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147
This paper presents the faunistic results of three short field excursions conducted in spring and early summer of 2018 in Kosovo, considering dragonfly fauna one of the most understudied countries of Europe. This study presents first systematic dragonfly research in Kosovo. Within a total of 13 field days between end of April and end of June 2018, 60 sites were surveyed and 44 dragonfly species were found. Significant results include the first documented report of 15 species for Kosovo. New data on several other species with a broader European concern or generally rare on the West Balkan peninsula, i.e. Coenagrion ornatum, Anax ephippiger, Caliaeschna microstigma, Cordulegaster heros, C. bidentata, Somatochlora flavomaculata, and Sympetrum flaveolum, are also presented. The overview of all visited sites is included. Altogether, 47 dragonfly species are now reported for Kosovo.
146
Previously unpublished records of Odonata from the states of Negeri Sembilan and Selangor in Peninsular Malaysia are presented. One hundred and eight species are listed, of which 77 were collected in Negeri Sembilan and 87 in Selangor. Fifteen of the species recorded from Negeri Sembilan and seven of those recorded from Selangor appear to be first records for the respective state. Notable records include Drepanosticta sp. cf hamadryas Laidlaw, 1931, Rhinocypha pelops Laidlaw, 1936, Acrogomphus ?malayanus Laidlaw, 1925, Heliogomphus kelantanensis (Laidlaw, 1902); Onychogomphus duaricus Fraser, 1924, Macromia cupricincta Fraser, 1924, Idionyx montana Karsch, 1891, Chalybeothemis chini Dow, Choong & Orr, 2007 and Hylaeothemis clementia
Ris, 1909 Ris, 1909. Provisional checklists of the Odonata known from Selangor plus the
Federal Territory of Kuala Lumpur (171 species) and for Negeri Sembilan (116 species)
are given in appendices.
145
Arising from a number of 2019 IUCN Red List assessments for a subset of Chinese Odonata, a series of corrections and taxonomic revisions were made to the World Odonata List. The rationale for these amendments is provided here. Paragomphus wuzhishanensis Liu, 1988 is shown to be a junior synonym of Paragomphus pardalinus Needham (1942). Epophthalmia kuani Jiang 1998 is synonymised as a junior synonym of Epophthalmia. elegans (Brauer, 1865) and Epophthalmia bannaensis Zha & Jiang, 2010 is treated as a junior synonym of Epophthalmia vittata Burmeister, 1839. Idionyx pseudovictor Xu, 2013 is shown to be junior synonym of Idionyx claudia Ris, 1912 and Sympetrum anomalum Needham, 1930 is treated as a junior synonym of Sympetrum maculatum Oguma, 1922.
144
We report here the results from field trips to collect Odonata in the northeastern parts of Kelantan state and the north of Terengganu state, Peninsular Malaysia. Eighty four species were collected, and four of these are new records for the state Kelantan and 10 are new records for the state of Terengganu. Notable records obtained from the field trips were Euphaea masoni, Coeliccia sameerae, Pseudagrion ?lalakense, Leptogomphus tioman and Macromia cupricincta. Checklists for Kelantan (140 species) and Terengganu (132 species) are given in an appendix.
143
Using a qualitative sampling approach to survey for Odonata along water systems inside and outside of four major protected areas in Southern Ghana (viz; Atewa Range Forest Reserve, Ankasa Forest Reserve, Owabi Wildlife Sanctuary and Bobiri Forest Reserve), a total of 66 Odonata species in seven families encompassing 28 Zygoptera and 38 Anisoptera were recorded. Libellulidae (n= 36) exhibited the highest number of species, followed by Coenagrionidae (n= 10), and Calopterygidae (n= 5). In considering the observed number of species exhibited by each protected area, the Atewa Range Forest Reserve exhibited the highest number of species (n= 51), followed by the Ankasa Forest Reserve (n= 47), the Owabi Wildlife Sanctuary (n= 44) and Bobiri Forest Reserve (n= 43). Disturbance tolerance species dominated in habitats outside the various forest reserves, whiles forest specialists predominated inside the reserves, indicating the quality of the forest cover therein.
142
141
Records of Odonata collected in Gunong Mulu National Park in Sarawak are presented. Between 2005 and 2019, in 12 surveys that lasted between one week and five months, 163 species were collected. The collections from Gunong Mulu National Park are of importance for the taxonomic study of dragonflies and damselflies in Borneo; several species have been described based on material collected in the Park.
