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96
In 2014, 56 localities in four provinces of Lesser Himalaya in Pakistan were studied. A total of 28 species have been recorded. A female of the data deficient, threatened species Coeliccia vacca was recorded from Charhaan. The record of Drepanosticta carmichaeli is a new addition to the list of Odonata of Pakistan, and expand the range of this species further to the west. The taxonomical status of Ischnura aurora aurora – considered common in Pakistan, following baseline literature of Fraser (1933) – now turns out to be Ischnura aurora rubilio.
164
Damselflies recorded before the administrative partition of the Indian Subcontinent and now housed at National Insect Museum (NIM), Islamabad were reviewed and catalogued. This collection is the divided part of National PUSA Collection (NPC) transferred to the Pakistan during 1947. Data for this collection had never been available or published. A record of 104 taxa is reported herein. Few of the species were found double named, misidentified and not updated as per valid classification. Some of the specimens were found unidentified. All such issues were resolved by following regional literature.
37
The paper compiles records from four excursions to study the Odonata fauna of southern Guizhou, China. Between 2007 and 2010 in Xiaoqikong Park and Maolan National Nature Reserve, 104 taxa have been recorded. Some interesting species are discussed, compared with sibling taxa, and information on habitats and habits is given.
103
Between 2009 and 2016, a total of 174 Odonata species (Tab. 1) have been recorded in the Dehong Dai and Jingpo Autonomous Prefecture, China. 21 of these species are new records for China, and additional 26 taxa have to be described as new to science. Brief comments on selected species refer to morphological characters, distribution and seasonality.
88
145
Arising from a number of 2019 IUCN Red List assessments for a subset of Chinese Odonata, a series of corrections and taxonomic revisions were made to the World Odonata List. The rationale for these amendments is provided here. Paragomphus wuzhishanensis Liu, 1988 is shown to be a junior synonym of Paragomphus pardalinus Needham (1942). Epophthalmia kuani Jiang 1998 is synonymised as a junior synonym of Epophthalmia. elegans (Brauer, 1865) and Epophthalmia bannaensis Zha & Jiang, 2010 is treated as a junior synonym of Epophthalmia vittata Burmeister, 1839. Idionyx pseudovictor Xu, 2013 is shown to be junior synonym of Idionyx claudia Ris, 1912 and Sympetrum anomalum Needham, 1930 is treated as a junior synonym of Sympetrum maculatum Oguma, 1922.
147
This paper presents the faunistic results of three short field excursions conducted in spring and early summer of 2018 in Kosovo, considering dragonfly fauna one of the most understudied countries of Europe. This study presents first systematic dragonfly research in Kosovo. Within a total of 13 field days between end of April and end of June 2018, 60 sites were surveyed and 44 dragonfly species were found. Significant results include the first documented report of 15 species for Kosovo. New data on several other species with a broader European concern or generally rare on the West Balkan peninsula, i.e. Coenagrion ornatum, Anax ephippiger, Caliaeschna microstigma, Cordulegaster heros, C. bidentata, Somatochlora flavomaculata, and Sympetrum flaveolum, are also presented. The overview of all visited sites is included. Altogether, 47 dragonfly species are now reported for Kosovo.
39
151
The paper presents new odonatological data from the formerly unstudied islands Patnanungan and Jomalig as well from the northeastern part of Polillo, The Philippines. Four species are new for the PolilloIsland group, including one new taxon Anax parthenope julius – for The Philippines. Currently, 78 odonate taxa are known from the Polillogroup.
119
An odonatological survey, based on sighting and photographic documenting, was conducted in Palawan and in Cuyo Island. Ten species were found in Cuyo Island raising the number of known species from five to eleven. There are six additions to the Cuyo Island fauna: Agriocnemis f. femina, Ischnura senegalensis, Pseudagrion microcephalum, Brachydiplax c. chalybea, Neurothemis fluctuans and Neurothemis t. terminata. The occurrence of Coeliccia boettcheri, known from Cuyo, but not recorded during this survey, is discussed in some detail. Forty species were recorded in Palawan. Four new additions to the Palawan fauna were recorded: Lestes p. praemorsus, Pseudagrion microcephalum, Xiphiagrion cyanomelas, and Anax guttatus. Neurobasis daviesi, a rare calopterygid damselfly endemic in Palawan, was encountered at the northern side of Cleopatra Needle during the survey.
