IDF-Report : newsletter of the International Dragonfly Fund
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169
Odonata recorded during two brief sampling trips to the Kapit Town area in Kapit Division, Sarawak in 2020 are reported on. Seventy nine species were recorded, of which at least six (Onychargia atrocyana Selys, 1865, Ischnura senegalensis (Rambur, 1842), Pseudagrion lalakense Orr & van Tol, 2001, Megalogomphus borneensis (Laidlaw, 1914), Agrionoptera insignis (Rambur, 1842) and Hydrobasileus croceus (Brauer, 1867)) are first records for the division, bringing the total number of Odonata known from the division to 160. Notable records include Coeliccia kenyah Dow, 2010, Teinobasis laidlawi Kimmins, 1936, Burmagomphus insularis Laidlaw, 1914, Leptogomphus sp. cf coomansi Laidlaw, 1936 and Macromia callisto Laidlaw, 1922. Remarks are made on the habitat preferences of Agrionoptera insignis and Camacinia gigantea (Brauer, 1867). A highly atypical population of Neurothemis Brauer, 1867 (species unclear at present) is reported.
168:43-55
Boudot (2022) criticised in unusually harsh language with reference to 'Resolution 74 (26) on the Right of Reply of the Committee of Ministers of the Council of Europe (https://rm.coe.int/CoERMPublicCommonSearchServices/DisplayDCTMContent?documentId=09000016805048e1)' in an e-mail with attached word document dated 31st January 2022, local time Zerf: 15.45h a small and rather insignificant publication written by Nataly Snegovaya and myself, and demanded that his counter statement be published. In my function as the responsible editor of IDF Report I granted this wish. I t has to be noted that all we had done was insisting on retaining the record of Bartenef (1912a) which we are considering a proven record of Gomphus vulgatissimus....
168
1-23: Odonata collected in 2021 in Azerbaijan, including new data on Gomphus schneiderii Selys, 1850 and Libellula pontica Selys, 1887 (Nataly Yu. Snegovaya); published: 21.01.2022 -
25-32: On the occurrence of Gomphus vulgatissimus (Linnaeus, 1758) and G. schneiderii Selys, 1850 in Azerbaijan – a brief discussion of the known status quo (Martin Schorr & Nataly Yu. Snegovaya); published: 24.01.2022 -
33-36: Reply to Schorr & Snegovaya (2022), this volume of IDF-Report (Jean-Pierre Boudot); published: 03.02.2022 -
37-42: Editor's response to Boudot (2022) (Martin Schorr); published: 16.02.2022 -
43-55: Call it schneiderii, but document which identification key you used (Martin Schorr); published: 17.02.2022 (Hinweis: dies ist eine geänderte Version des IDF-Report 168 mit Stand 17.02.2022, welche die früheren Versionen ersetzt).
168:1-23
This paper presents the results of a study on the odonate fauna in Azerbaijan The survey was conducted in the summer of 2021 and covered 24 localities in twelve districts. A total of 34 species from 9 families was recorded. New localities for Gomphus schneiderii Selys, 1850 and Libellula pontica Selys, 1887 are reported. A formerly published record of Gomphus vulgatissimus from Khachmaz, Nabran village, has to be corrected into G. schneiderii.
168:25-32
Boudot et al. (2021) extirpated, without any discussion, G. vulgatissimus from the list of Azerbaijan Odonata, subsuming all records of this taxon under G. schneiderii. This is contradictory to the fact that G. vulgatissimus was documented for Azerbaijan by Bartenef (1912). We discuss the current knowledge of the two taxa G. vulgatissimus and G. schneiderii, document a new record of G. vulgatissimus for Azerbaijan, and map all known findings of the two taxa that have been reported to date as well as the potential distribution (search area) of G. vulgatissimus in northern Azerbaijan.
167
This paper offers an explanation of each of the 44 scientific names given by Leopold Krüger (1861-1942) to odonate taxa together with that for the names of all the genera into which they are sorted now. But prior to that there is some information about the life and work of this scientist, and in the final part his preferences in odonatological nomenclature are compared with those in the names created by F.M. Brauer and F. Ris and some impressions of his studies on Neuroptera are presented and considerations about his aspirations in his work are given.
166
Inhalt:
Preparation of preserved field collected dried damselfly specimens for
character illustrations S. 1-7
Imaging preserved damselflies for scientific publications S. 9-20
166:1-7
Preparation of preserved field collected dried damselfly specimens for character illustrations
(2021)
165
Two new congeneric species endemic to Fiji are introduced: Nesobasis martina sp. nov. [holotype female, Viti Levu Is] and N. monika sp. nov. [holotype male, Taveuni Is]. The species are placed in the comosa- and erythrops-groups respectively (grouping following Donnelly 1990). Diagnostic features are proposed, but not discussed. Further discussion is left for an ongoing revision of the genus (Donnelly & Marinov in prep. ). The new species reported here are introduced ahead of this revision in a study which was made possible due to the International Dragonfly Fund (IDF) offering the opportunity to pick a new species' name for a donation.
164
Damselflies recorded before the administrative partition of the Indian Subcontinent and now housed at National Insect Museum (NIM), Islamabad were reviewed and catalogued. This collection is the divided part of National PUSA Collection (NPC) transferred to the Pakistan during 1947. Data for this collection had never been available or published. A record of 104 taxa is reported herein. Few of the species were found double named, misidentified and not updated as per valid classification. Some of the specimens were found unidentified. All such issues were resolved by following regional literature.
163
Im Jahr 2019 wurde ein etwa 500 m langer Abschnitt der Trattnach in Schlüßlberg (Bundesland Oberösterreich) restrukturiert. Ziel der libellenkundlichen Untersuchungen in den Jahren 2016 und 2021 war die Bewertung dieser ökologischen Aufwertung durch ein Prae- und ein Post-Monitoring. Die Verbesserungsmaßnahmen führten zu einer Verdopplung der Artengesamtzahl (von fünf auf zehn) und der Zahl der sicher, wahrscheinlich und möglicherweise bodenständigen Arten (von vier auf acht). Es konnten im Jahr 2021 vier der fünf gewässertyp-spezifischen Leitarten nachgewiesen werden: Calopteryx splendens, Calopteryx virgo, Gomphus vulgatissimus und Onychogomphus forcipatus. Das bodenständige Vorkommen dieser Arten bestätigt den hyporhithral/epipotamalen Übergangscharakter der Trattnach in diesem Bereich und indiziert die Ausprägung von geeigneten Strömungs- und Substratverhältnissen sowie von entsprechenden Vegetationsverhältnissen im Uferbereich. Durch die wasserbaulichen Eingriffe wurden insbesondere die rheophilen Spezies gefördert, limnophile Begleitarten waren nur vereinzelt nachzuweisen. Grund dafür ist der weiterhin in überwiegendem Maß gestreckte Verlauf des Flusses und das damit verbundene weitgehende Fehlen lenitischer Bereiche.
Die Funde von Exuvien und Sichtungen frisch emergierter Individuen von G. vulgatissimus und O. forcipatus belegen, dass das Gewässer unmittelbar nach Fertigstellung der Bauarbeiten von diesen Arten mit zweijähriger Entwicklungszeit besiedelt wurde. Die Bewertung des libellen-ökologischen Zustandes basierte auf der Berechnung des Odonata-Fließgewässer-Zonations-Index, in dem allfällige Abweichungen des aktuellen Odonatafauna von einem gewässertyp-spezifischen Referenzzustand (Klasse 1, „sehr guter libellen-ökologischer Zustand“) verrechnet werden. Das Ergebnis ist die Grundlage für die Einstufung in eine der fünf Klassen des ökologischen Zustandes gemäß EU Wasserrahmenrichtlinie. Die Restrukturierungen führten zu einer Verbesserung der libellen-ökologischen Zustandes: der betroffene Abschnitt (vormals Klasse 3, „mäßig“) wurde genauso wie die drei darin liegenden Einzelstrecken (vormals Klassen 4, „unbefriedigend“, und 3, „mäßig“) mit „gut“ (Klasse 2) bewertet. Auch der libellen-ökologische Zustand einer von den Aufwertungsmaßnahmen nicht betroffenen regulierten und in beiden Jahren kartierten oberstromig liegenden Kontrollstrecke veränderte sich von „unbefriedigend“ auf „mäßig“. Aus dem Restrukturierungsbereich wirkende Strahleffekte dürften für diese Verbesserung verantwortlich sein.