139
138
Description of Hemicordulia tuiwawai sp. nov. from Kadavu Island, Fiji (Odonata: Corduliidae)
(2019)
Hemicordulia tuiwawai sp. nov. (Odonata: Corduliidae) is described and diagnosed based on material collected from Kadavu Island, Fiji; holotype: Wainitayuki River about 750 m above Baidamudamu village, -19.0916, 178.1038; 37 m a.s.l., 06 June 2016, M. Marinov leg. This species is distinguished from its congeners in the field by the contrasting colouration – dark green metallic body with bright yellow spots on the synthorax and base of the abdomen. This pattern is comparable to H. pacifica Selys, 1871. However, Hemicordulia tuiwawai sp. nov. can be recognised by the larger size and unique shape of the caudal appendages and genital hamule (in males) and vulvar scale (in females).
137
Records of Odonata from the southwest of Sri Aman Division and the extreme east of Serian Division in Sarawak are presented. The sampled areas are interesting not only because they are poorly known for Odonata but also because many are just to the south and west of the Lupar Line which is a division between the ancient Sunda shelf and more recent geological formations. Differences between the odonate faunas on either side of the Lupar Line are discussed. Eightyfive species of Odonata were recorded during the surveys reported on. The single most notable record is that of Coeliccia southwelli Dow & Reels, 2011, which represents a considerable extension to the known range of this species. Other interesting records include Telosticta dupophila (Lieftinck, 1933), T. species cf longigaster Dow & Orr, 2012, Podolestes parvus Dow & Ngiam, 2019 and Heliogomphus species cf olivaceous Lieftinck, 1961. Variation in the markings of Stenagrion dubium (Laidlaw, 1912) across its range is discussed and a gene tree using the COI marker is presented to illustrate the high variability of this species in this marker. However the variability in COI does not appear to be correlated with other characters.
136
Odonatological results of a trip across the southern Far East of Russia, from nearly the border of North Korea to Khabarovsk from July 1st to 16th, 2014, are presented. In total, 50 species were recorded, including those with limited presence in Russia, such as Paracercion calamorum, P. hieroglyphicum, P. plagiosum (for this species the 3rd Russian locality is reported), Pseudocopera tokyoensis, Stylurus annulatus (2nd Russian locality), Sinictinogomphus clavatus; Trigomphus citimus, Macromia daimoji (3rd Russian locality), M. manchurica, Deielia phaon, Lyriothemis pachygastra (2nd Russian locality). For S. annulatus, M. manchurica and D. phaon the northernmost known localities in the world are reported. D. phaon, earlier reported from one locality in Russia, have been found in five localities in Primorye and for the first time reported for Khabarovskiy Kray in general and Bol'shekhekhtsirskiy State Nature Reserve in particular. A trend of mutual exclusion of two abundant Shaogomphus postocularis epophthalmus and S. schmidti is supposed: the lowermost Ussuri River / Amur River, respectively. Variation in Paracercion spp. and Macromia amphigena fraenata is discussed.
135
During four field trips in Georgia, Armenia and Azerbaijan from 2010 to 2018, the author collected data of a total of 55 species (see Tab. 2 in Appendix). This study provides first insights into new or rare species in this ecoregion. Ischnura fountaineae and Cordulia aenea were found for the first time in Armenia. We also highlight the rediscovery of some species that were mentioned in the older literature but had not been confirmed since. An autochthonous population of Lestes macrostigma was discovered in Azerbaijan sixteen years after the single previous record by Dumont (2004). Original information is provided on the distribution of some rare species encountered in these countries. Finally, the identification of a puzzling Cordulegaster sp. observed in south Armenia is discussed briefly.
134
The Odonata collection deposited at the Museum of Comparative Zoology (MCZ) includes specimens of 634 taxa labeled as types. Fifteen of these have been incorrectly labeled as types (pseudotypes) and eight are apparently lost, leaving a total of 611 types currently deposited at MCZ. From these, 489 represent primary namebearing types (syntype/s, holotype, lectotype and neotype), 21 are probable primary types, and 101 are secondary types (paratype/s, paralectotype/s).
