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The myrmecophilous Paederinae rove beetle genus Megastilicus Casey, 1889 from North America is reviewed based on museum specimens. Prior to this study, the genus was monotypic with one species Megastilicus formicarius Casey, 1889 described. Here, we provide a redescription of the genus and the type species, designate a lectotype, and provide pictures of habitus and illustrations of the aedeagus and genital segments. Additionally, we describe a new species for the genus, Megastilicus iowaensis sp. nov., include an identification key to the two species and present the distribution map of both of them, including new state records. We discuss the assignment of the genus to the subtribe Stilicina based on morphological features.
Selizitapia gen. nov. (Hemiptera: Fulgoromorpha: Flatidae) from tapia woodlands of Madagascar
(2021)
A new monotypic genus of flatid planthoppers (Hemiptera: Fulgoromorpha: Flatidae), Selizitapia gen. nov., is described for Selizitapia pennyi gen. et sp. nov. (type species) from the island of Madagascar. Habitus, male and female external and internal genital structures of the new species are illustrated and compared with similar taxa. Selizitapia pennyi gen. et sp. nov. is endemic to Madagascar where it is known to date only from one locality in the Central Plateau and is associated with tapia woodland formation.
A new monotypic genus of flatid planthoppers (Hemiptera: Fulgoromorpha: Flatidae), Medleria gen. nov., is described for Medleria caudata gen. et sp. nov. (type species) from the island of Socotra (Yemen). Habitus, male and female external and internal genital structures of the new species are illustrated and compared with similar taxa. Medleria caudata gen. et sp. nov. is probably endemic to Socotra where it is known to date from a small area of the Dixam mountain plateau only.
Bujurquina is the most widely distributed and species-rich genus of cichlids in the western Amazon of South America. In this study we describe a new species from Peru from a hypothesized reverse flowing river system. Prior to the origin of the modern Amazon River at 4.5 Ma, this river system had its headwaters on the Iquitos arch, one of several main structural arches (swells) in the Amazon. Prior to the origin of the modern Amazon these arches formed topographic barriers of drainage basins in lowland Amazonia. For our analyses we use morphological and molecular data, analyzed through multivariate statistics and molecular phylogenies, respectivelly. For all valid species in the genus (except B. cordemadi and B. pardus) we additionally for the first time provide photographs of live specimens. Based on DNA phylogeny and coloration patterns we demonstrate that Bujurquina is divided into two main clades and based on this we provide a dichotomous key for all the species.
Three subterranean leptodirine leiodid taxa, viz., Bozidaria Ćurčić & Pavićević gen. nov., Bozidaria serbooccidentalis Ćurčić & Pavićević gen. et sp. nov. and Proleonhardella (Proleonhardella) tarensis Ćurčić & Pavićević sp. nov., are described and diagnosed. Bozidaria Ćurčić & Pavićević gen. nov. belongs to the phyletic series of “Leonhardella”. The new beetle taxa differ from their closest relatives in numerous morphological characters. They most likely belong to phyletic lineages of Pliocene age. The new leiodid taxa are endemic to the Dinaric mountain chain of western Serbia. Keys to the leptodirine leiodid genera of the phyletic series of “Leonhardella” and to the taxa of the genus Proleonhardella Jeannel, 1910 are included.
Plagiognathus ozgurkocaki sp. nov. is described based on a long series of specimens from Karaman, Turkey. The new species is remarkable among its congeners in Palearctic Region due to the combination of following characters: remarkably small size, dense and unicolorous pale yellow vestiture, darkened cuneus and yellow first antennal segment with a basal ring and pre-apical dots. The new species is associated with the endemic Phlomis leucophracta P.H.Davis & Hub.-Mor. (Lamiaceae) which makes it unique among all its congeners. Additionally, Plagiognathus bipunctatus albicans (Reuter, 1901) and Plagiognathus marivanensis Linnavuori, 2010 are recorded from Karaman, former constitutes a new record for Turkey.
Review of Synapsis Bates (Scarabaeidae: Scarabaeinae: Coprini), with description of a new species
(2010)
Presented are a checklist, a discussion of and keys to species groups and their constituent species, and a description of one new species: Synapsis horaki. The species Synapsis cambeforti Krikken and S. thoas Sharp are synonymized with S. ritsemae Lansberge, Balthasar’s synonymy of S. yunnana Arrow with S. tridens Sharp is revived, and the status of six recently described species is left unresolved because of insufficient data.
The spider genus Zaitunia Lehtinen, 1967 (Araneae, Filistatidae) is revised. It was found to include 24 species distributed in the Eastern Mediterranean, Middle East and Central Asia: ♀ Z. afghana (Roewer, 1962) (Afghanistan), ♀ Z. alexandri Brignoli, 1982 (Iran), ♀ Z. akhanii Marusik & Zamani, 2015 (Iran), ♂♀ Z. annulipes (Kulczyński, 1908) (Cyprus), ♂♀ Z. beshkentica (Andreeva & Tyshchenko, 1969) (Tajikistan, Uzbekistan), ♀ Z. brignoliana sp. nov. (Iran), ♂♀ Z. ferghanensis sp. nov. (Kyrgyzstan, Uzbekistan), ♀ Z. feti sp. nov. (Turkmenistan), ♀ Z. halepensis sp. nov. (Syria), ♀ Z. huberi sp. nov. (Afghanistan), ♀ Z. inderensis Ponomarev, 2005 (Kazakhstan), ♂♀ Z. kunti sp. nov. (Cyprus, Turkey), ♂♀ Z. logunovi sp. nov. (Kazakhstan, Kyrgyzstan), ♂♀ Z. maracandica (Charitonov, 1946) (Uzbekistan, Kazakhstan), ♂♀ Z. martynovae (Andreeva & Tyshchenko, 1969) (Tajikistan, Turkmenistan), ♀ Z. medica Brignoli, 1982 (Iran), ♂♀ Z. minoica sp. nov. (Greece), ♀ Z. minuta sp. nov. (Uzbekistan), ♀ Z. persica Brignoli, 1982 (Iran), ♂ Z. psammodroma sp. nov. (Turkmenistan), ♂♀ Z. schmitzi (Kulczyński, 1911), the type species (Egypt, Israel), ♂♀ Z. spinimana sp. nov. (Kazakhstan, Turkmenistan), ♂♀ Z. wunderlichi sp. nov. (Kyrgyzstan) and ♀ Z. zonsteini Fomichev & Marusik, 1969 (Kazakhstan). Twelve above-listed species are newly described, and males of Z. annulipes, Z. beshkentica, Z. maracandica and Z. martynovae are described for the first time. Two new combinations are established: Z. annulipes (Kulczyński, 1908) comb. nov., ex Filistata, and Pholcoides monticola (Spassky, 1941) comb. nov., ex Zaitunia. New data on distribution of the considered taxa are provided.
The genus Raveniola Zonstein, 1987 is found to be represented in Western Asia by 16 species: ♂♀ R. adjarica sp. nov. (Georgia), ♂ R. anadolu sp. nov. (Turkey), ♂ R. arthuri Kunt & Yağmur, 2010 (Turkey), ♂ R. birecikensis sp. nov. (Turkey), ♂♀ R. dunini sp. nov. (Armenia, Azerbaijan, Iran), ♂♀ R. hyrcanica Dunin, 1988 (Azerbaijan), ♂ R. marusiki sp. nov. (Iran), ♂ R. mazandaranica Marusik, Zamani & Mirshamsi, 2014 (Iran), ♂♀ R. micropa (Ausserer, 1871) (Turkey), ♀ R. nana sp. nov. (Turkey), ♂♀ R. niedermeyeri (Brignoli, 1972) (Iran), ♂♀ R. pontica (Spassky, 1937) (Russia, Georgia), ♀ R. sinani sp. nov. (Turkey), ♂♀ R. turcica sp. nov. (Turkey), ♂♀ R. vonwicki Zonstein, 2000 (Iran) and ♂♀ R. zaitzevi (Charitonov, 1948) (Azerbaijan, Georgia) = ♀ Brachythele recki Mcheidze, 1983, syn. nov. Eight species are described as new; others are redescribed from types and/or conspecific material. Males of R. micropa and R. zaitzevi, hitherto unknown, are described for the first time. Data on the variability, relationships, distribution and ecology of all considered species are also provided.
The Central Asian spider genus Anemesia Pocock, 1895 is rediagnosed and revised. The genus was found to contain 14 species: ♂♀ A. andreevae sp. nov. (Uzbekistan, Tajikistan); ♂♀ A. birulai (Spassky, 1937) (Turkmenistan); ♂♀ A. castanea sp. nov.; ♂♀ A. incana Zonstein, 2001, ♂♀ A. infumata sp. nov.; ♂♀ A. infuscata sp. nov.; ♂♀ A. karatauvi (Andreeva, 1968) (all Tajikistan); ♂ A. koponeni Marusik, Zamani & Mirshamsi, 2014 (Iran); ♂♀ A. oxiana sp. nov.; ♂♀ A. pallida sp. nov.; ♂ A. parvula sp. nov. (all Tajikistan); ♂♀ A. pococki sp. nov. (Turkmenistan); ♂♀ A. sogdiana sp. nov. (Uzbekistan, Tajikistan) and ♂♀ A. tubifex (Pocock, 1889), the type species (Afghanistan, Turkmenistan). Nine species are newly described; others are redescribed from types and/or conspecific material. Males of A. tubifex and females of A. birulai, hitherto unknown, are described for the first time. Data on the variability, relationships, distribution, and ecology of all considered species are provided.