160:1-20
160:21-74
154:99-184
Phnom Kulen is a small and low plateau in the northern Cambodia still partly covered with evergreen forests and isolated from similar habitats by the Cambodian Lowland at least for 60 km. A preliminary checklist of its Odonata is provided, including 97 species. Burmagomphus sp. cf. willamsoni and Macromia callisto are for the first time reported for Cambodia.
154:29-98
Lake Tonle Sap in NW Cambodia is the largest freshwater lake in Southeast Asia and one of the most productive freshwater ecosystems in the world, so its banks are a home for ca 1,5 million people. It serves as a natural reservoir of the excess water of the Mekong River and cyclically changes its area from 2,500 km2 in May to 16,000 km2 in October. Its banks are naturally occupied by temporarily inundated forest and scrub, at present mostly replaced by rice fields. The present day semiaquatic vegetation of the lake is to a large extent formed by invasive plant species. The hitherto existing data on Odonata of the lake are very scarce. The author briefly examined the bank and floodplain at the NW part of the lake in February/March 2017, June and November 2018 and December 2019. Five main localities studied are described and illustrated
in detail. In total 41 odonate species of four families (22 in Libellulidae) were found. Most of them are common and widespread lentic species but Macrogomphus phalantus is a species hitherto known only by few specimens from swamped forests of Borneo and Sumatra; its Tonle Sap population was earlier described by the author as the subspecies M. phalantus jayavarman Kosterin, 2019. The earlier published report by Seehausen et al. (2016) of Sinictinogomphus clavatus (not found by the author) was a considerable extension of the known species’ range to the south. Six species were found in all main examined localities and ten only in one of them. At any season at the lake immediate bank (that is water front at the lowest level), Brachythemis contaminata predominates overwhelmingly, Orthetrum sabina and Crocothemis servilia are numerous, two damselfly species, Pseudagrion microcephalum and P. rubriceps, invariably occur at floating vegetation (mostly water hyacinth), and Trithemis pallidinervis, Urothemis signata, Rhyothemis phyllis, R. variegata and Tholymis tillarga are common at bushes. Agriocnemis nana, Ceriagrion praetermissum, Ischnura senegalensis, Macrogomphus phalantus, and Aethriamanta aethra were occasionally met at the lake bank. Other 26 species were found, with different occurrence and quantity, on the lake floodplain. Variation of the male occiput coloration of Amphiallagma parvum is commented.
154:1-28
Prey Long (Prey Lang) Forest is the largest remaining lowland rainforest in Indochina, shared by Stung Treng, Preah Vihear, Kampong Thom and Kratie Provinces of Cambodia, which has been persisted until present because of the lack of roads. It includes patches of unique evergreen swamp forests. Odonata of Prey Long forest, including Cheum Takong forest swamp, was briefly examined in December 2019, while the already deforested area was examined in June 2018. The former examination resulted in 40 species, the latter in 34 species, 60 species in total. Two species, Copera chantaburii Asahina, 1984 and Burmagomphus williamsoni Förster, 1914, are for the first time reported for Cambodia from the deforested area (Chey Saen District of Preah Vihear Province). The swamped forest of Cheum Takong provided 17 species, 5 of which were not found elsewhere in the considered area, 3 are rare and 4 generally Sondaic. Prey Long Forest should be re-examined in the rainy season soon after the road to Spong village is constructed.
153
The changes in coloration with age in males and females of Coeliccia mattii Phan & Kompier, 2016 from the Central Highlands of Vietnam are documented. Immature forms have extensive yellow markings on the thorax and S9 is yellowish, while in the fully mature male form, the thorax is largely black and covered in pruinosity and S9 is completely black. S9 is yellowish in the immature female while S9 is black in the fully mature female.
152
151
The paper presents new odonatological data from the formerly unstudied islands Patnanungan and Jomalig as well from the northeastern part of Polillo, The Philippines. Four species are new for the PolilloIsland group, including one new taxon Anax parthenope julius – for The Philippines. Currently, 78 odonate taxa are known from the Polillogroup.
150:3-20
A tenday collecting trip to Napo province was conducted between January 13 and 23, 2020, visiting localities where W. C. Macintyre originally collected Argia schneideri Garrison & von Ellenrieder, 2017 between 1935 and 1942, with the intention of documenting its life habits and obtaining photographs in life. A total of 65 odonate species in 36 genera were collected, including four new records for Napo province, but the target species was not found. A list of species recorded and color scans of species that have so far not been photographed are included. Color photographs and notes on the habitat of Argia schneideri are made available through the courtesy of colleagues who found it elsewhere, and its current known distribution is discussed.
149
148
An explanation is presented for each of the 135 scientific names given to Odonata by F. M. Brauer (fossils and synonyms included), in addition the names of the actual genera in which Brauer’s species are now classified are explained. Prior to that part biographical information is given and Brauer’s merits in enlightening the taxonomy of dragonflies are analysed. Conclusions are drawn as to his preferences in odonatological nomenclature and finally the difficulties are discussed, which Brauer had to face in his taxonomic work.
147
This paper presents the faunistic results of three short field excursions conducted in spring and early summer of 2018 in Kosovo, considering dragonfly fauna one of the most understudied countries of Europe. This study presents first systematic dragonfly research in Kosovo. Within a total of 13 field days between end of April and end of June 2018, 60 sites were surveyed and 44 dragonfly species were found. Significant results include the first documented report of 15 species for Kosovo. New data on several other species with a broader European concern or generally rare on the West Balkan peninsula, i.e. Coenagrion ornatum, Anax ephippiger, Caliaeschna microstigma, Cordulegaster heros, C. bidentata, Somatochlora flavomaculata, and Sympetrum flaveolum, are also presented. The overview of all visited sites is included. Altogether, 47 dragonfly species are now reported for Kosovo.
146
Previously unpublished records of Odonata from the states of Negeri Sembilan and Selangor in Peninsular Malaysia are presented. One hundred and eight species are listed, of which 77 were collected in Negeri Sembilan and 87 in Selangor. Fifteen of the species recorded from Negeri Sembilan and seven of those recorded from Selangor appear to be first records for the respective state. Notable records include Drepanosticta sp. cf hamadryas Laidlaw, 1931, Rhinocypha pelops Laidlaw, 1936, Acrogomphus ?malayanus Laidlaw, 1925, Heliogomphus kelantanensis (Laidlaw, 1902); Onychogomphus duaricus Fraser, 1924, Macromia cupricincta Fraser, 1924, Idionyx montana Karsch, 1891, Chalybeothemis chini Dow, Choong & Orr, 2007 and Hylaeothemis clementia
Ris, 1909 Ris, 1909. Provisional checklists of the Odonata known from Selangor plus the
Federal Territory of Kuala Lumpur (171 species) and for Negeri Sembilan (116 species)
are given in appendices.
145
Arising from a number of 2019 IUCN Red List assessments for a subset of Chinese Odonata, a series of corrections and taxonomic revisions were made to the World Odonata List. The rationale for these amendments is provided here. Paragomphus wuzhishanensis Liu, 1988 is shown to be a junior synonym of Paragomphus pardalinus Needham (1942). Epophthalmia kuani Jiang 1998 is synonymised as a junior synonym of Epophthalmia. elegans (Brauer, 1865) and Epophthalmia bannaensis Zha & Jiang, 2010 is treated as a junior synonym of Epophthalmia vittata Burmeister, 1839. Idionyx pseudovictor Xu, 2013 is shown to be junior synonym of Idionyx claudia Ris, 1912 and Sympetrum anomalum Needham, 1930 is treated as a junior synonym of Sympetrum maculatum Oguma, 1922.