133
A checklist of 49 damselfly species from 12 families (Odonata: Zygoptera) recorded from Kon Ka Kinh National Park is provided. A first description of the female Protosticta socculus Phan & Kompier, 2016, is given. Burmargiolestes cf. laidlawi Lieftinck, 1960 and three apparently new species, two Coeliccia and one Protosticta species, are recorded, all of which are to be described in the future.
132
Phnom Tumpor is a scarcely accessible basalt table mountain in the Cardamom Mts. in Pursat Province of Cambodia. On top surface it bears tall evergreen forest (ca 1100 m a.s.l.), concealing a slow rivulet, O'Gran, being a chain of deep pools. It was examined odonatologically on March 1418th, 2019. Six common species were recorded in dry and burnt scrub on the Phnom Tumpor slopes and ten on the forested upper surface at O'Gran, among them Polycanthagyna erythromelas (Selys, 1891) and Macromia sp. cf. pinratani Asahina, 1987 for the first time in Cambodia. The peculiarities of the males of Coeliccia kazukoae Asahina, 1984 from Phnom Tumpor and the problem of distinguishing females of M. pinratani and M. moorei are discussed.
131
Survey in three protected areas of Chhattisgarh reveals the presence of 50 species of Odonata belonging to 34 genera and 9 families. Specimens were sampled from different lotic and lentic ecosystems. 17 species were exclusively found in or around running water. Larvae or exuviae of 23 species were found and photographed. Habitat availability, larval abundance, species composition and phenology are discussed.
129
New data on Odonata of the Preah Sihanouk Province of Cambodia obtained in March 2017 and November 2018 are presented. The presence of Onychargia atrocyana Selys, 1865 in Cambodia is confirmed. Twentyone species are added to the known fauna of the Kbal Chhay Waterfall environs, 19 species to that of Ream Peninsula and 4 to that of Koh Rong Island. The total number of species registered for Kampong Saom Peninsula amounts to 74. The presented data are rather of historical importance since most of the remaining forest has been quickly and irreversibly logged a few months ago. The validity of Gynacantha demeter Ris, 1911 as a species distinct from G. dohrnii Krüger, 1899 is doubted.
128
Globally urban wetlands are under high anthropogenic pressure of degradation. Urban wetlands are hotspots for species losses and rapid turnover in species assemblages. Therefore, studying such wetlands may provide an estimate of the pace of local extinction, concerning wetlanddependent species such as odonates. We undertook a study to document odonate species across a tropical urbanization gradient. We sampled six localities across the gradient across the Mula River that flows through the Pune City, India. We sampled adult odonates using a newly devised Halfcircle Point Count method from September 2016 to March 2017. We took multiple temporal replicates per site. We also sampled larvae across six sites once in November 2016. We measured site characteristics such as canopy cover, solid waste, and water turbidity to understand the level of disturbance at each site. We recorded 41 odonates, six species (primarily Gomphidae members) exclusively from the larval sampling. We did not find the localization of species in a particular site across the urbanization gradient, possibly because we sampled a relatively short stretch of the river to capture the variation. Here, we update the Odonata list of Pune including data on larvae. We demonstrate that larval sampling complements Odonata surveys, especially in recording Gomphids.
We recommend future research to include a longer timespan and extensive sampling area.
127
The article presents new faunistic data on 33 Odonata species, based on the material collected by the author in 2012, 2016 and 2017 and a systematic research in 2018 throughout the Nakhichevan Autonomous Republic (AR). Onychogomphus assimilis (Schneider, 1845) is a new record for the fauna of Azerbaijan. Eight species were registered for the first time for the territory of Nakhichevan AR: Lestes virens Rambur, 1842, Coenagrion scitulum (Rambur, 1842), Aeshna mixta Latreille, 1805, Anaciaeschna isoceles (Müller, 1764), Anax parthenope (Selys, 1839), Sympetrum sanguineum (Müller, 1764), Crocothemis erythraea (Brullé, 1832), and Selysiothemis nigra (Vander Linden, 1825).
126
We present records of 99 Odonata species from the Réserve Naturelle des Gorilles de Lésio-Louna (RNGLL) in the Republic of Congo (Congo-Brazzaville) collected during a short survey from 14 January to 01 February 2017. It is the first systematic Odonata inventory for the RNGLL and for the Congolese part of the species-rich Batéké Plateau. A short introduction is given about the existing knowledge of dragonflies and damselflies from the country. Amongst the recorded species eight are new for the country list, raising it to at least 208 species. Some of the recorded species are endemic and characteristic for sandy streams and rivers of the Batéké Plateau. The potential diversity of the plateau in comparison to other regions of the country is discussed.