In this paper, four new cave-dwelling species of the subgenus Tachycines (Gymnaeta) from Guangxi, Hunan and Jiangxi are described, i.e., Tachycines (Gymnaeta) quadratus sp. nov., Tachycines (Gymnaeta) shiziensis sp. nov., Tachycines (Gymnaeta) sparsispinus sp. nov. and Tachycines (Gymnaeta) papilious sp. nov. The female of Tachycines (Gymnaeta) liboensis Zhu, Chen & Shi, 2020 is recorded for the first time. Moreover, images illustrating the morphology of the four new and three previously described species are provided.
A new species complex, the eparmata complex, is established within the subgenus Phortica s. str., based on eight known and five new species, all of which are endemic to the Oriental Region: P. bipartita (Toda & Peng, 1992), P. eparmata (Okada, 1977), P. lanuginosa Chen & Toda, 2007, P. latipenis Chen & Gao, 2005, P. pangi Chen & Wen, 2005, P. setitabula Chen & Gao, 2005, P. unipetala Chen & Wen, 2005 and P. zeta Chen & Toda, 2007; P. jadete sp. nov., P. kava sp. nov., P. mengda sp. nov., P. wongding sp. nov. and P. yena sp. nov. A key to all species of this complex is provided. Barcoding sequences (mitochondrial COI gene) were obtained for 22 specimens of five known and the five abovementioned new species. The intra- and inter-specific pairwise K-2P (Kimura’s two-parameter) distances of COI were determined. Phylogenetic analysis was performed using Bayesian inference based on COI sequences, confirming the monophyletic status of the eparmata complex, which is distinct from the species complexes of magna, omega, variegata and another two ungrouped species.
New species of genus Atractides Koch, 1837 (Acari: Hydrachnidiae, Hygrobatidae) from Qinghai, China
(2022)
The paper deals with five new species of the genus Atractides Koch, 1837 (Acari: Hydrachnidiae, Hygrobatidae) collected from Qinghai Province, P.R. China, Atractides (Atractides) biprojectus Zhang, Li & Guo sp. nov., Atractides (Atractides) smiti Zhang, Li & Guo sp. nov., Atractides (Atractides) menyuanensis Zhang, Li & Guo sp. nov., Atractides (Atractides) longiprojectus Zhang, Li & Guo sp. nov. and Atractides (Atractides) xianmiensis Zhang, Li & Guo sp. nov. All the new species are described and illustrated in detail, and all the type specimens are deposited in the Institute of Entomology, Guizhou University, Guiyang, China (GUGC).
Four new species of the genus Coecobrya, C. gejianbangi sp. nov., C. annulata sp. nov., C. ciliata sp. nov., and C. oculata sp. nov., are described from Guangxi caves as the representative of the genus in China. Coecobrya oculata sp. nov. of the boneti-group has 1+1 eyes and a serrate outer edge of the unguiculus. The other three species, devoid of eyes and with a tiny outer tooth on the unguiculus, are assigned here to the tenebricosa-group, assuming that the large tooth on the unguiculus is transformed into a tiny one in cave-obligate species. Clypeal chaetae in Entomobryoidea are systematically surveyed for the first time, and are found to be well diversified at species level. They have a potential taxonomical value in discriminating taxa of morphologically conserved groups.
Conidera Rivalier, 1961 is a well-defined subgenus of tiger beetles (Coleoptera: Cicindelidae) hitherto only known from two species endemic to the island of Luzon. This paper describes the first species from Mindoro, with a doubtful record from Mindanao. Cylindera (Conidera) mindoroana Zettel and Wiesner, new species, differs strongly by an unusual shape of the female’s elytra. A diagnosis for Conidera and a key to the species of this subgenus is presented.
The Iranian species of the genus Anomalon Panzer, 1804 (Hymenoptera: Ichneumonidae: Anomaloninae) are reviewed. Four species, Anomalon amseli (Hedwig, 1961), A. chinense (Kokujev, 1915), A. cruentatum (Geoffroy, 1785) and A. narinae Zardouei & Rakhshani sp. nov., are found to occur in Iran. The female of A. amseli is described for the first time. Anomalon chinense is a new record for Iran. A key to the known Anomalon species of Iran is provided.
A new fossil diatom species, Tertiarius minutulus sp. nov., is described from a sediment sequence DEEP-5045-1 of Lake Ohrid. The species is characterized by small valves (3.0–8.0 μm) with a round shape, a marginal area with radially arranged costae and a central uneven area with scattered areolae. Externally, the alveoli are occluded by cribra perforated by irregularly arranged pores. Internally, the alveoli are simple and areolae are occluded with domed cribra. The marginal fultoportulae are situated on costae close to the valve margin, located on every 5th or 7th thick internal costa. One to three fultoportulae are present on the valve face, each surrounded by two to three satellite pores. One rimoportula is present, positioned on a costa at the valve face / mantle junction. The species is compared with morphologically similar taxa and a detailed differential diagnosis is provided. Tertiarius minutulus sp. nov. is known only as a fossil taxon from Lake Ohrid, with a first occurrence during the early stages of lake basin development.
Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae)
(2014)
The North American species of the genus Eusphalerum Kraatz (Coleoptera, Staphylinidae, Omaliinae) are revised. The taxonomic history, natural history, geographical distribution of the genus, characters, species groups, diversity, and biogeography of North American species are presented. Two main phylogenetic lineages and 13 species groups are provisionally recognized. The following new synonymies are proposed: Eusphalerum farrarae (Hatch, 1944) = E. lawrencei Hatch, 1957; Eusphalerum californicum (Fauvel, 1878) = E. atriventre (Casey, 1894), = E. nigerrimum (Casey, 1894), = E. gilvipenne (Casey, 1894), = E. dichroum (Fall, 1922), = E. bonnelli (Hatch, 1944), = E. lunae Hatch, 1957; Eusphalerum fraternum (Casey, 1894) = E. minskae (Hatch, 1944); Eusphalerum rugulosum (Mäklin, 1853) = E. grayae (Hatch, 1944); Eusphalerum orientale (Bernhauer, 1912) = E. frosti (Bernhauer, 1928). The following lectotypes are designated: E. subangulatum (Casey), E. californicum (Fauvel), E. gilvipenne (Casey), E. diversicolle (Casey), E. convexum (Fauvel), E. fraternum (Casey), E. horni (Fauvel), E. orientale (Bernhauer), E. pothos (Mannerheim), and E. punctatum (Casey). The following new species are described: Eusphalerum pilosum (California); E. klimaszewskii (British Columbia); E. chatzimanolisi (California); E. carolinensis (Kentucky, Missouri, North Carolina, Ohio, Pennsylvania, Tennessee, Virginia); E. caterinoi (California); E. luteipes (California); E. thayeranum (Alberta, British Columbia, Idaho, Indiana (doubtful record), Oregon, Washington); E. margaretae (Tennessee); E. newtoni (British Columbia, Oregon, California); E. parvispiculum (California, Oregon); E. uncinatum (British Columbia, California, Oregon, Washington). Eusphalerum lapponicum (Mannerheim, 1830) is excluded from the North American fauna. The following new combination is proposed: Xylodromus segmentarius (Mäklin, 1852: 322) (ex Omalium), wrongly attributed to Eusphalerum in the literature. Omalium marginatum Say, 1832 is considered a doubtful species, probably not Eusphalerum. A key to the 27 recognized North American Eusphalerum species and a catalog of the species are provided.
The taxonomic and faunistic status of the spider family Oecobiidae in Iran, Afghanistan and Turkmenistan is revised. A new species, namely Uroctea gambronica sp. nov. (♂) is described from southern Iran, and the male of U. grossa Roewer, 1960 is described and illustrated for the first time. Additionally, new faunistic data are provided, including the first records of Oecobius putus O. Pickard-Cambridge, 1876 and U. grossa in Afghanistan and Turkmenistan, respectively, and the re-evaluation of previously misidentified and questionable records of this family in the region. The known distribution ranges of all species are mapped for these three countries.
The taxa of the subgenus Turritus of Cochlostoma (Cochlostomatidae) are analysed based on molecular and morphological data. The phylogenetic trees, based on ribosomal (16S) and nuclear (H3) DNA, indicate that the currently accepted taxonomy should be revised. Based on our data, there are 37 species in Turritus of which 5 are new to science: Cochlostoma (Turritus) pallgergelyi sp. nov., C. (T. ) muranyii sp. nov., C. (T. ) hallgassi sp. nov., C. (T. ) kontschani sp. nov. and C. (T. ) lacazei sp. nov. Of these, we describe the shells and the female genitalia and summarize the distributional data. Some samples (or set of samples) will remain undetermined for lack of data and these are reported in the appendix.