55
Notes on a small Odonata collection from Tawi-Tawi, Sanga-Sanga and Jolo islands, Philippines
(2012)
Sulu region is among the least explored faunal region in the Philippine archipelago. Odonatologically, this region is poorly studied until recently. Presently a survey conducted in July 1 – 14, 2011 revealed ten new records in Tawi-Tawi raising the total number of Odonata to 54. Three new species records were made for Sanga-Sanga raising the known number in that island to 34. Three species were recorded for the first time in Jolo raising the total number to 18. One new species of damselfly was found and several questionable and possible new species of dragonflies were documented.
60
The Odonata fauna of Balabac Island, Philippines was studied in March 2013. A total of 41 species under 33 genera were recorded. Twenty-eight species were recorded for the first time in the island. One genus – Mortonagrion was recorded for the first time in the Philippines. Mortonagrion astamii spec. nov. and Prodasineura poncei spec. nov. are new to science and are described. Three previously recorded species remained elusive and not seen during the survey.
59
Odonata survey was conducted in Talaingod, Davao del Norte, Mindanao Island. Four major sites were explored in Barangay Santo Niño from December 26 – 30, 2012. Thirty five species under eleven families including one new species were found representing the first odonatological record in the province of Davao del Norte. Three species need further study while Orthetrum glaucum represents a new record for the island of Mindanao. Coeliccia exoleta population, a vulnerable species in the IUCN Red List of Threatened Species, was found.
66
Balut and Sarangani islands are two small landmasses situated off the coast of Davao Occidental, Mindanao Island. Despite recent increase on odonatological data from various islands in the Philippines, these two remote islands have never been explored. Hence, a short survey was conducted on first week of April and November 7 – November 14, 2010 on all freshwater systems in these two islands. Twenty-five species under seven families and 21 genera were found representing the first Odonata record for the two islands.
34
Odonata were recorded and voucher specimens were collected between August 3- 13 and August 16-20, 2011. This account lists 51 species of Odonata for both islands, 47 species in Siargao and 24 species in Bucas Grande. Thirty seven species are new island records for Siargao Island while the 24 species recorded in Bucas Grande represent the first island records. Three species are new to science, and two of which (Drepanosticta schorri n. sp., Pseudagrion schieli n. sp.) are described in the present paper. Four species previously listed (Hämäläinen & Müller, 1997) remained elusive during the present survey.
38
Odonata fauna of Diomabok Lake and its surroundings, Davao Oriental, Mindanao Island, Philippines
(2011)
During three visits in October and December 2010 and May 2011, a total of 56 Odonata species was recorded. All species reported here represent first Odonata records in the area. The most noteworthy discoveries were one novelty (Hydrobasileus vittatus) to the Philippine fauna and two first records (Tetracanthagyna brunnea and Aethriamanta gracilis) from Mindanao Island. Seven species represent either new species to science or potentially new species; one Drepanosticta and one Amphicnemis are new to science, and another Drepanosticta, Amphicnemis, Pseudagrion, Gomphidia and Urothemis are potentially new to science.
23
Polillo is a small group of island (27 islands in total) east of central Luzon (Figure 1). It is made up of four main island viz. Polillo I, Patnanungan I, Jomalig I and Palasan I and several islets. This island group has relatively flat to gentle sloping terrain and the highest point is only 300 m asl (Mt. Maluhod) in Polillo Island (ca 700 km²) which is the largest in the group and the third largest island in greater Luzon biogeographic region.
27
Polillo Island is the largest among the Polillo group of islands. It is the third (ca 700sq km) largest island of the Luzon biogeographic region. The island lies in central eastern Luzon, at the western edge of Pacific Ocean and has a seasonal climate with distinct dry and wet season. The island is biologically rich with several insular species or subspecies (McGregor, 1910; Robinson, 1911; Manuel, 1957). The recent Odonata survey revealed the rich Odonata fauna of the island (Villanueva, 2010). This paper provides an update on the known Odonata based on recent fieldwork in 2010.
43
Six Odonata species were collected during night light trapping on the Mambilla Pla-teau, Taraba State Nigeria. Being predominantly diurnal insects, odonates captured in light traps have always been considered as an anomaly. The new data initiated an extensive interrogation of all records on Odonata collected near artificial light sources.A total of 415 records (402 published and 13 new) are presented here with a sum-mary of previous discussions and new discussion points. The general conclusion is that odonates are mainly confused by, rather than attracted to the light. New ave-nues for further research in this field are suggested based on previous important stu-dies undertaken on Odonata morphology and physiology.
118
A catalogue of all types of subfamilies Protoneurinae and Disparoneurinae currently housed in the entomological collection of the Museum für Naturkunde − Leibniz Institute for Evolution and Biodiversity Science in Berlin (Germany) is presented. It includes current status of the familygroup, genusgroup and speciesgroup names, transcriptions of data labels and references to the original descriptions.