125
A total of 20 Odonata species were recorded from 15 water bodies (3 natural water bodies and 12 manmade reservoirs) in a desert landscape west of the Namak Lake in the northwest of the Central Plateau of Iran, from 27 May to 20 June, 2017. The study area included Qom province and the northern part of Esfahan province (34.07 to 35.14 N, 51.33 to 50.89 E). For Qom province, 17 out of 18 species found are new provincial records. In addition Selysiothemis nigra, Orthetrum chrysostigma and Orthetrum taeniolatum are new findings for Esfahan province. The species around the manmade reservoirs are characterized by a broad ecological amplitude ("generalists") while species assemblages of natural water bodies consisted of more sensitive species. In the arid climate of central part of Iran, manmade reservoirs are major habitats for Odonata species. Although these reservoirs have increased the amount of habitat for generalist species, they do not foster sensitive and specialist species. Furthermore, considering the more intense droughts predicted for these areas in the future, and the current scenario of environmental degradation, some sensitive species may be in danger of local extinction.
124
A record of male and female specimens of Sinolestes editus Needham, 1930 collected in Dak Hro village (14°19'45'' N, 108°24'23'' E, alt. 1,420m a.s.l.), Dak Roong Commune, K’Bang District, Gia Lai Province, Central Highlands of Vietnam is present with a detailed description of the morphology of these two specimens.
123
More Odonata found at the Cardamonean foothills in Koh Kong Province of Cambodia in 2014-2018
(2018)
Still unpublished data on Odonata of the coastal foothills of the Cardamom Mts. in Koh Kong Province, SW Cambodia obtained in 2014-2018 are provided. Tetrathemis flavescens Kirby, 1889 is for the first time reported for Cambodia and Risiophlebia guentheri Kosterin, 2015 for SW Cambodia. The photographic records from this area of Gynacantha basiguttata Selys, 1882, Gynacantha demeter Ris, 1911, Heliaeschna crassa Krüger, 1899, Amphithemis curvistyla Selys, 1891 and Pornothemis serrata Krüger, 1902 have been confirmed by specimens. The known local faunas of two neighbouring rivulets in Koh Kong Province, rich in Odonata, are updated and summarised.
122
Records of Odonata from Gunung Melatai, Nanga Gaat and the Kastima Logging area, all in Sarawak’s Kapit Division, are presented. The most notable records are of Matronoides cyaneipennis Förster, 1897 and Heliogomphus blandulus Lieftinck, 1929. A distribution map for Matronoides cyaneipennis and updated distribution maps for three species from the Coeliccia borneensisgroup are given. Tentative identifications to species of previously published records of Idionyx females are given. The genus Heliogomphus in Borneo, Peninsular Malaysia and Singapore is reviewed and a simple one marker molecular analysis is presented for the genus in this region. Based on reexamination of specimens from the genus and the molecular results, an additional member of the genus is reported from Borneo: H. sp. cf olivaceus Lieftinck, 1961. Although both morphological and molecular results remain incomplete, it does appear likely that there is at least one more species of Heliogomphus present in Borneo than has been recognised until now, and that H. borneensis Lieftinck, 1964 may be a junior synonym of H. kelantanensis (Laidlaw, 1902).
121
Macromidia genialis buusraaensis subspecies nova (Odonata, Synthemisidae s.l.) from eastern Cambodia
(2018)
Macromidia genialis buusraaensis subspecies nova is described from 10 males and 2 females from three localities of Mondulkiri Province, the eastern Cambodia (the type locality: a brook downstream Buu Sraa Waterfall 12°34' N 107°25' E). The new subspecies is close to M. g. shanensis Fraser, 1927 and differs from it by an additional, and the broadest, yellow spot on S7. It is partly syntopic with Macromidia rapida Martin, 1907. A brief overview of the current knowledge of the genus Macromidia Martin, 1907 is provided.
120
Results of a brief examination of the spring aspect of Odonata at the Abrau Peninsula and some nearby localities on 2127th May 2018 are presented. Lestes dryas and Caliaeschna microstigma are reported for the Peninsula for the first time, revealing the world northernmost record of the latter. Some old data of Odonata at Novorossiysk and some recent erroneous data on the Krasnodarskiy Kray are referenced and discussed. The known Odonata fauna of the Abrau Peninsula reaches 38 species.