Fourteen new species of the Colletes fasciatus species group are described, all of them endemic to the winter rainfall area in South Africa: C. ascopalis sp. nov. ♀, C. carolinae sp. nov. ♀♂, C. cedarbergensis sp. nov. ♀, C. fabiani sp. nov. ♀♂, C. fuscitergus sp. nov. ♂, C. khoisanorum sp. nov. ♀, C. kogelbergensis sp. nov. ♀♂, C. littoralis sp. nov. ♀, C. longitarsus sp. nov. ♂, C. peerboomi sp. nov. ♀, C. richtersveldensis sp. nov. ♀, C. ruschia sp. nov. ♀♂, C. spinipes sp. nov. ♂, C. troetroeensis sp. nov. ♀. Two species are synonymized based on newly recognized sex associations: C. katharinae Kuhlmann, 2007 syn. nov. is synonymized with C. infracognitus Cockerell, 1937 and C. bokkeveldi Kuhlmann, 2007 syn. nov. with C. zygophyllum Kuhlmann, 2007. The previously unknown female of C. inornatus Cockerell, 1946 is described for the first time and new records of already described species are added. All of the currently known 37 species of the C. fasciatus-group are imaged and included in a key to facilitate their identification.
Antiochtha is a genus of the family Lecithoceridae characterized by the forewing with veins M2 and M3 coincident, the hindwing with vein M2 absent, and the male genitalia with the valva narrowed from base to apex. The genus comprises 23 species and is mainly distributed in the Oriental Region. In this paper, we describe three new species: Antiochtha erromera sp. nov., A. hainanensis sp. nov. and A. miniscula sp. nov., and propose a new synonym, A. rotunda Zhu & Li, 2009 syn. nov., of A. jianfengensis Zhu & Li, 2009. Antiochtha semialis Park, 2002 is newly recorded in China and its female is described for the first time.
The endophallic structure of the genus Laius is studied and discussed based on the examination of 19 species from Asia to the Indian Ocean. The structure contains two primary sclerites (named gonoporal piece and ligula), a secondary sclerite on the basal part of the gonoporal piece (named additional sclerite) in some species, and a membranous basal area closely covered with many spines (named spinous area). Five species groups are recognized based on the morphology of the endophallic sclerites. The sympatric species have different body sizes and quite distinguishable endophallic sclerites (= different species group), while the allopatric species have overlapping body sizes and similar endophallic sclerites (= same species group). Three new species are described and six previously known species are redescribed with endophallic sclerites, and the descriptions of endophallic sclerites of the remaining ten species are added. The larva of Laius rodriguesensis sp. nov. is also described. The genus
Nossibeus Evers, 1994 is synonymised with Laius Guérin-Méneville, 1830.
The Asian species of the subfamily Thaumastodinae Champion, 1924 are reviewed. Seven new species are described: Acontosceles borneensis sp. nov., Pseudeucinetus papuanus sp. nov., Mexico ogasawaraensis sp. nov., M. baliensis sp. nov., M. papuanus sp. nov., M. palauensis sp. nov. and M. borneensis sp. nov. The genus Babalimnichus Satô, 1994 is treated as a junior synonym of the genus Mexico Spilman, 1972, and three known species of the genus Babalimnichus are transferred to Mexico, viz. M. taiwanus (Satô, 1994) comb. nov., M. masamii (Satô, 1994) comb. nov. and M. splendens (Hernando & Ribera, 2003) comb. nov. Additional specimen data are shown, and new distributional records are as follows: Acontosceles chujoi Yoshitomi & Satô, 2005 from Vietnam; A. zetteli Pütz, 2008 from Laos; Pseudeucinetus javanicus Yoshitomi & Putra, 2010 from Lombok Island; Mexico taiwanus (Satô, 1994) comb. nov. from Lutao, Lanhsu and the Yonaguni-jima Islands; and M. masamii (Satô, 1994) comb. nov. from Kume-jima. A species list of the subfamily Thaumastodinae is given, with ZooBank LSIDs. The phylogenetic relationships of the thaumastodine genera are discussed.
Four new species of the genus Caccothryptus (Coleoptera: Limnichidae) are described: C. taiwanus from Taiwan; C. orion from Okinawa; C. tibetanus and C. chayuensis from Tibet. All the species belong to the testudo species group (sensu Hernando & Ribera 2014). Additional specimen data and an updated species list are also given, and C. testudo Champion, 1923 is newly recorded from Thailand.
The Chinese fauna of the pselaphine genus Sathytes Westwood (Batrisitae: Batrisini) currently includes 20 species. In this paper, 15 new species from various provinces of the country are described: S. alpicola sp. nov. (Xizang), S. australis sp. nov. (Guangdong, Guangxi), S. chayuensis sp. nov. (Xizang), S. chengzhifeii sp. nov. (Yunnan), S. huapingensis sp. nov. (Guangxi), S. linzhiensis sp. nov. (Xizang), S. maoershanus sp. nov. (Guangxi), S. nujiangensis sp. nov. (Yunnan), S. panzhaohuii sp. nov. (Xizang), S. shennong sp. nov. (Hubei), S. tianquanus sp. nov. (Sichuan), S. transversus sp. nov. (Xizang), S. valentulus sp. nov. (Guangxi), S. xingdoumontis sp. nov. (Hubei) and S. xizangensis sp. nov. (Xizang). New collection records are provided for S. longitrabis Yin & Li, 2012, S. tangliangi Yin & Li, 2012 and S. yunnanicus Yin & Li, 2012. Maps showing the distribution of the genus in China, and an updated checklist of the world species are provided.
A review of the genus Stratiomys from India is presented. The new species Stratiomys brunettii sp. nov. is described based on male and female specimens collected from the Kashmir Himalayas. The only other congener previously recorded in India, Stratiomys approximata, is redescribed. A key to the species is presented.
Japanese species of the genus Georissus are revised. Seven species, including two new, G. (Neogeorissus) takahashii sp. nov. and G. (Ne.) satoi sp. nov. are recognized. Georissus (Neogeorissus) sakaii Satô, 1972 and G. (Ne.) katsuoi Nakane, 1995 are synonymized with G. (Ne.) japonicus Satô, 1972. Based on the type series and additional specimens, three subgenera and seven species are (re)described with SEM micrographs of the external morphology and figures of male and female genitalia. The terminology of some structures of the head and the pronotum are redefined for convenience of taxonomy. A key to the Japanese species is also given.
The species of Stenothemus Bourgeois, 1907 from Southwest China are reviewed. Three species are transferred to this genus, S. davidi (Pic, 1926) comb. nov. (from Malthacus Kirby, 1837), S. minutissimus (Pic, 1933) comb. nov. (from Micropodabrus Pic, 1920) and S. singularicollis (Pic, 1933) comb. nov. (from Lycocerus Gorham, 1895). Two new synonyms are suggested, S. dinshuiensis Švihla, 2011 syn. nov. is synonymized with S. davidi and S. melleus Švihla, 2005 syn. nov. with S. singularicollis. Stenothemus shaanxiensis Švihla, 2004 stat. nov. is raised to species level from a subspecies of S. benesi Švihla, 2004. Five new species are described, S. gracilis Y. Yang & X. Yang sp. nov., S. chongqingensis Y. Yang & H. Liu sp. nov., S. flavicollis Y. Yang & S. Ge sp. nov., S. jindraimimus Y. Yang & X. Yang sp. nov. and S. laticornis Y. Yang & H. Liu sp. nov., which are illustrated with photographs of habitus, aedeagi, abdominal sternites VIII and internal genitalia of the female. Characters of the female reproductive system are described for the first time for the following species: S. diffusus Wittmer, 1974, S. dundai Švihla, 2004, S. grahami Wittmer, 1974, S. jindrai Švihla, 2004, S. tryznai Švihla, 2004 and S. singularicollis. Some additional distribution information is added for previously known species. A key for the identification of all studied species is provided.
The species of Stenothemus Bourgeois, 1907 from Southeast China are reviewed. Stenothemus fukienensis Wittmer, 1974 and S. kuatunensis Wittmer, 1979 are supplementarily described. Two new species are described, S. longicornis Y. Yang & H. Liu sp. nov. (China: Guangdong) and S. flavus Y. Yang & X. Yang sp. nov. (China: Zhejiang). Five new combinations are established: S. biimpressiceps (Pic, 1930) comb. nov. (from Cantharis L.), S. chinensis (Wittmer, 1982) comb. nov., S. limbatipennis (Pic, 1926) stat. rev. et comb. nov., S. nigriceps (Wittmer, 1955) comb. nov. and S. pallicolor (Wittmer, 1951) comb. nov. (from Lycocerus Gorham). Leiothorax atrosanguineus Švihla, 2005 syn. nov. is synonymized with S. chinensis, Lycocerus limatus Kazantsev, 2007 syn. nov. with S. limbatipennis. The above species are illustrated with habitus photos, aedeagi, abdominal sternites VIII and internal genitalia of female. A key for the identification of the above species is provided.