144
We report here the results from field trips to collect Odonata in the northeastern parts of Kelantan state and the north of Terengganu state, Peninsular Malaysia. Eighty four species were collected, and four of these are new records for the state Kelantan and 10 are new records for the state of Terengganu. Notable records obtained from the field trips were Euphaea masoni, Coeliccia sameerae, Pseudagrion ?lalakense, Leptogomphus tioman and Macromia cupricincta. Checklists for Kelantan (140 species) and Terengganu (132 species) are given in an appendix.
143
Using a qualitative sampling approach to survey for Odonata along water systems inside and outside of four major protected areas in Southern Ghana (viz; Atewa Range Forest Reserve, Ankasa Forest Reserve, Owabi Wildlife Sanctuary and Bobiri Forest Reserve), a total of 66 Odonata species in seven families encompassing 28 Zygoptera and 38 Anisoptera were recorded. Libellulidae (n= 36) exhibited the highest number of species, followed by Coenagrionidae (n= 10), and Calopterygidae (n= 5). In considering the observed number of species exhibited by each protected area, the Atewa Range Forest Reserve exhibited the highest number of species (n= 51), followed by the Ankasa Forest Reserve (n= 47), the Owabi Wildlife Sanctuary (n= 44) and Bobiri Forest Reserve (n= 43). Disturbance tolerance species dominated in habitats outside the various forest reserves, whiles forest specialists predominated inside the reserves, indicating the quality of the forest cover therein.
142
141
Records of Odonata collected in Gunong Mulu National Park in Sarawak are presented. Between 2005 and 2019, in 12 surveys that lasted between one week and five months, 163 species were collected. The collections from Gunong Mulu National Park are of importance for the taxonomic study of dragonflies and damselflies in Borneo; several species have been described based on material collected in the Park.
140
Neu angelegte Kleingewässer werden oft durch Libellen rasch als Lebensräume in Anspruch genommen. Allerdings ist über die dauerhafte Bildung von Populationen, vor allem durch seltene und gefährdete Arten, noch wenig bekannt. Ziel dieser Studie war die Untersuchung des Besiedlungsprozesses von künstlich angelegten Gewässern durch Libellen mit besonderem Fokus auf moortypische Arten. In den Jahren 2015–2018 wurden in der Niederschlesischen Heide an sechs solchen Gewässern, drei neuen und drei alten, Exuvien von Großlibellen (Anisoptera) quantitativ aufgesammelt. Die Erfassung des Besiedlungserfolges durch Kleinlibellen (Zygoptera) erfolgte qualitativ als Zusatzinformation zur Artenvielfalt der Gewässer. Insgesamt waren 26 Libellenarten – 9 Zygoptera und 17 Anisoptera – bodenständig. Allerdings variierte der Artenreichtum deutlich sowohl zwischen den Gewässern als auch den Untersuchungsjahren, so dass sich keine klar definierten Unterschiede in der Artenvielfalt zwischen den neuen und alten Gewässern erkennen ließen. Unter der bodenständigen Arten wurden vier moortypische Großlibellen (Aeshna juncea, Leucorrhinia dubia, L. albifrons, L. pectoralis) nachgewiesen. Ununterbrochene Entwicklung ließ sich nur für A. juncea und L. dubia in einzelnen Gewässern bestätigen. Leucorrhinia albifrons kam nur sporadisch vor und wurde im letzten Untersuchungsjahr (2018) nicht mehr gefunden. Dagegen vollendete L. pectoralis ihre Entwicklung erst in den Jahren 2017–2018. Generell bevorzugten diese Arten Gewässer, in denen die Vegetation mäßig bis gut entwickelt war. Die Ergebnisse bestätigen, dass künstliche Gewässer durch ein breites Spektrum von Arten erfolgreich besiedelt werden können. Allerdings ließ sich die dauerhafte Reproduktion in den jeweiligen Gewässern nur für wenige Großlibellen nachweisen. Deshalb sind die Gewässer in der Niederschlesichen Heide nicht als langfristige Ersatzbiotope, sondern vielmehr als temporäre Fortpflanzungsbiotope von Individuen der Nebenpopulationen zu sehen.
139
138
Description of Hemicordulia tuiwawai sp. nov. from Kadavu Island, Fiji (Odonata: Corduliidae)
(2019)
Hemicordulia tuiwawai sp. nov. (Odonata: Corduliidae) is described and diagnosed based on material collected from Kadavu Island, Fiji; holotype: Wainitayuki River about 750 m above Baidamudamu village, -19.0916, 178.1038; 37 m a.s.l., 06 June 2016, M. Marinov leg. This species is distinguished from its congeners in the field by the contrasting colouration – dark green metallic body with bright yellow spots on the synthorax and base of the abdomen. This pattern is comparable to H. pacifica Selys, 1871. However, Hemicordulia tuiwawai sp. nov. can be recognised by the larger size and unique shape of the caudal appendages and genital hamule (in males) and vulvar scale (in females).
137
Records of Odonata from the southwest of Sri Aman Division and the extreme east of Serian Division in Sarawak are presented. The sampled areas are interesting not only because they are poorly known for Odonata but also because many are just to the south and west of the Lupar Line which is a division between the ancient Sunda shelf and more recent geological formations. Differences between the odonate faunas on either side of the Lupar Line are discussed. Eightyfive species of Odonata were recorded during the surveys reported on. The single most notable record is that of Coeliccia southwelli Dow & Reels, 2011, which represents a considerable extension to the known range of this species. Other interesting records include Telosticta dupophila (Lieftinck, 1933), T. species cf longigaster Dow & Orr, 2012, Podolestes parvus Dow & Ngiam, 2019 and Heliogomphus species cf olivaceous Lieftinck, 1961. Variation in the markings of Stenagrion dubium (Laidlaw, 1912) across its range is discussed and a gene tree using the COI marker is presented to illustrate the high variability of this species in this marker. However the variability in COI does not appear to be correlated with other characters.
136
Odonatological results of a trip across the southern Far East of Russia, from nearly the border of North Korea to Khabarovsk from July 1st to 16th, 2014, are presented. In total, 50 species were recorded, including those with limited presence in Russia, such as Paracercion calamorum, P. hieroglyphicum, P. plagiosum (for this species the 3rd Russian locality is reported), Pseudocopera tokyoensis, Stylurus annulatus (2nd Russian locality), Sinictinogomphus clavatus; Trigomphus citimus, Macromia daimoji (3rd Russian locality), M. manchurica, Deielia phaon, Lyriothemis pachygastra (2nd Russian locality). For S. annulatus, M. manchurica and D. phaon the northernmost known localities in the world are reported. D. phaon, earlier reported from one locality in Russia, have been found in five localities in Primorye and for the first time reported for Khabarovskiy Kray in general and Bol'shekhekhtsirskiy State Nature Reserve in particular. A trend of mutual exclusion of two abundant Shaogomphus postocularis epophthalmus and S. schmidti is supposed: the lowermost Ussuri River / Amur River, respectively. Variation in Paracercion spp. and Macromia amphigena fraenata is discussed.
135
During four field trips in Georgia, Armenia and Azerbaijan from 2010 to 2018, the author collected data of a total of 55 species (see Tab. 2 in Appendix). This study provides first insights into new or rare species in this ecoregion. Ischnura fountaineae and Cordulia aenea were found for the first time in Armenia. We also highlight the rediscovery of some species that were mentioned in the older literature but had not been confirmed since. An autochthonous population of Lestes macrostigma was discovered in Azerbaijan sixteen years after the single previous record by Dumont (2004). Original information is provided on the distribution of some rare species encountered in these countries. Finally, the identification of a puzzling Cordulegaster sp. observed in south Armenia is discussed briefly.