119
An odonatological survey, based on sighting and photographic documenting, was conducted in Palawan and in Cuyo Island. Ten species were found in Cuyo Island raising the number of known species from five to eleven. There are six additions to the Cuyo Island fauna: Agriocnemis f. femina, Ischnura senegalensis, Pseudagrion microcephalum, Brachydiplax c. chalybea, Neurothemis fluctuans and Neurothemis t. terminata. The occurrence of Coeliccia boettcheri, known from Cuyo, but not recorded during this survey, is discussed in some detail. Forty species were recorded in Palawan. Four new additions to the Palawan fauna were recorded: Lestes p. praemorsus, Pseudagrion microcephalum, Xiphiagrion cyanomelas, and Anax guttatus. Neurobasis daviesi, a rare calopterygid damselfly endemic in Palawan, was encountered at the northern side of Cleopatra Needle during the survey.
118
A catalogue of all types of subfamilies Protoneurinae and Disparoneurinae currently housed in the entomological collection of the Museum für Naturkunde − Leibniz Institute for Evolution and Biodiversity Science in Berlin (Germany) is presented. It includes current status of the familygroup, genusgroup and speciesgroup names, transcriptions of data labels and references to the original descriptions.
117
In the period 18th – 31st May 2017, 33 localities were examined for Odonata in the Central Zagros area of Iran: 16 in Markazi Province, 14 in Lorestan Province and 3 in Esfahan Province; in 27 of those localities Odonata were found, 25 species in total. For Markazi Province, only one species, Calopteryx splendens, was previously reported (and also found by us), so of 17 species found there 16 are formally new provincial records (Epallage fatime, Lestes barbarus, Coenagrion ornatum, C. persicum, Enallagma cyathigerum, Ischnura elegans, I. intermedia, I. pumilio, Platycnemis kervillei, Anax imperator, Anaciaeschna isoceles, Caliaeschna microstigma, Libellula depressa, Onychogomphus lefebvrii, Orthetrum brunneum, Sympetrum fonscolombii). Of 17 species
found in Lorestan, 5 are new for this province (L. barbarus, Aeshna mixta, Orthetrum taeniolatum, Sympetrum arenicolor, S. striolatum). Only two species were seen in Esfahan Province, in which little time was spent. Notes on variation and taxonomy are provided for Sympecma paedisca, C. ornatum (considered to be a senior synonym of C. vanbrinkae because of variation in the presumed main diagnostic character), E. cyathigerum, I. elegans, Gomphus schneideri (including discussion of G. amseli), as well as notes on habitats of most species and the general characteristics of the area.
115
Records of Odonata from the Lanjak Entimau Wildlife Sanctuary (LEWS) in Sarawak, Malaysian Borneo are presented. Previous records of Odonata from LEWS are critically examined. One hundred and ten species have been recorded within the sanctuary, including three that have yet to be found outside (Drepanosticta adenani, Telosticta iban and "Elattoneura" mauros); records of four more species are regarded as in need of confirmation, those of another six are incorrect. In addition to the three species only known from LEWS, other notable records include: Drepanosticta sbong, Dysphaea lugens, Euphaea sp. cf basalis, Pericnemis kiautarum, Burmagomphus insularis, Gomphidia caesarea, Merogomphus species, Phaenandrogomphus safei, Macromia callisto and Idionyx montana. A fresh illustration of the anal appendages of Drepanosticta sbong in lateral view is provided to make up for inaccuracies in the original illustration. The taxonomy of Phaenandrogomphus safeii is briefly discussed. Zygonyx errans Lieftinck, 1953 is considered a subspecies of Z. ida not Z. iris Additional records from areas adjacent to LEWS are given in an appendix.
114
New field sampling was carried out in June 2011 in northern Moldova, the most unstudied area in the Republic of Moldova. A total of 19 Odonata species was recorded during a field trip, with Libellula fulva and Gomphus flavipes new for Moldova. The presence of species of European concern is discussed. Photos of all visited habitats are included. The fauna of Moldova mostly consists of species which are able to survive in biotopes with high anthropogenic pressure.
