanges knobbed (lack of any expansion of the digital tips), proportionally short hind limbs, and its advertisement call composed of series of two-pulsed notes, emitted at a high repetition rate. In addition, we report the occurrence of the new species in the southern foothills of the Serra do Cipó National Park, at elevations lower than 800 m above sea level (a.s.l). The sister species of this new taxon is P. mineira Lobo, 1994, which occurs in rupestrian grasslands above 1200 m a.s.l. at the Serra do Cipó National Park and in Serra do Cabral, both in the state of Minas Gerais, Brazil. We also redescribe the advertisement call of P. mineira based on recordings from topotypical males and compare it to calls of related species.
592
This paper describes a novel species of the genus Heterophoxus Shoemaker, 1925 from Brazil. The material examined was collected during the Mini Biological Trawl Project, off southeast and southern Brazil's coast off the states of Rio de Janeiro, Santa Catarina, and Rio Grande do Sul, ranging in depth from 25 to 128 meters. The new species, H. shoemakeri sp. nov., is similar to H. videns K.H. Barnard, 1930; however, the former species can be distinguished by: pereopod 7 basis posterior margin serrate with acute teeth, merus posterior margin not produced; epimeral plate 3 posteroventral corner with a longer acute projection, posterior margin almost straight; uropod 3 outer ramus article 2 clearly notable in both sexes. For Brazilian waters, this is the second record of the genus and the first species described with type material from the country. Besides that, we provide an identification key for the world species of Heterophoxus and a distribution map for the new species.
768
This is a first inventory of Isopoda Anthuroidea from near-shore marine sites in Sulawesi. The material was collected from reefs, sea-grass beds and mangroves of the Minahasa Peninsula in North Sulawesi. Specimens were washed out from benthos samples collected in shallow water. The new species Hyssura reptans sp. nov., Kupellonura indonesica sp. nov., Kupellonura macaroni sp. nov., Pendanthura bangkaensis sp. nov., Expanathura marcoi sp. nov., and Colanthura gerungi sp. nov. are described. We discovered that in Colanthura the mandible is fused to the cephalothorax, a unique feature among isopods. Expanathura collaris (Kensley, 1979) is described from Bangka Island and several differences between other Indo-Pacific populations are discussed.
134
Morphological (shell) and molecular examination of a large suite of specimens of pleurotomariids from around New Caledonia and the Coral Sea reveals the existence of four species in the complex of Perotrochus caledonicus: Perotrochus deforgesi Métivier, 1990 and P. pseudogranulosus sp. nov. live allopatrically on the plateaus and guyots of the Coral Sea; Perotrochus caledonicus Bouchet & Métivier, 1982 and Perotrochus wareni sp. nov. live sympatrically - but essentially not syntopically - on the slopes of New Caledonia, Norfolk Ridge and the Loyalty Ridge. All species live in the 300–500 m interval, and together form a significant component of the mollusc fauna living on hard bottoms in the SW Pacific, with individual dredge hauls containing up to 25 specimens of Perotrochus.
819: 90–107
A new species of the genus Caucaseuma Strasser, 1970, Caucaseuma strasseri Antić sp. nov., is described from a cave in the Western Caucasus, representing the eighth species of the genus, and the fourth presumed troglobiontic Caucaseuma. An updated key to and a distribution map of all eight species of Caucaseuma is presented, including new records. In addition, the cave millipede fauna of the Caucasus is briefly discussed, with the inclusion of the most relevant references.
908:39-76
The present paper is devoted to the description of three new species of the genus Acanthophorella Antić & Makarov, 2016 from Georgia, western Caucasus: A. devi Antić sp. nov. and A. valerii Antić sp. nov., both presumed troglobionts and each from a single cave, and the epigean A. aurita Antić sp. nov. The troglobiotic Acanthophorella barjadzei Antić & Makarov, 2016 is reported from two additional caves, with further descriptive notes given. Notes on the ecology and localities, and a distribution map for all six species of the genus are presented. A key is given to all members of the Flagellophorella complex. The relationships within the complex and the distribution and troglomorphism in the genus Acanthophorella are briefly discussed.