134
The Odonata collection deposited at the Museum of Comparative Zoology (MCZ) includes specimens of 634 taxa labeled as types. Fifteen of these have been incorrectly labeled as types (pseudotypes) and eight are apparently lost, leaving a total of 611 types currently deposited at MCZ. From these, 489 represent primary namebearing types (syntype/s, holotype, lectotype and neotype), 21 are probable primary types, and 101 are secondary types (paratype/s, paralectotype/s).
133
A checklist of 49 damselfly species from 12 families (Odonata: Zygoptera) recorded from Kon Ka Kinh National Park is provided. A first description of the female Protosticta socculus Phan & Kompier, 2016, is given. Burmargiolestes cf. laidlawi Lieftinck, 1960 and three apparently new species, two Coeliccia and one Protosticta species, are recorded, all of which are to be described in the future.
132
Phnom Tumpor is a scarcely accessible basalt table mountain in the Cardamom Mts. in Pursat Province of Cambodia. On top surface it bears tall evergreen forest (ca 1100 m a.s.l.), concealing a slow rivulet, O'Gran, being a chain of deep pools. It was examined odonatologically on March 1418th, 2019. Six common species were recorded in dry and burnt scrub on the Phnom Tumpor slopes and ten on the forested upper surface at O'Gran, among them Polycanthagyna erythromelas (Selys, 1891) and Macromia sp. cf. pinratani Asahina, 1987 for the first time in Cambodia. The peculiarities of the males of Coeliccia kazukoae Asahina, 1984 from Phnom Tumpor and the problem of distinguishing females of M. pinratani and M. moorei are discussed.
131
Survey in three protected areas of Chhattisgarh reveals the presence of 50 species of Odonata belonging to 34 genera and 9 families. Specimens were sampled from different lotic and lentic ecosystems. 17 species were exclusively found in or around running water. Larvae or exuviae of 23 species were found and photographed. Habitat availability, larval abundance, species composition and phenology are discussed.
130
In May 2018, 34 dragonfly species were recorded on Wolin (21), Usedom (30) and the adjacent mainland (21). The most frequent spring species in the area were Coenagrion puella, Libellula quadrimaculata, C. pulchellum, Ischnura elegans, Erythromma najas, Brachytron pratense and Cordulia aenea. C. lunulatum and Calopteryx virgo were recorded for the first time on Usedom, while Anax parthenope, Libellula fulva and Leucorrhinia caudalis were recorded for the first time on Wolin. The occurrence of Coenagrion armatum, which had been recorded in 2016 on Usedom, could not be confirmed.
The quantitative results of the survey are given for different types of water bodies (ditches, pools, ponds, lakes, bog lakes). The average number of species per water body was highest in bog lakes (7.5) and lowest (3.4) in temporary pools. In comparison to older studies from the area, the flight season started 13.7 days earlier in 2018 than in years documented prior to 1989. This might be explained by climatic changes. The species composition, however, has remained rather stable – at least on Usedom and the mainland. On Wolin, climatic factors as well as habitat loss and transformation have led to a species composition that is different from the one recorded in the first half of the 1970s.
129
New data on Odonata of the Preah Sihanouk Province of Cambodia obtained in March 2017 and November 2018 are presented. The presence of Onychargia atrocyana Selys, 1865 in Cambodia is confirmed. Twentyone species are added to the known fauna of the Kbal Chhay Waterfall environs, 19 species to that of Ream Peninsula and 4 to that of Koh Rong Island. The total number of species registered for Kampong Saom Peninsula amounts to 74. The presented data are rather of historical importance since most of the remaining forest has been quickly and irreversibly logged a few months ago. The validity of Gynacantha demeter Ris, 1911 as a species distinct from G. dohrnii Krüger, 1899 is doubted.
128
Globally urban wetlands are under high anthropogenic pressure of degradation. Urban wetlands are hotspots for species losses and rapid turnover in species assemblages. Therefore, studying such wetlands may provide an estimate of the pace of local extinction, concerning wetlanddependent species such as odonates. We undertook a study to document odonate species across a tropical urbanization gradient. We sampled six localities across the gradient across the Mula River that flows through the Pune City, India. We sampled adult odonates using a newly devised Halfcircle Point Count method from September 2016 to March 2017. We took multiple temporal replicates per site. We also sampled larvae across six sites once in November 2016. We measured site characteristics such as canopy cover, solid waste, and water turbidity to understand the level of disturbance at each site. We recorded 41 odonates, six species (primarily Gomphidae members) exclusively from the larval sampling. We did not find the localization of species in a particular site across the urbanization gradient, possibly because we sampled a relatively short stretch of the river to capture the variation. Here, we update the Odonata list of Pune including data on larvae. We demonstrate that larval sampling complements Odonata surveys, especially in recording Gomphids.
We recommend future research to include a longer timespan and extensive sampling area.
127
The article presents new faunistic data on 33 Odonata species, based on the material collected by the author in 2012, 2016 and 2017 and a systematic research in 2018 throughout the Nakhichevan Autonomous Republic (AR). Onychogomphus assimilis (Schneider, 1845) is a new record for the fauna of Azerbaijan. Eight species were registered for the first time for the territory of Nakhichevan AR: Lestes virens Rambur, 1842, Coenagrion scitulum (Rambur, 1842), Aeshna mixta Latreille, 1805, Anaciaeschna isoceles (Müller, 1764), Anax parthenope (Selys, 1839), Sympetrum sanguineum (Müller, 1764), Crocothemis erythraea (Brullé, 1832), and Selysiothemis nigra (Vander Linden, 1825).
126
We present records of 99 Odonata species from the Réserve Naturelle des Gorilles de Lésio-Louna (RNGLL) in the Republic of Congo (Congo-Brazzaville) collected during a short survey from 14 January to 01 February 2017. It is the first systematic Odonata inventory for the RNGLL and for the Congolese part of the species-rich Batéké Plateau. A short introduction is given about the existing knowledge of dragonflies and damselflies from the country. Amongst the recorded species eight are new for the country list, raising it to at least 208 species. Some of the recorded species are endemic and characteristic for sandy streams and rivers of the Batéké Plateau. The potential diversity of the plateau in comparison to other regions of the country is discussed.
125
A total of 20 Odonata species were recorded from 15 water bodies (3 natural water bodies and 12 manmade reservoirs) in a desert landscape west of the Namak Lake in the northwest of the Central Plateau of Iran, from 27 May to 20 June, 2017. The study area included Qom province and the northern part of Esfahan province (34.07 to 35.14 N, 51.33 to 50.89 E). For Qom province, 17 out of 18 species found are new provincial records. In addition Selysiothemis nigra, Orthetrum chrysostigma and Orthetrum taeniolatum are new findings for Esfahan province. The species around the manmade reservoirs are characterized by a broad ecological amplitude ("generalists") while species assemblages of natural water bodies consisted of more sensitive species. In the arid climate of central part of Iran, manmade reservoirs are major habitats for Odonata species. Although these reservoirs have increased the amount of habitat for generalist species, they do not foster sensitive and specialist species. Furthermore, considering the more intense droughts predicted for these areas in the future, and the current scenario of environmental degradation, some sensitive species may be in danger of local extinction.
124
A record of male and female specimens of Sinolestes editus Needham, 1930 collected in Dak Hro village (14°19'45'' N, 108°24'23'' E, alt. 1,420m a.s.l.), Dak Roong Commune, K’Bang District, Gia Lai Province, Central Highlands of Vietnam is present with a detailed description of the morphology of these two specimens.