113
Sympetrum fonscolombii is for the first time reported for Tuva (Tyva Republic, Russia), as found in the UbsuNur Depression. New data are provided on Odonata of the Turan or TuranUyuk Depression of Tuva, including the first record of Somatochlora exuberata in Tuva beyond the Todzha Depression and Coenagrion armatum, C. ecornutum, Aeshna juncea, A. grandis, Somatochlora graeseri and Libellula quadrimaculata for the first time reported for the Turan Depression. New distributional data and comments on Ophiogomphus spinicornis Selys, 1878 are added. Somatochlora alpestris found at Lake Oyskoe is for the first time reported for the southern Krasnoyarskiy Kray.
112
We describe three new species of damselflies from streams draining into Lake Kutubu in Southern Highlands Province, Papua New Guinea. They are Drepanosticta johncanni sp. nov. (Platystictidae), Pseudagrion parafarinicolle sp. nov. (Coenagrionidae) and Wahnesia kutubuensis sp. nov. (Argiolestidae). Diagnostic characters of the males and, where available of the females, are illustrated and the probable affinities of the new species are discussed.
111
Part 1, Quoc Toan Phan and Van Quang To, page 1-6:
The genus Megalestes Selys, 1862 in Vietnam, with first description of female of Megalestes australis Karube, 2014 (Odonata: Zygoptera: Synlestidae)
Abstract: The male secondary genitalia and the caudal appendages of males and females of the three confirmed Vietnamese Megalestes species (M. australis, M. haui and M. micans) are figured. The female sex of M. australis is described for the first time.
Part 2, Quoc Toan Phan and Van Quang To, page 7-19:
Odonata checklist from Son Tra Nature Reserve, Da Nang city, central Vietnam
Abstract: A checklist of 44 odonate species (21 Zygoptera and 23 Anisoptera) from Son Tra Nature Reserve, central Vietnam is provided. Idionyx thailandica Hämäläinen, 1985 is recorded for the Vietnamese fauna for the first time, a new Coeliccia sp. is awaiting description in the near future, and the taxonomic status of specimens of the genus Leptogomphus remains unsettled and will have to be solved in the future.
110
Results are presented of a brief odonatological examination of the Abrau and Taman' Peninsulas at the northwesternmost Caucasian Black Sea coast in Krasnodarskiy Kray, Russia, on July 2026, 2017. Twenty three species have been observed at the former peninsula and five at the latter. The Odonata records at the Abrau Peninsula are summarised, to include 34 species.
108
A survey of Odonata on the Indonesian island of Belitung is reported. The work of Belitung Biodiversity Observer on Odonata is briefly outlined. Sixty four species were recorded during the survey, including two new records for the island. A checklist of the known odonate fauna, consisting of 105 species, of the island is given in an appendix.
107
Part 1, Oleg E. Kosterin and Vladimir I. Solovyev, page 1-43:
Odonata found in mid-summer 2015 and 2016 at the north-westernmost Black Sea Coast of the Caucasus, with the first record of Cordulegaster picta Selys, 1854 in Russian Federation
Abstract: Results are presented of brief odonatological examination of the Black Sea coastal northwesternmost spurs of the Caucasus between Anapa and Gelendzhik (mostly at Kabardinka village), Russia, in late July/early August 2015 and early-mid July 2016. In total, 28 Odonata species were found, including Cordulegaster picta for the first time in Russia. For C. picta and Caliaeschna microstigma, the world’s northernmost records were made. New localities of species rare in this area are reported: one for Coenagrion scitulum and three for Selysiothemis nigra, including their breeding habitat. Numerous migrant individuals of Pantala flavescens were observed in many localities in 2015 but none in 2016. Observations on trophic activity of Aeshna affinis and A. mixta are reported, the former showing predominantly matutinal and vespertinal activity and the latter diurnal activity. Occurrence of the Chalcolestes spp. in the Caucasus is discussed.
Part 2, Oleg E. Kosterin, page 45 - 57:
Calopteryx virgo feminalis subsp. nov., a long known under the same name but hitherto formally nameless subspecies from the Caucasian Black Sea Coast
Abstract: The populations of Calopteryx virgo of the Black Sea Coast of the Caucasus have females with the distal hindwing part darkened and males with the underside of S10 and appendages whitish, They are known for more than a century and deserve a subspecific status but since the name feminalis Bartenev, 1910 proposed to them is unavailable, a new subspecies is formally erected under the same name, Calopteryx virgo feminalis Kosterin subsp. nov., with the following type locality: Russia, Krasnodarskiy Kray, Gelendzhik Municipality, Kabardinka village, the Doob River lowermost reaches, 44°38'26-53'', 37°55'55''-57'58'' E.