Twelve species of the subgenus Vestiplex (Diptera, Tipulidae) were previously known to occur in Tibet (= Xizang), China. Here, six species are described and illustrated as new to science: Tipula (Vestiplex) bucera sp. nov., Tipula (Vestiplex) magatama sp. nov., Tipula (Vestiplex) motuoensis sp. nov., Tipula (Vestiplex) nayogabuensis sp. nov., Tipula (Vestiplex) platyphylla sp. nov. and Tipula (Vestiplex) uncinella sp. nov. The following three species are redescribed: Tipula (Vestiplex) himalayensis Brunetti, 1911, Tipula (Vestiplex) nigroapicalis Brunetti, 1911 and Tipula (Vestiplex) zayulensis Alexander, 1963. A key to the species of Tipula (Vestiplex) from Tibet is presented.
Three novel species collected from Dinghushan Biosphere Reserve (DHSBR) in southern China, Russula cylindrica Y.Song sp. nov. and R. lacteocarpa Y.Song sp. nov. in subgenus Archaeae and R. reticulofolia Y.Song sp. nov. in subg. Compactae, are described based on morphological and molecular data. In addition, Russula leucobrunnea Y.Song nom. nov. is proposed in replacement of R. leucocarpa nom. illeg. in subg. Brevipedum, as R. leucocarpa (T.Lebel) T.Lebel had been described earlier. Differences between the three novel species and their closely related taxa were analyzed. Another two known species in subg. Brevipedum, R. callainomarginis J.F.Liang & J.Song and R. japonica Hongo were also identified among specimens from DHSBR and are described and illustrated. Phylogenetic analyses of ITS and a five-locus phylogeny (concatenated LSU, mtSSU, rpb1, rpb2 and tef1) support the recognition of these taxa.
Three species of the genus Cryptochetum Rondani, 1875 from China are described and figured as new to science: C. euthyiproboscise sp. nov., C. glochidiatusum sp. nov., and C. longilingum sp. nov. An identification key to the known species of Cryptochetum from China is presented. The type specimens of the new species are deposited in the Henan Agricultural University.
Two new genera and species of tiger beetles from Baltic amber (Coleoptera: Carabidae: Cicindelinae)
(2017)
Two fossil tiger beetle species (Coleoptera, Carabidae, Cicindelinae) are described from Eocene Baltic amber using light microscopic and X-ray microscopic techniques. Both species are considered representatives of the subtribe Iresina Rivalier, 1971 due to the shared combination of character states: glabrous head, six labral and four suborbital setae, and glabrous pronotum. Palaeopronyssiformia groehni Wiesner, Will, and Schmidt, new genus, new species, is characterized by a glabrous and furrowed head with six labral setae, large eyes, presence of two supraorbital setae on each side, mandibles with two teeth of the incisor region, and a glabrous and furrowed pronotum. Palaeoiresina cassolai Wiesner, Will, and Schmidt, new genus, new species, is characterized by a unicolored, undentated labrum, mandibles with two teeth of the incisor region, glabrous head with six labral setae, two clypeal setae, two supraorbital setae on each side, and a glabrous pronotum, mesepisternum, mesepimeron, and metepisternum. The species described here represent the only known tiger beetle fossils preserved in Baltic amber.
Prothyma (Genoprothyma) thandamoeae Wiesner, Phyu and Hori, new species, Prothyma (Genoprothyma) sotai Wiesner, Phyu and Hori, new species, and Prothyma (Genoprothyma) asamii Wiesner, Phyu and Hori, new species (Coleoptera: Cicindelidae) are described from Myanmar. A key to all members of the genus known to occur in Myanmar is given.
Cylindera (Cylinderina) sierramadrensis Wiesner and Dheurle, new species (Coleoptera: Cicindelidae) is described from Luzon. Re-descriptions of the other seven species of this subgenus (C. (C.) nanula (Horn, 1937), C. (C.) nana (Schaum, 1862), C. (C.) pseudonana (Horn, 1924), C. (C.) werneri Wiesner, 1988, C. (C.) rothschildi (Horn, 1896), C. (C.) genieri Cassola and Werner, 2003, C. (C.) vandenberghei Dheurle, 2016) are provided together with a key and pictures of their habitus, labrum and aedeagus.
A key is provided for the twelve taxonomic groups within the genus Therates Latreille (Coleoptera: Cicindelidae). The chennelli group is reviewed here and a key to the 53 species known for this group is provided. Each species is illustrated, and a detailed description of its morphology and distributional records are provided. Therates khaoyaii, Th. pearsoni, Th. safraneki, Th. schuelei and Th. sigridgeissleri are described as new to science. Therates differens Sawada and Wiesner, 1999 is placed into synonymy under Th. concinnus Gestro, 1888. Therates tonkinensis kubani Wiesner, 1988 is placed into synonymy under Th. tonkinensis Horn, 1902 and Th. belokobylskiyi Matalin and Wiesner, 2006 is placed into synonymy under Th. haucki Moravec and Wiesner, 2001. Lectotypes are designated for Th. waagenorum Horn, 1900, Th. clavicornis Horn, 1902, Th. tonkinensis Horn, 1902 and Th. annandalei Horn, 1908. Therates baolocensis Wiesner, 1996 is elevated to species rank.
Examination of leucothoid amphipods of the Red Sea has revealed seven species not previously reported from this location. Leucothoe minoculis sp. nov., Leucothoe pansa sp. nov., Leucothoe reimeri sp. nov., and Paranamixis sommelieri sp. nov. are described and the range of Leucothoe predenticulata Ledoyer, 1978, L. acutilobata Ledoyer, 1978 and L. squalidens Ledoyer, 1978 is expanded to include the Red Sea. Clarification of reports of L. acanthopus Schellenberg, 1928 and L. bannwarthi (Schellenberg, 1928) is provided and Leucothoe alani sp. nov. is described from outside the Red Sea.
The concept of the jumping spider genus Pochytoides Berland & Millot, 1941 is reviewed, based on the examination of described and undescribed species. Pochytoides is elevated from the subgeneric to the generic rank and a short diagnosis and description of the genus are presented. Redescriptions or descriptions of all species are provided together with a key to the species. Two new combinations are proposed: Pochytoides perezi (Berland & Millot, 1941) comb. nov. and P. poissoni (Berland & Millot, 1941) comb. nov. (both from Pochyta). Pochyta remyi Berland & Millot, 1941 originally placed in the subgenus Pochytoides is excluded; new combination Thiratoscirtus remyi (Berland & Millot, 1941) comb. nov. is proposed for it (but its generic status is uncertain). Six new species are described: Pochytoides monticola sp. nov., P. obstipa sp. nov., P. lamottei sp. nov., P. patellaris sp. nov., P. securis sp. nov. and P. spiniger sp. nov. The genus has a West African distribution.
Seven new species of the giant pill-millipede genus Zoosphaerium Pocock, 1895 are described from Madagascar: Z. nigrum sp. nov., Z. silens sp. nov., Z. ambatovaky sp. nov., Z. beanka sp. nov., Z. voahangy sp. nov., Z. masoala sp. nov. and Z. spinopiligerum sp. nov. All species are described based on drawings and scanning electron microscopy, while genetic barcoding of the COI gene was successful for six of the seven new species. Additional COI barcode information is provided for the first time for Z. album Wesener, 2009 and Z. libidinosum (de Saussure & Zehntner, 1897). Zoosphaerium nigrum sp. nov. and Z. silens sp. nov. belong to the Z. libidinosum species-group, Z. ambatovaky sp. nov. to the Z. coquerelianum species-group, Z. beanka sp. nov., Z. voahangy sp. nov. and Z. masoala sp. nov. to the Z. platylabum species-group and Z. spinopiligerum sp. nov. to the Z. piligerum species-group. Updated identification keys are provided for each species-group. Two western dry forest species, Z. silens sp. nov. and Z. voahangy sp. nov. are recorded from two localities, while the other five species are currently only known from their type localities. Of special conservation concern might be Z. ambatovaky sp. nov. from the lowland rainforest fragment of Ambatovaky, a nowadays isolated lowland rainforest, rapidly shrinking due to slash and burn agriculture. In addition to the new species, new locality data is provided for 11 species and numerous unidentifiable species of Zoosphaerium: Z. neptunus (Butler, 1872), Z. platylabum (de Saussure & Zehntner, 1902) and Z. piligerum (de Saussure & Zehntner, 1897) from the central eastern montane forests, as well as Z. ambrense Wesener, 2009, Z. aureum Wesener, 2009, Z. libidinosum, Z. corystoides Wesener, 2009, Z. discolor Wesener, 2009, Z. smaragdinum Wesener, 2009, Z. villosum Wesener & Sierwald, 2005 and Microsphaerotherium anjozorobe Wesener, 2009.