205
A new troglobitic species of the previously monotypic genus Biokoviella Mršić, 1992, B. mosorensis Antić & Dražina sp. nov., is described from caves on Mt. Mosor, Croatia. In addition to this, B. mauriesi Mršić, 1992, is partially re-described, and the taxonomic status of the family Biokoviellidae is re-considered. The genus Biokoviella is placed in the subfamily Biokoviellinae Mršić, 1992 stat. nov. within the family Anthogonidae Ribaut, 1913. The relationship of the genus Biokoviella with other anthogonids is briefly discussed, and a distribution map of the genus is presented. Notes on ecology and coinhabitants of the genus Biokoviella, and new data on some Balkan anthogonids are also included in the paper.
713
The Caucasian leucogeorgiinine genera Archileucogeorgia Lohmander, 1936 and Leucogeorgia Verhoeff, 1930 are revised, with Leucogeorgia being considered as a senior subjective synonym of Archileucogeorgia, syn. nov. The following new combinations are thus warranted: Leucogeorgia abchasica (Lohmander, 1936) and L. satunini (Lohmander, 1936), both comb. nov. ex Archileucogeorgia. All four previously described species, viz., L. longipes Verhoeff, 1930, L. abchasica, L. satunini and L. rediviva Golovatch, 1983, are redescribed based on new material, partly also on the type series, with a lectotype being designated for L. longipes. In addition, eleven new species of Leucogeorgia are described, both with normal (L. borealis sp. nov., L. gioi sp. nov., L. golovatchi sp. nov., L. lobata sp. nov., L. oculata sp. nov. and L. prometheus sp. nov.) and modified mouthparts (L. caudata sp. nov., L. mystax sp. nov., L. profunda sp. nov., L. redivivoides sp. nov. and L. turbanovi sp. nov., all clearly troglobionts). Additionally, a new monotypic genus, Martvilia gen. nov., is erected, with M. parva gen. et sp. nov. as the type species, another presumed troglobiont. An identification key to both genera and all 16 species of Caucasian Leucogeorgiini is presented. Several other members of this tribe are also re-examined, including a syntype male of Telsonius nycteridonis Strasser, 1976, from Greece, herewith designated as the lectotype. Troglomorphisms and mouthpart modifications, as well as the distributions and relationships within both Leucogeorgia and Leucogeorgiini, and a Leucogeorgiini species richness estimate for the western Caucasus are discussed.
244
The Sergipe-Alagoas Basin has one of the most complete, exposed lithological successions of the Cretaceous period in the continental margin of Brazil. It captures several phases of the evolution of the South Atlantic Ocean, including rift, gulf and drift. The upper Aptian–Albian Riachuelo Formation corresponds to the first stages of the southern proto-Atlantic Ocean invasion in that basin. The present study reviews the taxonomic identification and ecology of 39 ostracod species of this formation, proposing a new genus – Gabonorygma gen. nov. – and three new species – Praebythoceratina deltalata sp. nov., Gabonorygma sergipana gen. et sp. nov. and Brachycythere smithsoniana sp. nov. Other taxa include Conchoecia? sp. 1, Cytherella sp. 1, C. besrineensis comb. nov., Cytherelloidea aff. globosa, C. btaterensis, Bairdoppilata sp. 1, Bairdoppilata sp. 2, B. comanchensis comb. nov., B. pseudoseptentrionalis, Robsoniella falklandensis, Cetacella sp. 1, Paracypris eniotmetos, Harbinia sinuata?, H. crepata, Liasina sp. 1, Praebythoceratina amsittenensis comb. nov., P. trinodosa comb. nov., Patellacythere sp. 1, P. shimonensis comb. nov., Xestoleberis? sp. 1, Xestoleberis? sp. 2, Apatocythere? sp. 1, Neocythere? aff. pseudovanveeni, N. (Physocythere) tenuis, Aracajuia antiqua comb. nov., A. benderi, A. fragilis comb. nov., Eocytheropteron sp. 1, Metacytheropteron aff. minuta, Microceratina? sp. 1, M. azazoulensis, Veenia guianensis, Algeriana? sp. 1, Quasihermanites? sp. 1 and Sergipella viviersae.