106
A new species of the genus Teinobasis Kirby is described from the Muller Range in Western Province, Papua New Guinea. Its male is distinguished from all other Teinobasis species by having a pale labrum, an extensively bright orange thorax, and ventrally bowed superior anal appendages that are markedly shorter than the plump, apically rounded inferiors. Characters of the male are illustrated, and the affinities of the new species are discussed.
105
Results are presented of an odonatological survey of 23 localities in Thala Barivat District of Stung Treng Province, northern Cambodia, on July 26 – August 1, 2016. Most localities were situated in areas of open low deciduous dipterocarp forests on gravel soils, some at hillside areas of tall evergreen dipterocarp forest. The great Mekong River right bank was studied within 7 km downstream of its Nimith (Nimet, Khon Thai, Labak Koun, Khone Pha Pheng) Waterfall. In total, 55 species were found, of which 52 identified to species and three to genus. Two species, Gynacantha saltatrix Martin, 1909 and Macrogomphus matsukii Asahina, 1986, were recorded in Cambodia for the first time. Five obligatory lotic species were found at the Mekong River, namely Dysphaea gloriosa, Prodasineura coerulescens, Burmagomphus asahinai, Nychogomphus duaricus and Onychothemis testacea, including tenerals of P. coerulescens and B. asahinai. Most probably these species breed in the Mekong reach which is enriched with oxygen downstream of the great waterfall cascade.
104
Drepanosticta kosterini sp. nov. (holotype ♂, from Gunung Penrissen, Kuching Division, Sarawak, Malaysian Borneo, deposited in RMNH) is described from both sexes. It is the sister species of D. actaeon Laidlaw, 1934; a fresh description of the male of D. actaeon and the first description of the female are given, along with discussion of variation in this species. Both D. actaeon and D. kosterini are considered to belong to a species group also including D. rufostigma (Selys, 1886) and a preliminary discussion of variation in this species is given, along with illustrations of both sexes. A neighbour joining COI gene tree for D. actaeon and D. kosterini is presented. The relationships of D. actaeon, D. kosterini and D. rufostigma to other members of the Platystictidae are briefly discussed.
103
Between 2009 and 2016, a total of 174 Odonata species (Tab. 1) have been recorded in the Dehong Dai and Jingpo Autonomous Prefecture, China. 21 of these species are new records for China, and additional 26 taxa have to be described as new to science. Brief comments on selected species refer to morphological characters, distribution and seasonality.
102
DragonflyIndia Meet 2016
(2017)
101
The Odonata fauna of flat marshy areas of the Gulf of Siam coast in Koh Kong Province of Cambodia, containing 55 species, is considered. The published data of 2010- 2013 and new data of 2014 and 2016 on the surroundings of Koh Kong town are compiled and the first data on the area of large swampy Melaleuca forests at Andoung Tuek village are presented as well as some occasional photographic records. Gynacantha bayadera, Lyriothemis mortoni and Pornothemis serrata were for the first time recorded for Cambodia. Mortonagrion falcatum was found unexpectedly abundant at Andoung Tuek.
99
Palaeosynthemis opaca sp. nov., a new dragonfly from Papua New Guinea (Anisoptera: Synthemistidae)
(2016)
A new species of the synthemistid genus Palaeosynthemis is described from the northern slopes of Papua New Guinea’s central cordillera. It is distinguished from all congeners by having females with large, nearly black patches at the bases of both wings. The male is most similar to P. cervula and P. feronia, but it differs from those species in having superior anal appendages less than three times as long as S10, basally enlarged and otherwise unarmed vs basally not enlarged (feronia) and almost four times as long as S10 and armed (cervula). Characters of the adults (both sexes) are illustrated and the affinities of the species are discussed.