The Bolivian species of Polyrhaphis Audinet-Serville, 1835, (Coleoptera, Cerambycidae, Lamiinae) are reviewed and illustrated, with P. skillmani new species described. A key is presented to the six species recorded from Bolivia (P. angustata Buquet, 1853; P. argentina Lane, 1978; P. gracilis Bates, 1862; P. pilosa Lane, 1965; P. spinosa (Drury, 1773); and P. skillmani Wappes and Santos-Silva, new species). Their collection localities, based on recently identified specimens examined by the authors, are plotted to show the distribution of species, and displayed next to an ecoregion map of Bolivia to illustrate biogeographical information for Polyrhaphis.
New species and taxonomical notes in Gorybia Pascoe, 1866 (Coleoptera: Cerambycidae: Cerambycinae)
(2019)
Three new Gorybia Pascoe, 1866 (Coleoptera: Cerambycidae: Cerambycinae: Piezocerini), species from Bolivia are described: G. martinsi Wappes, Botero and Santos-Silva new species; G. galileoae Wappes, Botero and Santos-Silva, new species; and G. clarkeorum Wappes, Botero and Santos-Silva, new species. In addition, G. bispinosa Martins, Galileo and Limeira-de-Oliveira, 2009 is proposed as a synonym of G. castanea (Gounelle, 1909) and G. maculosa Martins, 1976 as a synonym of G. apatheia Martins, 1976.
Serratacosa, a new genus of Lycosidae (Araneae) from the southern slopes of the Eastern Himalayas
(2021)
A new wolf spider genus, Serratacosa gen. nov., is erected based on the type and new species, S. medogensis gen. et sp. nov. and two newly combined species, S. himalayensis (Gravely, 1924) (from Hogna Simon, 1885) and S. multidontata (Qu, Peng & Yin, 2010) (from Pardosa C.L. Koch, 1847). All of them are from the southern slopes of the Eastern Himalayas. Descriptions of the new genera and species, and a redescription of S. multidontata are provided together with digital images, illustrations and a distribution map.
Six species of the spider genus Spinirta Jin & Zhang, 2020 from southern China (Araneae: Corinnidae)
(2024)
In the current study, six species of the genus Spinirta from southern China are recognized, four new species are described: S. hongyui Wang, Lu & Z.S. Zhang sp. nov. (♂♀), S. liuae Wang, Lu & Z.S. Zhang sp. nov. (♂♀), S. simianshan Wang, Lu & Z.S. Zhang sp. nov. (♂♀) and S. yintiaoling Wang, Lu & Z.S. Zhang sp. nov. (♂). Females of S. aviforma Jin & Zhang, 2020 (♂♀) and S. quadrata Jin & Zhang, 2020 (♂♀) are described for the first time. Photos of the body and copulatory organs, as well as the locality map are provided.
Neopanorpa van der Weele, 1909 is the second largest genus in Panorpidae, and over 170 Oriental species have been reported hitherto. In this paper, we describe two new species: Neopanorpa luojishana sp. nov. and Neopanorpa xingmini sp. nov. from Southwest China. We also report the first discovery of the male of Neopanorpa ocellaris (Navás, 1908) from Guangxi and Guizhou, China, a species recorded from Sikkim previously. Neopanorpa brevivalvae Chou & Wang, 1988 is synonymized with Neopanorpa lungtaushana Cheng, 1957. In addition, Neopanorpa furcula nom. nov. is proposed for Neopanorpa furcata Zhou, 2005, a junior homonym preoccupied by Neopanorpa furcata (Hardwicke, 1825). The biogeographical implications of some Neopanorpa species are discussed in brief.
Three species of the genus Prorops Waterson, 1923 occur in Madagascar. Prorops nasuta Waterson, 1923 is recorded for the first time from Madagascar and two new species are described and illustrated: P. sparsa sp. nov. and P. impotens sp. nov., both based on the morphology of males and females. A brief discussion of the status of the genus, illustrations, and a key to Madagascan species of Prorops are provided.
Review of the Panorpa wormaldi group (Mecoptera: Panorpidae), with descriptions of two new species
(2022)
Panorpa Linnaeus, 1758 is the largest genus in the scorpionfly family Panorpidae. In this paper, a taxonomic review of the Panorpa wormaldi group is provided, with two new species described from China: Panorpa fengyanga Wang & Suzuki, sp. nov. from Zhejiang, and Panorpa zhuohengi Wang & Suzuki, sp. nov. from Guangdong. The male of Panorpa implicata Cheng, 1957 is discovered and described for the first time. A distributional map and keys to species are also provided for this group. Species number in this group is updated from 17 to 19. In addition, their biogeographical and evolutionary implications are briefly discussed.
The justification of the 4 genera that currently compose the class Pavlovophyceae is based on a low number of species and a relative paucity of available, traceable and referenced cultures. Previous integrative phylogeny work revealed strains that can refine and strengthen our knowledge of the genera in the class. The application of multiple light and electron microscopy techniques allowed us to prioritize the cytomorphological characters (pyrenoid, thylakoid, stigma, knob-scales, life stage / life cycle) used for the taxonomy of these algae and to describe two new species: Exanthemachrysis fresneliae Véron sp. nov. and Rebecca billardiae Véron sp. nov. Consequently, revisions of the two genera Exanthemachrysis Lepailleur emend. Véron and Rebecca Green emend. Véron were made. In addition, the genus Pavlova Butcher emend Véron is revised in the light of these characters. Particular emphasis is placed on the life stages and habitat of the species.
Revision of the Merodon bombiformis group (Diptera: Syrphidae) – rare and endemic African hoverflies
(2021)
In the present work, the Afrotropical species of the bombiformis species group, part of the aureus lineage, are revised. Six species are recognized, based on a combination of morphological and genetic features. Three of these species are new to science: Merodon lotus Vujić & Radenković sp. nov., M. vittatus Vujić & Likov sp. nov., and M. zebra Vujić & Radenković sp. nov. Redescriptions are provided for the other three species: M. bombiformis Hull, 1944, M. multifasciatus Curran, 1939, and M. nasicus Bezzi, 1915. The female of M. bombiformis is described. The name Merodon edentulus Macquart, 1855 is considered here as a nomen dubium. One new synonymy is proposed: M. apimima Hull, 1944 syn. nov. (junior synonym of M. multifasciatus). The distribution of the bombiformis species group is discussed. The larval host plant of M. multifasciatus is identified as Gladiolus. A key to the identification of both males and females of the bombiformis group is provided.
Coelogynoporidae (Platyhelminthes) includes comparatively large and slender Proseriata, usually occurring in shallow benthic environments. Coelogynopora Steinböck, 1924 is the most frequently reported genus and the one with the highest species diversity. Notwithstanding that, the genus has never been reported from the Southern Hemisphere. A recent analysis of sediment from the Magellan Strait shores (Chile) resulted in the discovery of a new species of Coelogynopora, the first representative of the genus to be found in austral waters. The new species is defined by the following combination of characters: sclerotised copulatory system consisting of a slender, ventrally curved stylet with a broad base and three pairs of symmetrically arranged spines, the proximal ends of which are fused laterally to the base of the stylet; distal ends of the three pairs of spines hooked, with apophyses at progressively longer distances from the tip; accessory spines and solar organ absent. Based on the morphological characters, the new species appears to be more related to species from the Pacific Ocean than to those from the Atlantic Ocean. The present work suggests a vast biogeographic disjunction in the genus Coelogynopora, which may be described as a bipolar or amphitropical pattern of distribution.
In February 2012, Odonata were recorded and voucher specimens collected in Luzon, The Philippines. The focus of study was set on localities near Dinapigue and San Mariano (Isabela Province), sites in Casiguran (Aurora Province) and on Polillo Island (Quezon Province). 60 Odonata species were recorded. Three are new to science and have been formally to be described. Four species were recorded for the first time in Luzon. Amphicnemis furcata and Diplacodes nebulosa were rediscovered after several decades since they were last documented from Luzon.
Notes on a small Odonata collection from Tawi-Tawi, Sanga-Sanga and Jolo islands, Philippines
(2012)
Sulu region is among the least explored faunal region in the Philippine archipelago. Odonatologically, this region is poorly studied until recently. Presently a survey conducted in July 1 – 14, 2011 revealed ten new records in Tawi-Tawi raising the total number of Odonata to 54. Three new species records were made for Sanga-Sanga raising the known number in that island to 34. Three species were recorded for the first time in Jolo raising the total number to 18. One new species of damselfly was found and several questionable and possible new species of dragonflies were documented.
The Odonata fauna of Balabac Island, Philippines was studied in March 2013. A total of 41 species under 33 genera were recorded. Twenty-eight species were recorded for the first time in the island. One genus – Mortonagrion was recorded for the first time in the Philippines. Mortonagrion astamii spec. nov. and Prodasineura poncei spec. nov. are new to science and are described. Three previously recorded species remained elusive and not seen during the survey.