153
The changes in coloration with age in males and females of Coeliccia mattii Phan & Kompier, 2016 from the Central Highlands of Vietnam are documented. Immature forms have extensive yellow markings on the thorax and S9 is yellowish, while in the fully mature male form, the thorax is largely black and covered in pruinosity and S9 is completely black. S9 is yellowish in the immature female while S9 is black in the fully mature female.
133
A checklist of 49 damselfly species from 12 families (Odonata: Zygoptera) recorded from Kon Ka Kinh National Park is provided. A first description of the female Protosticta socculus Phan & Kompier, 2016, is given. Burmargiolestes cf. laidlawi Lieftinck, 1960 and three apparently new species, two Coeliccia and one Protosticta species, are recorded, all of which are to be described in the future.
32
During three field trips in 2009-2010 to Xuan Son National Park in Phu Tho province in northern Vietnam, a total of 13 species of damselflies of the superfamily Calo-pterygoidea were recorded. These records are documented here with field photo-graphs of living damselflies. Observations on their biology and behaviour are also noted. Three of the species are reported from Vietnam for the first time: Rhinocypha arguta, an undescribed Matrona species and Vestalaria miao. The last species was first found in Huu Lien Nature Reserve in Lang Son province in June 2008.
124
A record of male and female specimens of Sinolestes editus Needham, 1930 collected in Dak Hro village (14°19'45'' N, 108°24'23'' E, alt. 1,420m a.s.l.), Dak Roong Commune, K’Bang District, Gia Lai Province, Central Highlands of Vietnam is present with a detailed description of the morphology of these two specimens.
112
We describe three new species of damselflies from streams draining into Lake Kutubu in Southern Highlands Province, Papua New Guinea. They are Drepanosticta johncanni sp. nov. (Platystictidae), Pseudagrion parafarinicolle sp. nov. (Coenagrionidae) and Wahnesia kutubuensis sp. nov. (Argiolestidae). Diagnostic characters of the males and, where available of the females, are illustrated and the probable affinities of the new species are discussed.
77
99
Palaeosynthemis opaca sp. nov., a new dragonfly from Papua New Guinea (Anisoptera: Synthemistidae)
(2016)
A new species of the synthemistid genus Palaeosynthemis is described from the northern slopes of Papua New Guinea’s central cordillera. It is distinguished from all congeners by having females with large, nearly black patches at the bases of both wings. The male is most similar to P. cervula and P. feronia, but it differs from those species in having superior anal appendages less than three times as long as S10, basally enlarged and otherwise unarmed vs basally not enlarged (feronia) and almost four times as long as S10 and armed (cervula). Characters of the adults (both sexes) are illustrated and the affinities of the species are discussed.
106
A new species of the genus Teinobasis Kirby is described from the Muller Range in Western Province, Papua New Guinea. Its male is distinguished from all other Teinobasis species by having a pale labrum, an extensively bright orange thorax, and ventrally bowed superior anal appendages that are markedly shorter than the plump, apically rounded inferiors. Characters of the male are illustrated, and the affinities of the new species are discussed.
64
71
A new species of the synthemistid genus Palaeosynthemis is described from the Trauna River valley in Western Highlands Province, Papua New Guinea. The new species is most similar to P. cyrene from which it can be distinguished, among other characters, by the coloration of the pterostigma (jet-black in the new species vs brownish yellow in P. cyrene) and of the wing bases (not darkened vs strongly darkened). The new species also differs from P. cyrene in having a narrow, almost parallel-sided yellow lateral synthoracic stripe and a well-defined yellow marking along most of the ventral margin of the metepimeron. In P. cyrene the lateral synthoracic stripe is markedly wider and tapered, and the yellow element along the ventral margin of the metepimeron is absent. Characters of the adult male are illustrated and the affinities of the species are discussed.
90
Two new species of damselflies are described from central Halmahera in North Maluku Province, Indonesia. They are Drepanosticta pararudicula sp. nov. (Holotype MZB. ODON. 19257) and Nososticta halmahera sp. nov. (Holotype MZB. ODON. 19265). The two species are most similar to the Moluccan taxa D. rudicula and N. moluccensis respectively and their descriptions bring the total number of Drepanosticta species known from Halmahera to five and of Nososticta to two.
58
149
6.1
Dragonflies were collected at ten localities in five counties (sous-préfectures) in southern Bénin and are presently identified. A total of 73 species were found, including 45 new records for the country. The list of the odonates of Bénin currently comprises 86 species although more than 100 can probably be expected.