98
Results of an odonatological survey of Mondulkiri Province of Cambodia, at the foothills and Central Plateau of the Annamese Mts. in June 8 – June 17, 2014 are presented. Of 106 Odonata species met (46 zygopterans, 60 anisopterans), 97 were identified to previously known named species, of which 15 are reported for the first time for Cambodia, namely Mnais mneme Ris, 1916, Rhinocypha seducta Hämäläinen et Karube, 2001, Philoganga loringae Fraser, 1927, Rhinagrion hainanense Wilson et Reels, 2001, Amphiallagma parvum (Selys, 1876), Ceriagrion chaoi Asahina, 1967, Paracercion malayanum (Selys, 1876), Prodasineura doisuthepensis Hoess, 2007, Protosticta grandis Asahina, 1985, Tetracanthagyna waterhousei McLachlan, 1898, Gomphidia kruegeri Martin, 1904, Heliogomphus chaoi Karube, 2004, Leptogomphus baolocensis Karube, 2001, Microgomphus jurzitzai Karube, 2000, Onychothemis culminicola Förster, 1904, and two species for which the specimens collected on this trip were described as new subspecies: Indolestes gracilis expressior Kosterin, 2015, Coeliccia poungyi dasha Kosterin, 2016. Five species collected on this trip have been described elsewhere as new to science, namely Onychargia priydak Kosterin, 2015, Prodasineura hoffmanni Kosterin, 2015, Asiagomphus reinhardti Kosterin et Yokoi, 2016, Euthygomphus schorri Kosterin, 2016 and Risiophlebia guentheri Kosterin, 2015. So the total number of the first country records of named species made on this trip is 22. Still four species found may be undescribed. The number of named species recorded in Cambodia has reached 178. Remarks on taxonomy and variation of Euphaea masoni Selys, 1879, E. ochracea Selys, 1859, Aciagrion approximans (Selys, 1876), and Lamelligomphus castor Lieftinck, 1941 are provided. Characters of mature males of R. seducta are updated. Notes on habitats and habits of some species are provided. Onychothemis culminicola and O. testacea Laidlaw, 1902 seem to exclude each other at rivers, the former occupying smaller and more elevated ones; a putative hybrid male was observed. General notes on the area and field impressions are briefly outlined.
97
Coeliccia poungyi dasha subsp. nov. (Odonata, Platycnemididae, Calicnemiinae) from eastern Cambodia
(2016)
Coeliccia poungyi dasha is described from the Annamense Mts. in the eastern Cambodia (holotype: Cambodia, Mondulkiri Province, a brook, a left tributary of the main river downstream Buu Sraa Waterfall, 12°34'0119'' N 107°24'50''25' 03'' E, 416490 m a.s., 15.06. 2014, RMNH). The new subspecies differs from the nominotypical one in coloration of the mesepisternum in males and end of the abdomen in both sexes, as well as in the length of the terminal lobe of the genital ligula.
96
In 2014, 56 localities in four provinces of Lesser Himalaya in Pakistan were studied. A total of 28 species have been recorded. A female of the data deficient, threatened species Coeliccia vacca was recorded from Charhaan. The record of Drepanosticta carmichaeli is a new addition to the list of Odonata of Pakistan, and expand the range of this species further to the west. The taxonomical status of Ischnura aurora aurora – considered common in Pakistan, following baseline literature of Fraser (1933) – now turns out to be Ischnura aurora rubilio.
95
As a result of increased interest in dragonflies and close cooperation between odonatologists on the Balkan Peninsula, the Balkan Odonatological Meeting (BOOM) has been established in 2011. This report presents the results of the field trip during 4th Balkan Odonatological Meeting that was held in Bosnia and Herzegovina from 1st to 8th August 2014. With 69 surveyed localities and 496 records of 47 species, this was the most successful BOOM until now. The noteworthy results are records of several nationally rare species: Chalcolestes viridis, Coenagrion scitulum, Erythromma najas and Ceriagrion tenellum, and new populations of Cordulegaster heros, a species mentioned in the Annexes of the EU Habitats Directive. The distribution of Chalcolestes spp. in Bosnia and Herzegovina is also presented and discussed.
94
Distribution data of dragonflies and damselflies from western Bhutan collected during a trip from 10 October 2015 to 22 October 2015 are presented. In total 53 species were recorded of which eleven are new to the country (Aciagrion pallidum, Anisopleura lestoides, Megalestes irma, Gynacantha incisura, Gynacantha khasiaca, Gynacanthaeschna sikkima, Lamelligomphus risi, Somatochlora daviesi, Crocothemis erythraea, Sympetrum fonscolombii, Tholymis tillarga). Another three species, one Megalestes and two Cephalaeschna, were not identified to species level but are also addition to the list of species recorded from Bhutan.