Odonata survey was conducted in Talaingod, Davao del Norte, Mindanao Island. Four major sites were explored in Barangay Santo Niño from December 26 – 30, 2012. Thirty five species under eleven families including one new species were found representing the first odonatological record in the province of Davao del Norte. Three species need further study while Orthetrum glaucum represents a new record for the island of Mindanao. Coeliccia exoleta population, a vulnerable species in the IUCN Red List of Threatened Species, was found.
Odonata were recorded and voucher specimens were collected between August 3- 13 and August 16-20, 2011. This account lists 51 species of Odonata for both islands, 47 species in Siargao and 24 species in Bucas Grande. Thirty seven species are new island records for Siargao Island while the 24 species recorded in Bucas Grande represent the first island records. Three species are new to science, and two of which (Drepanosticta schorri n. sp., Pseudagrion schieli n. sp.) are described in the present paper. Four species previously listed (Hämäläinen & Müller, 1997) remained elusive during the present survey.
Odonata fauna of Diomabok Lake and its surroundings, Davao Oriental, Mindanao Island, Philippines
(2011)
During three visits in October and December 2010 and May 2011, a total of 56 Odonata species was recorded. All species reported here represent first Odonata records in the area. The most noteworthy discoveries were one novelty (Hydrobasileus vittatus) to the Philippine fauna and two first records (Tetracanthagyna brunnea and Aethriamanta gracilis) from Mindanao Island. Seven species represent either new species to science or potentially new species; one Drepanosticta and one Amphicnemis are new to science, and another Drepanosticta, Amphicnemis, Pseudagrion, Gomphidia and Urothemis are potentially new to science.
Four taxa belonging to the complex of species around Achnanthidium minutissimum were found during the ongoing taxonomic revision of the Antarctic freshwater and limno-terrestrial diatom flora. Two taxa were previously described as Achnanthidium lailae and A. sieminskae. Two others were formerly identified as A. minutissimum but detailed light and scanning electron microscopical observations revealed sufficient morphological differences compared to the type of A. minutissimum, to justify their separation and description as new taxa: Achnanthidium indistinctum and A. maritimo-antarcticum. The morphology and ecology of all four taxa are discussed comparing the species with morphologically similar taxa. The biogeographical consequences of the splitting of the former A. minutissimum complex in the Antarctic Region are discussed.
The adult stage of Helioandesia tarregai gen. et sp. nov. (Lepidoptera: Yponomeutoidea: Heliodinidae) is described and illustrated from the arid western slopes of the Andes of northern Chile. The larvae of H. tarregai gen. et sp. nov. feed as leaf skeletonizers on Mirabilis acuta (Reiche) Heimerl (Nyctaginaceae). The mostly gray forewing of H. tarregai gen. et sp. nov., ornamented with strongly bulging metallic spots, resembles that of the representatives of the mainly Nearctic Lithariapteryx Chambers, 1876. However, the latter lacks CuP in the forewing, has a single bristle in the female frenulum, and lacks a well-developed cornutus. Helioandesia gen. nov. clustered as sister to Neoheliodines Hsu, 2004 in a cladistic analysis, although no synapomorphies were found for this cluster, while Lithariapteryx was sister to Helioandesia gen. nov. + Neoheliodines based on two synapomorphies. The genetic distance between a DNA barcode sequence of H. tarregai gen. et sp. nov. and representatives of other genera of Heliodinidae Heinemann, 1877 was 9.0–12.5% (K2P), and a maximum likelihood analysis based on this molecular marker confirmed the placement of H. tarregai gen. et sp. nov. as a member of this micromoth family. This contribution represents the first confirmed record of Heliodinidae for Chile.
Two Mastogloia Thwaites ex W.Sm. taxa were found during a survey of the diatom flora of Lac de Guiers, Senegal. Based on all currently available literature, one taxon could be identified as M. belaensis M.Voigt, a species formerly described from Pakistan. The second species showed some resemblance to M. braunii Grunow. Analysis of the type of M. braunii revealed, however, important morphologic differences, leading to the description of a new species from the Senegal population: M. senegalensis Van de Vijver, Fofana, Sow & Ector sp. nov. The present paper describes this new species and discusses and illustrates the morphology of M. belaensis and the type of M. braunii. All taxa are discussed with morphologically similar taxa.
New species of “giant” plume moths of the genus Platyptilia (Lepidoptera, Pterophoridae) from Uganda
(2016)
This paper describes two new species of plume moths from the group of the so-called “giant” Platyptilia Hubner, 1825: Platyptilia fletcheri Ustjuzhanin & Kovtunovich sp. nov. and P. stanleyi Ustjuzhanin & Kovtunovich sp. nov. Both species were collected in the Rwenzori Mountains in Uganda and Rwanda, respectively. Platyptilia stanleyi Ustjuzhanin & Kovtunovich sp. nov. exceeds all the known African species of Pterophoridae in its wingspan of 49 mm.
Two new species of aulacid wasps, Pristaulacus iuliae Turrisi & Nobile sp. nov. from South India (Karnataka) and Pristaulacus ninae sp. nov. from Peninsular Malaysia (Pahang), are described, figured and compared with most related species. Based on the present addition, the Indomalayan area currently includes 27 species of Pristaulacus Kieffer out of 61 known from the whole Oriental Region, which is however a largely underestimated number. An identification key to species and an updated checklist of Pristaulacus occurring in the Indomalayan area are provided.
A new species of the genus Mesobiotus is described from the Republic of South Africa using a traditional morphological approach (light and scanning electron microscopy) combined with molecular analysis (18S rRNA, 28S rRNA, ITS-2 and COI markers). Mesobiotus anastasiae sp. nov. differs from all known Mesobiotus species by having a unique combination of characters of the adult animals and the eggs. Adults of the new species have an oral cavity armature without elongate teeth in the second band, while the processes of the egg chorion have a basal collar and distinct rows of large pores. An updated key to the species of the genus Mesobiotus, including 66 of 70 currently described species, is given. An aquatic mite species from the Lobohalacarus weberi complex (freshwater Halacaridae) co-occurs with M. anastasiae sp. nov., suggesting that the newly described tardigrade inhabits constantly wet moss cushion habitats.
A new genus with a new species of soft-winged flower beetle, Pectotibialis paghmanensis Tshernyshev gen. et sp. nov. are described from Afghanistan. The new genus can be distinguished from the congeners of the tribe Apalochrini by the dark pectination in the apices of tibiae in both sexes, and the anterior tibiae which are hollowed at distal half, flattened and rounded femora, bituberculate basal parts of head and pronotum, two lamellate metathoracic appendages, tarsal comb above second tarsomere of anterior legs, and serrate antennae in the male. Based on the metathoracic appendages and comb in anterior legs would attribute this new species to the new genus Dromanthomorphus Pic, 1921, but all the other above-mentioned characters define its independent status and the designation of a new genus; Pectotibialis Tshernyshev gen. nov. The external appearance, special male characters and genitalia of the type species of the new genus are illustrated, and a distribution map is provided. A key to the Apalochrus-section of the tribe Apalochrini is provided.
Soft-winged flower beetles from the United Arab Emirates are studied for the first time. Three new genera, Tonyattalus gen. nov. of the tribe Attalini, Tonycolotes gen. nov. of the tribe Colotini, and Arabotroglops gen. nov. of the tribe Troglopini, and two species, Tonyattalus vanharteni gen. et sp. nov. and Arabotroglops longantennatus gen. et sp. nov., are described. Tonycolotes kovari (Švihla, 1987) gen. et. comb. nov. is recorded from the United Arab Emirates for the first time and transferred from the genus Colotes Erichson, 1840. All species discussed in the paper are described and illustrated. The fauna of soft-winged flower beetles of the Arabian Peninsula is reviewed.
Palaeosynthemis opaca sp. nov., a new dragonfly from Papua New Guinea (Anisoptera: Synthemistidae)
(2016)
A new species of the synthemistid genus Palaeosynthemis is described from the northern slopes of Papua New Guinea’s central cordillera. It is distinguished from all congeners by having females with large, nearly black patches at the bases of both wings. The male is most similar to P. cervula and P. feronia, but it differs from those species in having superior anal appendages less than three times as long as S10, basally enlarged and otherwise unarmed vs basally not enlarged (feronia) and almost four times as long as S10 and armed (cervula). Characters of the adults (both sexes) are illustrated and the affinities of the species are discussed.
A new species of the genus Teinobasis Kirby is described from the Muller Range in Western Province, Papua New Guinea. Its male is distinguished from all other Teinobasis species by having a pale labrum, an extensively bright orange thorax, and ventrally bowed superior anal appendages that are markedly shorter than the plump, apically rounded inferiors. Characters of the male are illustrated, and the affinities of the new species are discussed.