123
More Odonata found at the Cardamonean foothills in Koh Kong Province of Cambodia in 2014-2018
(2018)
Still unpublished data on Odonata of the coastal foothills of the Cardamom Mts. in Koh Kong Province, SW Cambodia obtained in 2014-2018 are provided. Tetrathemis flavescens Kirby, 1889 is for the first time reported for Cambodia and Risiophlebia guentheri Kosterin, 2015 for SW Cambodia. The photographic records from this area of Gynacantha basiguttata Selys, 1882, Gynacantha demeter Ris, 1911, Heliaeschna crassa Krüger, 1899, Amphithemis curvistyla Selys, 1891 and Pornothemis serrata Krüger, 1902 have been confirmed by specimens. The known local faunas of two neighbouring rivulets in Koh Kong Province, rich in Odonata, are updated and summarised.
122
Records of Odonata from Gunung Melatai, Nanga Gaat and the Kastima Logging area, all in Sarawak’s Kapit Division, are presented. The most notable records are of Matronoides cyaneipennis Förster, 1897 and Heliogomphus blandulus Lieftinck, 1929. A distribution map for Matronoides cyaneipennis and updated distribution maps for three species from the Coeliccia borneensisgroup are given. Tentative identifications to species of previously published records of Idionyx females are given. The genus Heliogomphus in Borneo, Peninsular Malaysia and Singapore is reviewed and a simple one marker molecular analysis is presented for the genus in this region. Based on reexamination of specimens from the genus and the molecular results, an additional member of the genus is reported from Borneo: H. sp. cf olivaceus Lieftinck, 1961. Although both morphological and molecular results remain incomplete, it does appear likely that there is at least one more species of Heliogomphus present in Borneo than has been recognised until now, and that H. borneensis Lieftinck, 1964 may be a junior synonym of H. kelantanensis (Laidlaw, 1902).
121
Macromidia genialis buusraaensis subspecies nova (Odonata, Synthemisidae s.l.) from eastern Cambodia
(2018)
Macromidia genialis buusraaensis subspecies nova is described from 10 males and 2 females from three localities of Mondulkiri Province, the eastern Cambodia (the type locality: a brook downstream Buu Sraa Waterfall 12°34' N 107°25' E). The new subspecies is close to M. g. shanensis Fraser, 1927 and differs from it by an additional, and the broadest, yellow spot on S7. It is partly syntopic with Macromidia rapida Martin, 1907. A brief overview of the current knowledge of the genus Macromidia Martin, 1907 is provided.
120
Results of a brief examination of the spring aspect of Odonata at the Abrau Peninsula and some nearby localities on 2127th May 2018 are presented. Lestes dryas and Caliaeschna microstigma are reported for the Peninsula for the first time, revealing the world northernmost record of the latter. Some old data of Odonata at Novorossiysk and some recent erroneous data on the Krasnodarskiy Kray are referenced and discussed. The known Odonata fauna of the Abrau Peninsula reaches 38 species.
119
An odonatological survey, based on sighting and photographic documenting, was conducted in Palawan and in Cuyo Island. Ten species were found in Cuyo Island raising the number of known species from five to eleven. There are six additions to the Cuyo Island fauna: Agriocnemis f. femina, Ischnura senegalensis, Pseudagrion microcephalum, Brachydiplax c. chalybea, Neurothemis fluctuans and Neurothemis t. terminata. The occurrence of Coeliccia boettcheri, known from Cuyo, but not recorded during this survey, is discussed in some detail. Forty species were recorded in Palawan. Four new additions to the Palawan fauna were recorded: Lestes p. praemorsus, Pseudagrion microcephalum, Xiphiagrion cyanomelas, and Anax guttatus. Neurobasis daviesi, a rare calopterygid damselfly endemic in Palawan, was encountered at the northern side of Cleopatra Needle during the survey.
118
A catalogue of all types of subfamilies Protoneurinae and Disparoneurinae currently housed in the entomological collection of the Museum für Naturkunde − Leibniz Institute for Evolution and Biodiversity Science in Berlin (Germany) is presented. It includes current status of the familygroup, genusgroup and speciesgroup names, transcriptions of data labels and references to the original descriptions.
117
In the period 18th – 31st May 2017, 33 localities were examined for Odonata in the Central Zagros area of Iran: 16 in Markazi Province, 14 in Lorestan Province and 3 in Esfahan Province; in 27 of those localities Odonata were found, 25 species in total. For Markazi Province, only one species, Calopteryx splendens, was previously reported (and also found by us), so of 17 species found there 16 are formally new provincial records (Epallage fatime, Lestes barbarus, Coenagrion ornatum, C. persicum, Enallagma cyathigerum, Ischnura elegans, I. intermedia, I. pumilio, Platycnemis kervillei, Anax imperator, Anaciaeschna isoceles, Caliaeschna microstigma, Libellula depressa, Onychogomphus lefebvrii, Orthetrum brunneum, Sympetrum fonscolombii). Of 17 species
found in Lorestan, 5 are new for this province (L. barbarus, Aeshna mixta, Orthetrum taeniolatum, Sympetrum arenicolor, S. striolatum). Only two species were seen in Esfahan Province, in which little time was spent. Notes on variation and taxonomy are provided for Sympecma paedisca, C. ornatum (considered to be a senior synonym of C. vanbrinkae because of variation in the presumed main diagnostic character), E. cyathigerum, I. elegans, Gomphus schneideri (including discussion of G. amseli), as well as notes on habitats of most species and the general characteristics of the area.
115
Records of Odonata from the Lanjak Entimau Wildlife Sanctuary (LEWS) in Sarawak, Malaysian Borneo are presented. Previous records of Odonata from LEWS are critically examined. One hundred and ten species have been recorded within the sanctuary, including three that have yet to be found outside (Drepanosticta adenani, Telosticta iban and "Elattoneura" mauros); records of four more species are regarded as in need of confirmation, those of another six are incorrect. In addition to the three species only known from LEWS, other notable records include: Drepanosticta sbong, Dysphaea lugens, Euphaea sp. cf basalis, Pericnemis kiautarum, Burmagomphus insularis, Gomphidia caesarea, Merogomphus species, Phaenandrogomphus safei, Macromia callisto and Idionyx montana. A fresh illustration of the anal appendages of Drepanosticta sbong in lateral view is provided to make up for inaccuracies in the original illustration. The taxonomy of Phaenandrogomphus safeii is briefly discussed. Zygonyx errans Lieftinck, 1953 is considered a subspecies of Z. ida not Z. iris Additional records from areas adjacent to LEWS are given in an appendix.
114
New field sampling was carried out in June 2011 in northern Moldova, the most unstudied area in the Republic of Moldova. A total of 19 Odonata species was recorded during a field trip, with Libellula fulva and Gomphus flavipes new for Moldova. The presence of species of European concern is discussed. Photos of all visited habitats are included. The fauna of Moldova mostly consists of species which are able to survive in biotopes with high anthropogenic pressure.
113
Sympetrum fonscolombii is for the first time reported for Tuva (Tyva Republic, Russia), as found in the UbsuNur Depression. New data are provided on Odonata of the Turan or TuranUyuk Depression of Tuva, including the first record of Somatochlora exuberata in Tuva beyond the Todzha Depression and Coenagrion armatum, C. ecornutum, Aeshna juncea, A. grandis, Somatochlora graeseri and Libellula quadrimaculata for the first time reported for the Turan Depression. New distributional data and comments on Ophiogomphus spinicornis Selys, 1878 are added. Somatochlora alpestris found at Lake Oyskoe is for the first time reported for the southern Krasnoyarskiy Kray.
112
We describe three new species of damselflies from streams draining into Lake Kutubu in Southern Highlands Province, Papua New Guinea. They are Drepanosticta johncanni sp. nov. (Platystictidae), Pseudagrion parafarinicolle sp. nov. (Coenagrionidae) and Wahnesia kutubuensis sp. nov. (Argiolestidae). Diagnostic characters of the males and, where available of the females, are illustrated and the probable affinities of the new species are discussed.