782:21-54
The unique Neotropical species of the alderfly genus Protosialis van der Weele, 1909, P. bifasciata (Hagen, 1861), is herein transferred to the newly described genus Caribesialis gen. nov. This new taxon is proposed to be sister to the clade Protosialis + Sialis Latreille, 1802, after a phylogenetic analysis that included male genital characters scored on a previous morphological matrix of the family. Also, Ilyobius nigrocephalus sp. nov., a remarkable new species from Ecuador, is described and its phylogenetic position is discussed. Furthermore, Ilyobius bimaculatus (Banks, 1920) from Bolivia, known solely from the female holotype, is redescribed. Based on the present study, the Neotropical fauna of Sialidae is proposed to be constituted by two genera, one insular (Cuba) and one continental (Mexico to Chile and Argentina).
142
Sharon gen. nov. is here described to include Asaphes? amoenus Philippi, 1861 comb. nov. from Chile. A redescription of the species is based on the female holotype and material from different geographic locations. Candèze (1891) placed Asaphes amoenus and Parasaphes elegans in the suprageneric group Asaphites. We discuss differences between Sharon gen. nov. and Hemicrepidius Germar, 1839, where Asaphes amoenus was later placed by Blackwelder (1944). Based on morphological characters, Sharon gen. nov. appears to be related to Parasaphes Candèze, 1881, Wynarka Calder, 1986, and Tasmanelater Calder, 1996, all from Australia, suggesting Gondwanan relationships.
96
The taxonomy of the family Desmodoridae (Nematoda: Desmodorida) is partially revised based on morphology. The diagnoses of the Desmodoridae and the subfamilies Desmodorinae and Spiriniinae are emended to accommodate re-analyzed morphological features. Eight known species are redescribed and the implication of the new findings for the taxonomy of the group is discussed. Amphispira and Metadesmodora are confirmed as genera inquirendae. Alaimonema and Sigmophoranema, and their corresponding type species, are proposed as inquirendae due to poor descriptions of the type material. The other three species of Sigmophoranema are transferred to the genus Onyx because they bear the diagnostic features of this group: spear-like dorsal tooth and s-shape precloacal supplements. Echinodesmodora, Paradesmodora and Stygodesmodora are transferred to the Spiriniinae based on the absence of a head capsule and on the amphidial fovea being surrounded by cuticle striation. Paradesmodora toreutes is transferred to the genus Acanthopharyngoides as A. toreutes comb. nov. The genus Onepunema does not fit in the family Desmodoridae because of diorchic males; thus, it is regarded as taxon incertae sedis.
Lists of valid genera for the two subfamilies are provided. A dichotomic key for the identification of the 14 genera within the Spiriinae is provided.
832
Lycosa Latreille, 1804 (Araneae, Lycosidae) of Israel, with a note on Geolycosa Montgomery, 1904
(2022)
Lycosa (Araneae, Lycosidae) is a wolf spider genus typical of subtropical latitudes in the western Palearctic. Despite being erected over 200 years ago, the taxonomy of Lycosa is still unclear. Many species formerly ascribed to it are currently being moved to other genera, while new species are still being described. The species of Lycosa of the western Mediterranean basin are relatively well known, yet the Levantine region, the easternmost part of the Mediterranean basin, has not received much attention since the early 20th century. Here, we study Lycosa from the southern Levant using morphological, molecular and behavioral characteristics, to delimit the species found in this region. We describe two new species: L. hyraculus sp. nov. and L. gesserit sp. nov. We re-describe the widespread and polymorphic species, Lycosa piochardi Simon, 1876. Lycosa piochardi infraclara Strand, 1913 is synonymized with Lycosa piochardi. By adding novel data to the molecular phylogeny of Lycosa created by Planas et al. (2013) and re-analyzing it, we explore the relationship of the Levantine species to other Mediterranean species of Lycosa. We discuss habitat preferences of the two species of Lycosa. Additionally, we report the burrowing species Geolycosa vultuosa (C.L. Koch, 1838) as a new record to Israel, thus extending the distribution of this species and genus into the Levant.
750:29-51
The following four species new to science are described: Platygaster azarbaijanica Buhl & Asadi sp. nov., Platygaster lotfalizadehi Buhl & Asadi sp. nov., Platygaster karimpouri Asadi & Buhl sp. nov. and Synopeas calecai Buhl & Asadi sp. nov. Diagnostic characters are discussed, and figures are provided to distinguish the new species. In addition, ten species of Platygastrinae Howard, 1892 belonging to the genera Platygaster Latreille, 1809, Synopeas Förster, 1856 and Leptacis Förster, 1856 are reported as new records for the fauna of Iran. Four species of Platygaster and one species of Synopeas are recorded as ʻconferʼ.