Two new species of damselflies are described from central Halmahera in North Maluku Province, Indonesia. They are Drepanosticta pararudicula sp. nov. (Holotype MZB. ODON. 19257) and Nososticta halmahera sp. nov. (Holotype MZB. ODON. 19265). The two species are most similar to the Moluccan taxa D. rudicula and N. moluccensis respectively and their descriptions bring the total number of Drepanosticta species known from Halmahera to five and of Nososticta to two.
Eight new species of the plant bug genus Coridromius are described: C. basilanus sp. nov. from the Philippines, C. eremnos sp. nov. from Sabah, Malaysia, C. fomangsu sp. nov. and C. tafo sp. nov. from Ghana, C. norfolkensis sp. nov. from Norfolk Island, Australia, C. mulu sp. nov. from Sarawak, Malaysia, C. macchabeeus sp. nov. from Mauritius, and C. taravao sp. nov. from Tahiti, French Polynesia.
The species of the ant genus Strumigenys Smith, 1860 from Southeast Asia are reviewed based on recent sampling efforts as well as unreported historical material from southern mainland China, Japan, Taiwan, Thailand and Vietnam. We report 42 new species records for these regions. A total of 20 species new to science are described: S. anhdaoae sp. nov., S. claviseta sp. nov., S. crinigera sp. nov., S. decumbens sp. nov., S. delicata sp. nov., S. densissima sp. nov., S. doydeei sp. nov., S. fellowesi sp. nov., S. intermedia sp. nov., S. jaitrongi sp. nov., S. liuweii sp. nov., S. longidens sp. nov., S. mediocris sp. nov., S. rongi sp. nov., S. scutica sp. nov., S. strummeri sp. nov., S. xenopilus sp. nov., S. yamanei sp. nov., S. zanderi sp. nov. and S. zhenghuii sp. nov. The descriptions of the existing species S. elegantula (Terayama & Kubota, 1989) and S. nathistorisoc Tang et al., 2019 are revised. Strumigenys formosensis Forel, 1912 syn. nov. is synonymized with S. feae Emery 1895 in the light of recently collected specimens. Three species complexes are created within the S. leptothrix-group: elegantula-complex, leptothrix-complex and zanderi-complex, based on differences in dentition. A new species group, S. nathistorisoc-group, is introduced. The key to Strumigenys of East Asia (as Pyramica) by Bolton is partially revised to accommodate species from these species groups that were described since the publication of the key. Finally, the implication of our results to the region and the current limitation of species groups and complexes of the genus is discussed.
The bathyal kinorhynch fauna along the Northwest American continental rise is explored, with emphasis on species of Echinoderidae Zelinka, 1894. Seven species of Echinoderes Claparède, 1863 are described as new to science: E. anniae sp. nov., E. dubiosus sp. nov., E. hamiltonorum sp. nov., E. hviidarum sp. nov., E. juliae sp. nov., E. lupherorum sp. nov. and E. yamasakii sp. nov. Three known species, Echinoderes hakaiensis Herranz, Yangel & Leander, 2017, E. cf. unispinosus Yamasaki, Neuhaus & George, 2018 and Fissuroderes higginsi Neuhaus & Blasche, 2006, are reported. The numerous new species indicate that the deep-sea still holds a great, undiscovered diversity of kinorhynchs, and that Echinoderes, as is also the case in more shallow, coastal waters, represents an important component of the deep-sea kinorhynch fauna. The presence of E. hakaiensis in the deepsea sediments demonstrates that the species may occur at a great depth range, and suggests that depth may play a less important role for the distribution of some kinorhynch species. The finding of the Northeast Atlantic species E. cf. unispinosus and the Southwest Pacific species Fissuroderes higginsi could indicate that kinorhynch species in the deep-sea may cover considerably larger distributional ranges than is assumed for coastal species of Echinoderidae.
En este estudio se describe una nueva especie de mosca blanca Aleuropleurocelus chamaedoreaelegans Sánchez-Flores y Carapia-Ruiz sp. nov. (Hemiptera: Aleyrodidae) encontrada en el envés de las hojas de Chamaedorea elegans Mart. en el Estado de Puebla (México). Se proporcionan microfotografías de estructuras morfológicas de puparios, se discute la separación con otras especies cercanas de Aleuropleurocelus Drews y Sampson, y se propone una clave para especies del grupo de forma oval.
Examination of Coccinellidae material recently collected from Napo Province in Ecuador revealed 38 species of ladybird beetles, of which six are new: Chnoodes yanayacu sp. nov., Cyrea mcclarini sp. nov., Eupalea borowieci sp. nov., Exoplectra misahualli sp. nov., Hyperaspis rutai sp. nov., Toxotoma gonzalezi sp. nov. Seven species are recorded for the first time from Ecuador: Cyrea noticollis (Mulsant, 1850), Eupalea formosa Mulsant, 1850, Epilachna transverselineata (Mader, 1858), Psyllobora marshalli Crotch, 1874, Toxotoma fuscopilosa (Weise, 1902), Toxotoma taeniola (Gordon, 1975), Zenoria linteolata Mulsant, 1850. For three species: Epilachna obtusiforma Gordon, 1975, Hinda ecuadorica Gordon & Canepari, 2013 and Siola atra González, 2015, supplementary notes on the original descriptions are provided. New name combinations are proposed for six species, which are transferred to the genus Toxotoma Weise, 1900: Toxotoma aequatorialis (Gordon, 1975), Toxotoma chigata (Gordon, 1975), Toxotoma flavocirculus (González, 2015), Toxotoma fuscopilosa, Toxotoma hybridula (Gordon, 1975), Toxotoma taeniola. The occurrence of Scymnus interruptus (Goeze, 1777) in Ecuador is also confirmed.
Two species of Xorides Latreille, 1809 are reported parasitizing wood-boring insects in trunks and relatively larger twigs of Juglans mandshurica Maxim. in Kuandian Manzu Autonomous County, Liaoning, in the Palaearctic part of China. Two new species are described: X. juglanse Sheng, Broad & Sun sp. nov. and X. kuandianense Sheng, Broad & Sun sp. nov. One species, X. sapporensis (Uchida, 1928), was associated with wood-borers in J. mandshurica Maxim. for the first time. A key to the 46 species of Xorides Latreille known from China is provided.
Ten species of Campodorus Förster, 1869 are reported from China and five species are new to science: C. albilineatus Sheng, Sun & Li sp. nov. from Guangxi Zhuang Autonomous Region in the Oriental part of China, C. punctatus Sheng, Sun & Li sp. nov. and C. rasilis Sheng, Sun & Li sp. nov. from Beijing, C. shandongicus Sheng, Sun & Li sp. nov. from Shandong Province and C. truncatus Sheng, Sun & Li, sp. nov. from Liaoning Province. Three species, C. ciliatus (Holmgren, 1857) and C. dauricus Kasparyan, 2005 collected from Liaoning Province and Mesoleius faciator Kasparyan, 2001 from Beijing, are new records for China. A key to species of Campodorus and a related genus known in China is provided.
Eleven species of Ctenopelma Holmgren, 1857 are reported from China. Five species are new to science: C. labiatum Sheng, Sun & Li sp. nov., from Ningxia Hui Autonomous Region, C. lii Sheng, Sun & Li sp. nov., from Liaoning province, C. rufofasciatum Sheng, Sun & Li sp. nov., reared from Cephalcia lariciphila (Wachtl, 1898) from Beijing, C. pineatum Sheng, Sun & Li sp. nov., reared from Acantholyda posticalis (Matsumura, 1912) and Cephalcia lariciphila from Beijing and Henan, Shanxi, Shan’xi, Liaoning, Jilin and Heilongjiang provinces, and C. spiraculare Sheng, Sun & Li sp. nov., reared from Cephalcia lariciphila from Henan and Shanxi provinces. One species, C. nigrum Holmgren, 1857, reared from Cephalcia lariciphila in Beijing, is a new record for China. Ctenopelma tomentosum (Desvignes, 1856) was reared from Neurotoma sibirica Gussakovskij, 1935 (new host record) in Liaoning province. A key to species of Ctenopelma known in China is provided.
The conspicuous Mediterranean brittle star Ophioderma longicauda (Bruzelius, 1805) has been discovered to represent a cryptic species complex, consisting of six nuclear clusters with contrasting reproductive modes (broadcast spawners and brooders). Here, O. longicauda is re-described. It is distinguished by a dark reddish-brown colouration both dorsally and on the ventral disc, and multiple tumid dorsal arm plates. One eastern Mediterranean brooding cluster is described as O. zibrowii sp. nov., characterized by a dark olive-green colour both dorsally and on the ventral disc, and single dorsal arm plates. Another brooder is described from Tunisia as O. hybrida sp. nov., with a highly variable morphology that reflects its origin by hybridization of O. longicauda and a brooder (possibly O. zibrowii sp. nov.), leaving the third brooding cluster as morphologically indistinguishable at this point and possibly conspecific with one of the others. The West-African O. guineense Greef, 1882 is resurrected as a valid species, differing morphologically from O. longicauda by predominantly single dorsal arm plates and light green or creamy white ventral side. Also from West Africa, O. africana sp. nov. is described, characterized by a dark brown colour, dorsally and ventrally, and single dorsal arm plates.