111
Part 1, Quoc Toan Phan and Van Quang To, page 1-6:
The genus Megalestes Selys, 1862 in Vietnam, with first description of female of Megalestes australis Karube, 2014 (Odonata: Zygoptera: Synlestidae)
Abstract: The male secondary genitalia and the caudal appendages of males and females of the three confirmed Vietnamese Megalestes species (M. australis, M. haui and M. micans) are figured. The female sex of M. australis is described for the first time.
Part 2, Quoc Toan Phan and Van Quang To, page 7-19:
Odonata checklist from Son Tra Nature Reserve, Da Nang city, central Vietnam
Abstract: A checklist of 44 odonate species (21 Zygoptera and 23 Anisoptera) from Son Tra Nature Reserve, central Vietnam is provided. Idionyx thailandica Hämäläinen, 1985 is recorded for the Vietnamese fauna for the first time, a new Coeliccia sp. is awaiting description in the near future, and the taxonomic status of specimens of the genus Leptogomphus remains unsettled and will have to be solved in the future.
110
Results are presented of a brief odonatological examination of the Abrau and Taman' Peninsulas at the northwesternmost Caucasian Black Sea coast in Krasnodarskiy Kray, Russia, on July 2026, 2017. Twenty three species have been observed at the former peninsula and five at the latter. The Odonata records at the Abrau Peninsula are summarised, to include 34 species.
109
Gewässer anthropogenen Ursprungs haben potenziell einen hohen Wert als Lebensraum für die Libellenfauna. Im Jahr 2017 wurden alte Restgewässer im Gebiet einer Tongrube bei Gozdnica (Woiwodschaft Lubuskie, SW Polen) auf Libellen untersucht. Dabei stand im Vordergrund die Überprüfung der Gewässer hinsichtlich des potenziellen Vorkommens von Leucorrhinia caudalis (Charpentier, 1840). Weiterhin sollte untersucht werden, ob sie einen Beitrag für die regionale Libellenvielfalt leisten. Die durchgeführte Kartierung erbrachte zwei neue Fundorte von L. caudalis mit Nachweis eines bodenständigen Vorkommens an einem Gewässer und Beobachtung von geschlechtsreifen Imagines an beiden Fundorten. Bemerkenswert sind darüber hinaus bodenständige Nachweise von zwei weiteren nach nationalem und europäischen Recht geschützten Arten: Leucorrhinia pectoralis (Charpentier, 1825) und Leucorrhinia albifrons (Burmeister, 1839). Außerdem ist das gesamte Gebiet für Pionierarten, wie Ischnura pumilio (Charpentier, 1825) und Libellula depressa Linnaeus, 1758 sowie thermophile Arten, wie Orthetrum coerulescens (Fabricius, 1798) und Crocothemis erythraea (Brullé, 1832) als Lebensraum besonders geeignet. Insgesamt konnten 30 Libellenarten nachgewiesen werden. Für 16 Arten wurde der Reproduktionsnachweis erbracht. Die untersuchten Restgewässer stellen einen wichtigen Lebensraum sowohl für weit verbreitete Generalisten als auch für seltene Spezialisten dar. Damit tragen sie auch signifikant zur Erhaltung der regionalen Libellenvielfalt bei.
108
A survey of Odonata on the Indonesian island of Belitung is reported. The work of Belitung Biodiversity Observer on Odonata is briefly outlined. Sixty four species were recorded during the survey, including two new records for the island. A checklist of the known odonate fauna, consisting of 105 species, of the island is given in an appendix.
107
Part 1, Oleg E. Kosterin and Vladimir I. Solovyev, page 1-43:
Odonata found in mid-summer 2015 and 2016 at the north-westernmost Black Sea Coast of the Caucasus, with the first record of Cordulegaster picta Selys, 1854 in Russian Federation
Abstract: Results are presented of brief odonatological examination of the Black Sea coastal northwesternmost spurs of the Caucasus between Anapa and Gelendzhik (mostly at Kabardinka village), Russia, in late July/early August 2015 and early-mid July 2016. In total, 28 Odonata species were found, including Cordulegaster picta for the first time in Russia. For C. picta and Caliaeschna microstigma, the world’s northernmost records were made. New localities of species rare in this area are reported: one for Coenagrion scitulum and three for Selysiothemis nigra, including their breeding habitat. Numerous migrant individuals of Pantala flavescens were observed in many localities in 2015 but none in 2016. Observations on trophic activity of Aeshna affinis and A. mixta are reported, the former showing predominantly matutinal and vespertinal activity and the latter diurnal activity. Occurrence of the Chalcolestes spp. in the Caucasus is discussed.
Part 2, Oleg E. Kosterin, page 45 - 57:
Calopteryx virgo feminalis subsp. nov., a long known under the same name but hitherto formally nameless subspecies from the Caucasian Black Sea Coast
Abstract: The populations of Calopteryx virgo of the Black Sea Coast of the Caucasus have females with the distal hindwing part darkened and males with the underside of S10 and appendages whitish, They are known for more than a century and deserve a subspecific status but since the name feminalis Bartenev, 1910 proposed to them is unavailable, a new subspecies is formally erected under the same name, Calopteryx virgo feminalis Kosterin subsp. nov., with the following type locality: Russia, Krasnodarskiy Kray, Gelendzhik Municipality, Kabardinka village, the Doob River lowermost reaches, 44°38'26-53'', 37°55'55''-57'58'' E.
106
A new species of the genus Teinobasis Kirby is described from the Muller Range in Western Province, Papua New Guinea. Its male is distinguished from all other Teinobasis species by having a pale labrum, an extensively bright orange thorax, and ventrally bowed superior anal appendages that are markedly shorter than the plump, apically rounded inferiors. Characters of the male are illustrated, and the affinities of the new species are discussed.
105
Results are presented of an odonatological survey of 23 localities in Thala Barivat District of Stung Treng Province, northern Cambodia, on July 26 – August 1, 2016. Most localities were situated in areas of open low deciduous dipterocarp forests on gravel soils, some at hillside areas of tall evergreen dipterocarp forest. The great Mekong River right bank was studied within 7 km downstream of its Nimith (Nimet, Khon Thai, Labak Koun, Khone Pha Pheng) Waterfall. In total, 55 species were found, of which 52 identified to species and three to genus. Two species, Gynacantha saltatrix Martin, 1909 and Macrogomphus matsukii Asahina, 1986, were recorded in Cambodia for the first time. Five obligatory lotic species were found at the Mekong River, namely Dysphaea gloriosa, Prodasineura coerulescens, Burmagomphus asahinai, Nychogomphus duaricus and Onychothemis testacea, including tenerals of P. coerulescens and B. asahinai. Most probably these species breed in the Mekong reach which is enriched with oxygen downstream of the great waterfall cascade.
104
Drepanosticta kosterini sp. nov. (holotype ♂, from Gunung Penrissen, Kuching Division, Sarawak, Malaysian Borneo, deposited in RMNH) is described from both sexes. It is the sister species of D. actaeon Laidlaw, 1934; a fresh description of the male of D. actaeon and the first description of the female are given, along with discussion of variation in this species. Both D. actaeon and D. kosterini are considered to belong to a species group also including D. rufostigma (Selys, 1886) and a preliminary discussion of variation in this species is given, along with illustrations of both sexes. A neighbour joining COI gene tree for D. actaeon and D. kosterini is presented. The relationships of D. actaeon, D. kosterini and D. rufostigma to other members of the Platystictidae are briefly discussed.
103
Between 2009 and 2016, a total of 174 Odonata species (Tab. 1) have been recorded in the Dehong Dai and Jingpo Autonomous Prefecture, China. 21 of these species are new records for China, and additional 26 taxa have to be described as new to science. Brief comments on selected species refer to morphological characters, distribution and seasonality.
102
DragonflyIndia Meet 2016
(2017)
101
The Odonata fauna of flat marshy areas of the Gulf of Siam coast in Koh Kong Province of Cambodia, containing 55 species, is considered. The published data of 2010- 2013 and new data of 2014 and 2016 on the surroundings of Koh Kong town are compiled and the first data on the area of large swampy Melaleuca forests at Andoung Tuek village are presented as well as some occasional photographic records. Gynacantha bayadera, Lyriothemis mortoni and Pornothemis serrata were for the first time recorded for Cambodia. Mortonagrion falcatum was found unexpectedly abundant at Andoung Tuek.