141
Records of Odonata collected in Gunong Mulu National Park in Sarawak are presented. Between 2005 and 2019, in 12 surveys that lasted between one week and five months, 163 species were collected. The collections from Gunong Mulu National Park are of importance for the taxonomic study of dragonflies and damselflies in Borneo; several species have been described based on material collected in the Park.
78
Records of larval rearing in the Gunung Mulu National Park, Sarawak, Malaysia carried out in 2014 are presented. In total, larvae of 27 species were collected. Larvae of eleven species (22 individuals) were successfully reared out, one individual is currently still being reared. An additional three species were collected right after emergence, with the adult still sitting on its exuvia. Most notable are the samples of Orthetrum borneense, Leptogomphus cf. pendleburyi, Coeliccia cf. nemoricola 1, Coeliccia cf. nemoricola 2, Heliocypha biseriata and Elattoneura analis whose final instar larvae are undescribed.
168
1-23: Odonata collected in 2021 in Azerbaijan, including new data on Gomphus schneiderii Selys, 1850 and Libellula pontica Selys, 1887 (Nataly Yu. Snegovaya); published: 21.01.2022 -
25-32: On the occurrence of Gomphus vulgatissimus (Linnaeus, 1758) and G. schneiderii Selys, 1850 in Azerbaijan – a brief discussion of the known status quo (Martin Schorr & Nataly Yu. Snegovaya); published: 24.01.2022 -
33-36: Reply to Schorr & Snegovaya (2022), this volume of IDF-Report (Jean-Pierre Boudot); published: 03.02.2022 -
37-42: Editor's response to Boudot (2022) (Martin Schorr); published: 16.02.2022 -
43-55: Call it schneiderii, but document which identification key you used (Martin Schorr); published: 17.02.2022 (Hinweis: dies ist eine geänderte Version des IDF-Report 168 mit Stand 17.02.2022, welche die früheren Versionen ersetzt).
127
The article presents new faunistic data on 33 Odonata species, based on the material collected by the author in 2012, 2016 and 2017 and a systematic research in 2018 throughout the Nakhichevan Autonomous Republic (AR). Onychogomphus assimilis (Schneider, 1845) is a new record for the fauna of Azerbaijan. Eight species were registered for the first time for the territory of Nakhichevan AR: Lestes virens Rambur, 1842, Coenagrion scitulum (Rambur, 1842), Aeshna mixta Latreille, 1805, Anaciaeschna isoceles (Müller, 1764), Anax parthenope (Selys, 1839), Sympetrum sanguineum (Müller, 1764), Crocothemis erythraea (Brullé, 1832), and Selysiothemis nigra (Vander Linden, 1825).
142
85
Two new Cordulegaster species are described and illustrated by drawings, photographs and SEM images. The first one, C. plagionyx sp. nov., was discovered in NW Azerbaijan in a low-mountain forest landscape; the second, C. nachitschevanica sp. nov., occurs in the subalpine zone of Nakhichevan AR, south of the main territory of Azerbaijan. Both new taxa look generally similar to C. insignis Schneider, 1845; however, each of them reveals unique features and distinctive combinations of characters that set them apart from other species of the genus. Both new species exhibits new types of sex dimorphism previously unknown in Cordulegaster. Some traits related to the structure of male appendages and important details of colouration prevent both C. plagionyx sp. nov. and C. nachitschevanica sp. nov., from being reliably classified under any of two widely accepted groups of species within the genus Cordulegaster: the boltonii-group and the bidentata-group.
87
This article contains new faunistic data on 53 Odonata species based on material collected by the authors in Azerbaijan between 2013-2014 and added by revising an old collection made by A.V. Bogachev in the 1930-1940s. Of these, 13 species are new for the country: Lestes dryas, L. sponsa, L. virens, Coenagrion hastulatum, C. lunulatum, C. ornatum, C. pulchellum, Aeshna affinis, Brachytron pratense, Cordulegaster picta, Somatochlora flavomaculata, Sympetrum flaveolum, S. vulgatum. The new locality of Cordulegaster picta is the easternmost for the species. Two new populations of a very rare species Cordulegaster vanbrinkae (discovered in the country in 2011) are found. Somatochlora flavomaculata and Brachytron pratense are generally very rare in the entire Caucasus. Pantala flavescens has been rediscovered in the country 100 years after the first record. A large population of Caliaeschna microstigma, a species very rare in Azerbaijan, has been found in the northern part of the country; it represents a peculiar dark morph whose features are described in detail. Particulary discussed are the variability of Coenagrion puella-complex in Azerbaijan and diagnostic features of two other Coenagrion species closely related to each other, C. ornatum and C. vanbrinkae.