774:155-177
The present study focuses on shrimps belonging to the genus Lysmata Risso, 1816, collected from Madagascar during the Atimo Vatae expedition carried out in 2010. Lysmata malagasy sp. nov. is a new species belonging to the clade named “long accessory ramous” or “cosmopolitan” in previous phylogenetic studies. The new species can be distinguished from the only two other representatives of this group in the Indo-west Pacific, L. ternatensis De Man, 1902, and L. trisetacea (Heller, 1861), by the accessory ramus of the lateral antennular flagellum consisting of four elongated articles. Lysmata lipkei Okuno & Fiedler, 2010 is reported here from Madagascar with a remarkable extension of its known range after its original description from Japan. This species has also been reported from Singapore and, as alien species, from Brazil. Lastly, L. kuekenthali De Man, 1902 known from numerous localities in the Indo-West Pacific biogeographic area, is reported for the first time from Madagascar. Results of the present morphological and molecular analyses suggest that L. hochi Baeza & Anker, 2008 from the Caribbean Sea is a synonym of the Indo-West Pacific L. kuekenthali, and thus the latter species is alien in the western Atlantic.
774:42-57
A new alpheid shrimp, Aretopsis sandybrucei sp. nov., is described on the basis of three specimens collected from three deep-water banks in the Chesterfield Plateau of the Coral Sea, between New Caledonia and Queensland, Australia, at a depth range of 280–550 m. The new species is the first known deep-water species of the genus Aretopsis De Man, 1910, with its two congeners, A. amabilis De Man, 1910 and A. manazuruensis Suzuki, 1971, being confined to the lower intertidal and shallow subtidal areas, to a maximum depth of 50 m. Based on morphological grounds, A. sandybrucei sp. nov. can be easily separated from A. amabilis and A. manazuruensis by the generally much stouter third to fifth pereiopods, with their dactyli each bearing a very small secondary unguis on the flexor margin, in comparison to a much stronger one in the other two species. Aretopsis sandybrucei sp. nov. also differs from A. amabilis and A. manazuruensis by the less contrasting colour pattern, including the more translucent, dull yellowish chelae and tail fan. An association of A. sandybrucei sp. nov. with a deep-water hermit crab appears to be highly likely due to the presence of several large hermit crabs (Paguridae) in at least one of the three dredge/bottom trawl hauls, which was also containing a paratype of the new species. The taxonomic status of A. amabilis and A. manazuruensis is discussed.
888:77-96
Panaeolus punjabensis M. Asif, Q. Firdous, A. Izhar, Niazi & Khalid sp. nov. was collected from three different localities (Bahawalnagar, Kasur, and Lahore) in Punjab, Pakistan. Morphological observations and phylogenetic analyses based on nuclear encoded internal transcribed spacers (ITS1-5.8S-ITS2 = ITS) and D1/D2 domain of large subunit (28S) rDNA confirmed the taxonomic distinctness of this species. The new species is potentially hallucinogenic and characterized by a parabolic pileus with a light brown center, broadly fusiform basidiospores, presence of cheilocystidia, pileocystidia, and caulocystidia, and absence of pleurocystidia and clamp connections. The DNA sequences of the species clustered together in a well-supported distinct clade. We present a detailed description, photographs, and line drawings, and elucidate and discuss the phylogenetic position of the new species. Morphological comparisons with phylogenetically and morphologically allied species are discussed.
826:176–187
During mycological surveys of different areas of Punjab, Pakistan, we collected a new species from the genus Candolleomyces. This is the first report of any species of this genus from Pakistan. Candolleomyces asiaticus M.Asif, A.Izhar, Niazi & Khalid sp. nov. is characterized by ellipsoid to oblong-ellipsoid (8.2 × 5 µm) basidiospores with indistinct germ pores, lageniform to utriform cheilocystidia without adhering deposits at the apices, absence of pleuro-, pileo- and caulocystidia, pileipellis with spherical, subglobose to columnar cells, and presence of clamp connections. The morpho-anatomical and molecular phylogenetic analysis of nrITS revealed its distinct phylogenetic position, further supporting the recognition of a new species.