100
In der Arbeit werden Ergebnisse der fortgeführten Libellenkartierung in den Jahren 2015 und 2016 in drei MoorNaturschutzgebieten ("Zacisze", "Przygiełkowe Moczary" und "Żurawie Bagno") in der Niederschlesischen Heide dargestellt. Insgesamt konnten 41 Arten nachgewiesen werden. Acht Arten wurden neu nachgewiesen, darunter sechs Arten mediterraner Herkunft. Die Beobachtung von Orthetrum albistylum (Selys, 1848) zeigt, dass diese Art bereits die westliche Grenze Polens erreicht hat und eine weitere Expansion nach Ostdeutschland zu erwarten ist. Die Untersuchungen erbrachten keine neuen Funde von Aeshna subarctica Walker, 1908 im Untersuchungsgebiet; somit bleibt der Status dieser Art immer noch unklar. Von den seltenen Arten wurde Leucorrhinia caudalis (Charpentier, 1840) im NSG „Zacisze“ belegt. Es ist die erste Feststellung dieser Art in der Niederschlesichen Heide und soweit die einzige im südlichen Teil der Lubuskie Woiwodschaft; jedoch bleibt der Status noch unbekannt. Die Studie bestätigt, dass die Biodiversität in den Schutzgebieten sehr hoch ist. Die thermophilen Arten tragen zur Erhöhung der lokalen Diversität in den Mooren bei, sind jedoch gleichzeitig regional neue Faunenelemente, die eine starke Veränderung der moorigen Libellengemeinschaft verursachen können. Zur Beobachtung dieser Entwicklung sind weitere systematische Erfassungen in den Gebieten notwendig.
99
Palaeosynthemis opaca sp. nov., a new dragonfly from Papua New Guinea (Anisoptera: Synthemistidae)
(2016)
A new species of the synthemistid genus Palaeosynthemis is described from the northern slopes of Papua New Guinea’s central cordillera. It is distinguished from all congeners by having females with large, nearly black patches at the bases of both wings. The male is most similar to P. cervula and P. feronia, but it differs from those species in having superior anal appendages less than three times as long as S10, basally enlarged and otherwise unarmed vs basally not enlarged (feronia) and almost four times as long as S10 and armed (cervula). Characters of the adults (both sexes) are illustrated and the affinities of the species are discussed.
98
Results of an odonatological survey of Mondulkiri Province of Cambodia, at the foothills and Central Plateau of the Annamese Mts. in June 8 – June 17, 2014 are presented. Of 106 Odonata species met (46 zygopterans, 60 anisopterans), 97 were identified to previously known named species, of which 15 are reported for the first time for Cambodia, namely Mnais mneme Ris, 1916, Rhinocypha seducta Hämäläinen et Karube, 2001, Philoganga loringae Fraser, 1927, Rhinagrion hainanense Wilson et Reels, 2001, Amphiallagma parvum (Selys, 1876), Ceriagrion chaoi Asahina, 1967, Paracercion malayanum (Selys, 1876), Prodasineura doisuthepensis Hoess, 2007, Protosticta grandis Asahina, 1985, Tetracanthagyna waterhousei McLachlan, 1898, Gomphidia kruegeri Martin, 1904, Heliogomphus chaoi Karube, 2004, Leptogomphus baolocensis Karube, 2001, Microgomphus jurzitzai Karube, 2000, Onychothemis culminicola Förster, 1904, and two species for which the specimens collected on this trip were described as new subspecies: Indolestes gracilis expressior Kosterin, 2015, Coeliccia poungyi dasha Kosterin, 2016. Five species collected on this trip have been described elsewhere as new to science, namely Onychargia priydak Kosterin, 2015, Prodasineura hoffmanni Kosterin, 2015, Asiagomphus reinhardti Kosterin et Yokoi, 2016, Euthygomphus schorri Kosterin, 2016 and Risiophlebia guentheri Kosterin, 2015. So the total number of the first country records of named species made on this trip is 22. Still four species found may be undescribed. The number of named species recorded in Cambodia has reached 178. Remarks on taxonomy and variation of Euphaea masoni Selys, 1879, E. ochracea Selys, 1859, Aciagrion approximans (Selys, 1876), and Lamelligomphus castor Lieftinck, 1941 are provided. Characters of mature males of R. seducta are updated. Notes on habitats and habits of some species are provided. Onychothemis culminicola and O. testacea Laidlaw, 1902 seem to exclude each other at rivers, the former occupying smaller and more elevated ones; a putative hybrid male was observed. General notes on the area and field impressions are briefly outlined.
97
Coeliccia poungyi dasha subsp. nov. (Odonata, Platycnemididae, Calicnemiinae) from eastern Cambodia
(2016)
Coeliccia poungyi dasha is described from the Annamense Mts. in the eastern Cambodia (holotype: Cambodia, Mondulkiri Province, a brook, a left tributary of the main river downstream Buu Sraa Waterfall, 12°34'0119'' N 107°24'50''25' 03'' E, 416490 m a.s., 15.06. 2014, RMNH). The new subspecies differs from the nominotypical one in coloration of the mesepisternum in males and end of the abdomen in both sexes, as well as in the length of the terminal lobe of the genital ligula.
96
In 2014, 56 localities in four provinces of Lesser Himalaya in Pakistan were studied. A total of 28 species have been recorded. A female of the data deficient, threatened species Coeliccia vacca was recorded from Charhaan. The record of Drepanosticta carmichaeli is a new addition to the list of Odonata of Pakistan, and expand the range of this species further to the west. The taxonomical status of Ischnura aurora aurora – considered common in Pakistan, following baseline literature of Fraser (1933) – now turns out to be Ischnura aurora rubilio.
95
As a result of increased interest in dragonflies and close cooperation between odonatologists on the Balkan Peninsula, the Balkan Odonatological Meeting (BOOM) has been established in 2011. This report presents the results of the field trip during 4th Balkan Odonatological Meeting that was held in Bosnia and Herzegovina from 1st to 8th August 2014. With 69 surveyed localities and 496 records of 47 species, this was the most successful BOOM until now. The noteworthy results are records of several nationally rare species: Chalcolestes viridis, Coenagrion scitulum, Erythromma najas and Ceriagrion tenellum, and new populations of Cordulegaster heros, a species mentioned in the Annexes of the EU Habitats Directive. The distribution of Chalcolestes spp. in Bosnia and Herzegovina is also presented and discussed.
94
Distribution data of dragonflies and damselflies from western Bhutan collected during a trip from 10 October 2015 to 22 October 2015 are presented. In total 53 species were recorded of which eleven are new to the country (Aciagrion pallidum, Anisopleura lestoides, Megalestes irma, Gynacantha incisura, Gynacantha khasiaca, Gynacanthaeschna sikkima, Lamelligomphus risi, Somatochlora daviesi, Crocothemis erythraea, Sympetrum fonscolombii, Tholymis tillarga). Another three species, one Megalestes and two Cephalaeschna, were not identified to species level but are also addition to the list of species recorded from Bhutan.