143
Using a qualitative sampling approach to survey for Odonata along water systems inside and outside of four major protected areas in Southern Ghana (viz; Atewa Range Forest Reserve, Ankasa Forest Reserve, Owabi Wildlife Sanctuary and Bobiri Forest Reserve), a total of 66 Odonata species in seven families encompassing 28 Zygoptera and 38 Anisoptera were recorded. Libellulidae (n= 36) exhibited the highest number of species, followed by Coenagrionidae (n= 10), and Calopterygidae (n= 5). In considering the observed number of species exhibited by each protected area, the Atewa Range Forest Reserve exhibited the highest number of species (n= 51), followed by the Ankasa Forest Reserve (n= 47), the Owabi Wildlife Sanctuary (n= 44) and Bobiri Forest Reserve (n= 43). Disturbance tolerance species dominated in habitats outside the various forest reserves, whiles forest specialists predominated inside the reserves, indicating the quality of the forest cover therein.
93
Specimens of Neurothemis disparilis Kirby, 1889, N. fluctuans (Fabricius, 1793), N. fulvia (Drury, 1773), N. ramburii (Brauer, 1866), N. stigmatizans (Fabricius, 1775) and N. terminata Ris, 1911, including their subspecies, were studied with the main focus on the morphology of the vesica spermalis, wing maculation, wing venation, abdominal markings and vulvar scales. The results were compared with species descriptions and directly with type specimens where possible. The vesica spermalis, especially the medial process, is useful at least in separating species groups and supports the traditional differentiation methods using wing maculation and venation. The use of other characters in accessing specific status, coupled with known distribution patterns, is discussed. The following taxonomic changes are proposed: Neurothemis manadensis (Boisduval, 1835) stat. nov., Neurothemis papuensis (Lieftinck, 1942) stat. nov. and Neurothemis taiwanensis sp. nov. is described (27.5.1998, Kenting, Pingtung County/Taiwan, L. M. Juang leg.; holotype is deposited at Taiwan Forestry Research Institute, Taipeh, Taiwan). The type of Polyneura palliata Rambur, 1842 was rediscovered at MNHN and designated as lectotype; a lectotype for Neurothemis nicobarica Brauer, 1867 housed at NHMW is designated. The holotype of Neurothemis incerta Brauer, 1867 was rediscovered and synonymized with N. ramburii.
70
A selection of duplicates from the collection of Michel Edmond de Selys Longchamps was found at the Übersee-Museum Bremen/Germany (UMB). Selys determined a lot of Odonata in the UMB collection and sent 80 European and 76 exotic species to Bremen on 23 April, 1875. According to the labels 121 specimens could be assigned to this shipment and eleven specimens must have been sent to UMB in later years. This collection includes two paralectotypes (Progomphus gracilis Hagen inSelys, 1853;Palaemnema nathalia Selys, 1886) and seven syntypes (Rhinocypha trifasciata Selys, 1853; Dysphaea dimidiata limbata Selys, 1859; Argia sordida Hagen inSelys, 1865; Oxyagrion dissidens Selys, 1876; Oxyagrion haematinum Selys, 1876; Oxyagrion pavi-dum Hagen in Selys, 1876; Telagrion longum Selys, 1876). In addition, a male specimen of Euphaea tricolor subcostalis Selys, 1873 might also belong to the original syntype series. Altogether three specimens with labeled nomina nuda(Diplax catharina Selys, Diplax fausta Selys, Dythemis bilineata Hagen) and two labeled with manuscript names (Diplax marcellina Selys, Perithemis ovate Bates) are in this collection.
41
Based on data collected from literature, museum collections, national databases and personal communications, an up-to-date map of the worldwide distribution of Somatochlora sahlbergi is presented. A new hypothesis is presented indicating that occurrences are at least regionally correlated with palsa mires. Two examples of larval habitats in Europe are illustrated and described, including the first observation of reproduction in Norway and the first record of co-occurrence with Zygoptera in Europe. The exuvia of S. sahlbergi is illustrated and distinguishing features briefly discussed.
28
Based on the results of fieldwork and collecting in 2008 and 2009 and the evaluation of literature an updated national checklist of the Odonata of Kyrgyzstan is presented. The list comprises a total of 63 species, whereas 55 species were encountered in the field by the author, including five new for the country: Aeshna serrata, Onychogomphus lefebvrii, Orthetrum sabina, Crocothemis servilia, Selysiothemis nigra. 826 specimens of 49 species have been collected (dep. in coll. A. Schröter). All 55 species recorded in 2008 and 2009 are listed and annotated. Moreover, the unclear or controversial taxonomical status of several species is briefly debated. Interesting ecological observations include the emergence of Libellula quadrimaculata from running water and cleptoparasitism by Ischnura forcipata in spider webs.