93
Specimens of Neurothemis disparilis Kirby, 1889, N. fluctuans (Fabricius, 1793), N. fulvia (Drury, 1773), N. ramburii (Brauer, 1866), N. stigmatizans (Fabricius, 1775) and N. terminata Ris, 1911, including their subspecies, were studied with the main focus on the morphology of the vesica spermalis, wing maculation, wing venation, abdominal markings and vulvar scales. The results were compared with species descriptions and directly with type specimens where possible. The vesica spermalis, especially the medial process, is useful at least in separating species groups and supports the traditional differentiation methods using wing maculation and venation. The use of other characters in accessing specific status, coupled with known distribution patterns, is discussed. The following taxonomic changes are proposed: Neurothemis manadensis (Boisduval, 1835) stat. nov., Neurothemis papuensis (Lieftinck, 1942) stat. nov. and Neurothemis taiwanensis sp. nov. is described (27.5.1998, Kenting, Pingtung County/Taiwan, L. M. Juang leg.; holotype is deposited at Taiwan Forestry Research Institute, Taipeh, Taiwan). The type of Polyneura palliata Rambur, 1842 was rediscovered at MNHN and designated as lectotype; a lectotype for Neurothemis nicobarica Brauer, 1867 housed at NHMW is designated. The holotype of Neurothemis incerta Brauer, 1867 was rediscovered and synonymized with N. ramburii.
92
A catalogue of 1290 persons commemorated in the scientific names of extant dragonflies (Odonata) is presented together with brief biographical information for each entry, typically the full name and year of birth and death (in case of a deceased person). For each individual a list is given of all available species, subspecies, genus or subgenus names erected in his or her honour. In total 2021 available names which qualify as eponyms are listed. These comprise 1966 species-group and 55 genus-group names including synonyms and homonyms. It is calculated that of the ca 8550 available species-group names in extant Odonata, ca 23 % are eponyms. Of the 1065 new species-group names introduced between 1 January 1995 and 31 December 2015, 435 (40.8 %) are eponyms.
91
New faunistic data is provided on the Odonata inhabiting the three main islands within the Samoan archipelago, namely Savai’i, Upolu and Tutuila as well as the smaller islands of Aunu’u and the Manu’a group. The specimens collected or observed in the field were compared to samples from other nearby Pacific island groups such as Fiji and Tonga. This study makes important contributions towards resolving taxonomic issues regarding the Ischnura species described as endemic to Samoa and their relations to other Coenagrionidae genera. New diagnostic features for distinguishing between females of the endemic genera Amorphostigma and Pacificagrion, subspecies separation in the Pacific Tramea transmarina and distinguishing between Samoan Hemicordulia species are suggested. Anaciaeschna melanostoma is proposed as junior synonym of A. jaspidea. A possible new subspecies of Lathrecista asiatica, confined to the Samoan archipelago, is discussed. The validity of Agriocnemis interrupta as a separate species from A. exsudans is questioned. Pacific Pseudagrion is believed to be represented within the region by one species only, with separate subspecies in Fiji, Tonga and Samoa, although more specimens from Fiji are required to resolve this issue.
90
Two new species of damselflies are described from central Halmahera in North Maluku Province, Indonesia. They are Drepanosticta pararudicula sp. nov. (Holotype MZB. ODON. 19257) and Nososticta halmahera sp. nov. (Holotype MZB. ODON. 19265). The two species are most similar to the Moluccan taxa D. rudicula and N. moluccensis respectively and their descriptions bring the total number of Drepanosticta species known from Halmahera to five and of Nososticta to two.
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The Cardamom foothills were re-assessed for Odonata in the late dry season of 2015 within E Thailand and SW Cambodia. In the narrow coastal strip of Trat Province of Thailand bordering to Cambodia, 44 species (1 unidentified) were recorded, of which 15, namely Agriocnemis nana, Archibasis viola, Ischnura senegalensis, Pseudagrion microcephalum, P. williamsoni, Acisoma panorpoides, Brachythemis contaminata, Brachydiplax farinosa, Hydrobasileus croceus, Macrodiplax cora, Rhyothemis plutonia, R. variegata, Tholymis tillarga and Trithemis pallidinervis were recorded for Trat Province for the first time. That increased the number of species recorded for the province to 61. Preliminary checklists of Odonata of Ream Peninsula (that is of Ream National Park) and of Koh Rong Island were complied, mostly on the data of this trip, to count 45 species (2 unidentified) and 17 species, respectively. As many as 36 species were recorded at the village of O’Som, Pursat Province. Copera marginipes is added to species recorded from Bokor Hill Station. Superficially similar males of Pseudagrion australasiae and P. microcephalum were observed in the same locality in Ream National Park.
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This article contains new faunistic data on 53 Odonata species based on material collected by the authors in Azerbaijan between 2013-2014 and added by revising an old collection made by A.V. Bogachev in the 1930-1940s. Of these, 13 species are new for the country: Lestes dryas, L. sponsa, L. virens, Coenagrion hastulatum, C. lunulatum, C. ornatum, C. pulchellum, Aeshna affinis, Brachytron pratense, Cordulegaster picta, Somatochlora flavomaculata, Sympetrum flaveolum, S. vulgatum. The new locality of Cordulegaster picta is the easternmost for the species. Two new populations of a very rare species Cordulegaster vanbrinkae (discovered in the country in 2011) are found. Somatochlora flavomaculata and Brachytron pratense are generally very rare in the entire Caucasus. Pantala flavescens has been rediscovered in the country 100 years after the first record. A large population of Caliaeschna microstigma, a species very rare in Azerbaijan, has been found in the northern part of the country; it represents a peculiar dark morph whose features are described in detail. Particulary discussed are the variability of Coenagrion puella-complex in Azerbaijan and diagnostic features of two other Coenagrion species closely related to each other, C. ornatum and C. vanbrinkae.
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Part 1, Oleg E. Kosterin, page 1-28:
On the Odonata of North Kazakhstan Province. I. First data on Petropavlovsk
Abstract: The fauna of Odonata of the environs of Petropavlovsk, North Kazakhstan, was for the first time examined on two short trips in late June and mid August 2015. Thirty five species were revealed. Coenagrion ecornutum was recorded in Kazakhstan for the first time, Gomphus vulgatissimus the second time and Stylurus flavipes the third time. Range expansion of C. ecornutum is discussed. Comparison is attempted of the known local Odonata faunas of the environs of Petropavlovsk, Omsk and Novosibirsk cities residing at the same latitude in the West Siberian Lowland. The Petropavlovsk fauna is very close to that of Omsk. The earlier published Kazakh records of G. vulgatissimus and S. flavipes are clarified and corrected. Breeding of Aeshna viridis in Ishim River (lacking water soldier) is supposed.
Part 2, Oleg E. Kosterin, page 29-46:
Odonata registered on a short excursion to Kyshtovka District, Novosibirsk Province, Russia
Abstract: During a fourday trip to Kyshotovka District, the most northwestern district of Novosibirsk Province, 21 species of Odonata were recorded. Two significant findings were made: that of Coenagrion ecornutum is most northern in West Siberia, and that of Lestes macrostigma is perhaps the northernmost in its range. The latter species was found over small, shallow, freshwater pools along a roadside. The diversity of this species’ habitats in Siberia in comparison to its uniform habitats at brackish water in Western Europe is discussed.
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Two new Cordulegaster species are described and illustrated by drawings, photographs and SEM images. The first one, C. plagionyx sp. nov., was discovered in NW Azerbaijan in a low-mountain forest landscape; the second, C. nachitschevanica sp. nov., occurs in the subalpine zone of Nakhichevan AR, south of the main territory of Azerbaijan. Both new taxa look generally similar to C. insignis Schneider, 1845; however, each of them reveals unique features and distinctive combinations of characters that set them apart from other species of the genus. Both new species exhibits new types of sex dimorphism previously unknown in Cordulegaster. Some traits related to the structure of male appendages and important details of colouration prevent both C. plagionyx sp. nov. and C. nachitschevanica sp. nov., from being reliably classified under any of two widely accepted groups of species within the genus Cordulegaster: the boltonii-group and the bidentata-group.
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Records of Odonata from two areas in the upper Baram area in Sarawak's Miri Division are presented. Sixty five species are recorded from the Sungai Sii area and sixty three from the Ulu Moh area. Notable records include Telosticta ulubaram, Coeliccia southwelli, Leptogomphus new species, Macromia corycia and Tramea cf. virginia. Rhyothemis regia is recorded from Sarawak for the first time.