Based on intensive collecting from various sites in Sweden, the genus Dasyhelea Kieffer, 1911 was reviewed and the number of species now known from the country increased from five to twenty. Among the new species recorded there are two species described as new to science, D. dominiakae sp. nov. and D. gothlandica sp. nov., both in the subgenus Dicryptoscena Enderlein, 1936. The following subgenera are now documented from Sweden: Dasyhelea, Dicryptoscena, Pseudoculicoides Malloch, 1915, Prokempia Kieffer, 1913 and Sebessia Remm, 1979, the two latter subgenera being reported for the first time.
The first Oriental species of the genus Trichopsomyia Williston, 1888, Trichopsomyia pilosa sp. nov. (Java), has been discovered and is now described. This Oriental species of Trichopsomyia has several characters strongly differing from the other species within this genus. It is hypothesized that it forms a separate group within Trichopsomyia. One species, Trichopsomyia formiciphila Downes, Skevington & Thompson, 2017, from Australia, is similar to the Oriental species described here, and the group is named after this first described species, hence the formiciphila group. The characters for a future phylogenetic analysis are discussed. The character states of the pilosity of the katepisternum and the shape of the metasternum hitherto used in a phylogenetic analysis of Syrphidae Latreille, 1802 are discussed too.
The Oriental species of the genus Sphegina Meigen, 1822 are revised. The following 43 new species are described: Sphegina (Sphegina) abbreviata sp. nov. (Nepal), S. (S.) angustata sp. nov. (Nepal), S. (Asiosphegina) albolobata sp. nov. (Vietnam), S. (Asiosphegina) amplistylus sp. nov. (Philippines), S. (A.) atrimanus sp. nov. (Vietnam), S. (A.) bifida sp. nov. (Sabah, Malaysia), S. (A.) bracon sp. nov. (Vietnam), S. (A.) brevipilis sp. nov. (China), S. (A.) clavigera sp. nov. (Vietnam), S. (A.) collicola sp. nov. (Malaysia), S. (A.) crinita sp. nov. (Java, Indonesia; Malaysia), S. (A.) dentata sp. nov. (Taiwan), S. (A.) distincta sp. nov. (Vietnam), S. (A.) exilipes sp. nov. (Java, Indonesia), S. (A.) farinosa sp. nov. (Sabah, Malaysia), S. (A.) fimbriata sp. nov. (Thailand), S. (A.) furcillata sp. nov. (Vietnam), S. (A.) ghatsi sp. nov. (India), S. (A.) gigantea sp. nov. (China), S. (A.) granditarsis sp. nov. (China), S. (A.) hamulata sp. nov. (India), S. (A.) hauseri sp. nov. (Nepal), S. (A.) incretonigra sp. nov. (Vietnam), S. (A.) inflata sp. nov. (Philippines), S. (A.) inventum sp. nov. (Sabah, Malaysia), S. (A.) karnataka sp. nov. (India), S. (A.) licina sp. nov. (Thailand), S. (A.) lobulata sp. nov. (Vietnam), S. (A.) lucida sp. nov. (Vietnam), S. (A.) nigrotarsata sp. nov. (Vietnam), S. (A.) nubicola sp. nov. (Thailand), S. (A.) ornata sp. nov. (China), S. (A.) perlobata sp. nov. (Taiwan), S. (A.) plautus sp. nov. (China), S. (A.) prolixa sp. nov. (Malaysia, Thailand), S. (A.) setosa sp. nov. (Nepal, India), S. (A.) spathigera sp. nov. (Philippines), S. (A.) spenceri sp. nov. (Vietnam), S. (A.) strigillata sp. nov. (Vietnam), S. (A.) taiwanensis sp. nov. (Taiwan), S. (A.) umbrosa sp. nov. (China), S. (A.) verrucosa sp. nov. (Vietnam) and S. (A.) vietnamensis sp. nov. (Vietnam). Sphegina. (A.) tenuis Brunetti, 1915 is not a synonym of S. (A.) javana de Meijere, 1914. The males of S. (A.) apicalis Shiraki, 1930, S. (A.) tricoloripes Brunetti, 1915 and S. (A.) varidissima Shiraki, 1930 and the females of S. (A.) achaeta Hippa, van Steenis & Mutin, 2015, S. (A.) index Hippa, van Steenis & Mutin, 2015, S. (A.) mirifica Hippa, van Steenis & Mutin, 2015 and S. (S.) quadriseta Huo & Ren, 2006 are described for the first time. Sphegina (S.) quadriseta is recorded for the first time in the Oriental region. A key to all the Oriental species of Sphegina is provided. The Oriental fauna of Sphegina now comprises 94 species.
In this study, I describe two new species of Macrobiotus based on morphological data collected through light and scanning electron microscopy. Both species are accompanied by DNA sequences from four commonly used molecular markers (18S rDNA, 28S rDNA, ITS-2, and COI). Macrobiotus ovovittatus sp. nov. was discovered in Greenland and can be distinguished from similar taxa of Macrobiotus by its continuous, solid, and clearly wrinkled egg surface, adorned with sparse, very small and irregularly spaced pores. Additionally, the terminal discs of egg processes are covered in multiple light-refracting dots, resembling crocheted napkins. Macrobiotus mileri sp. nov. was found in Israel and is characterized by unique pore arrangements in its body cuticle, expressed in two distinct animal forms: (i) forma porata with large pores arranged in five distinct patches and (ii) forma aporata with single, almost undetectable pores. It also features weakly defined convex terminal discs with smooth edges. Furthermore, the phylogenetic analyses conducted in this study offer the most updated phylogeny of superclade I within the family Macrobiotidae. This facilitates additional discussion concerning the interrelationships among species within the genus Macrobiotus and the circumscription of species groups within it.
The New World euparine scarab genera Parataenius Balthasar, 1961 and Pseudataenius Brown, 1927 are revised. Ataenius brunneus Schmidt is transferred to the genus Parataenius becoming Parataenius brunneus (Schmidt), new combination. The monospecific genus Ataenioides Petrovitz, 1973, is synonymized with Pseudataenius Brown, 1927, (new synonymy) and the type species, Ataenioides gracilitarsis Petrovitz, is given the new combination Pseudataenius gracilitarsis (Petrovitz). New species of Parataenius are described from southern South America: Parataenius selvae, P. estero, and P. martinezi. Keys for species of both genera are presented and pertinent morphological details are illustrated.
A revision of the genus Acentroptera Guérin-Méneville, 1844 (Coleoptera: Chrysomelidae: Cassidinae)
(2014)
The species of the genus Acentroptera Guérin-Méneville, 1844 are revised. Thirteen species are treated as valid and are illustrated. A neotype is designated for A. tessellata Baly. Acentroptera maculata Pic from Brazil and A. rubronotata Pic from Brazil are treated as incertae sedis. Acentroptera bita n. sp. and A. lineata n. sp. both from Panama are described as new. A key to the 13 treated species is presented. Five species appear to be associated with bromeliads (Bromeliaceae).
First records of Elachistinae are given from Thailand. Ten species of Elachistinae are reported, eight of which are described as new: Urodeta longa Sruoga & Kaila sp. nov., Elachista buszkoi Sruoga & Kaila sp. nov., E. oryx Sruoga & Kaila sp. nov., E. pellineni Sruoga & Kaila sp. nov., E. capricornis Sruoga & Kaila sp. nov., E. phichaiensis Sruoga & Kaila sp. nov., E. loeiensis Sruoga & Kaila sp. nov. and E. siamensis Sruoga & Kaila sp. nov. The new species are diagnosed and illustrated with photographs of the adults and genitalia. One species remains unidentified to species level.
The ‘gigas’ group of dragon millipedes, formerly placed in the genus Desmoxytes Chamberlin, 1923, is revised and assigned to the new genus Gigaxytes gen. nov. Desmoxytes gigas Golovatch & Enghoff, 1994 is the type species of the new genus and is redescribed as G. gigas (Golovatch & Enghoff, 1994) gen. et comb nov. Three new species are described: G. fusca gen et sp. nov. from Thailand and Myanmar; G. parvoterga gen et sp. nov. and G. suratensis gen et sp. nov. from Thailand. All Gigaxytes species are endemic to small distribution areas in limestone habitats in South Thailand and South Myanmar. Illustrations of external morphological characters and an identification key to all known species are provided as well as a distribution map.
The ‘acantherpestes’ group of dragon millipedes, formerly placed in the genus Desmoxytes Chamberlin, 1923, is revised and assigned to the new genus Nagaxytes Srisonchai, Enghoff & Panha gen. nov. Desmoxytes acantherpestes Golovatch & Enghoff, 1994 is the type species of the new genus and is redescribed as N. acantherpestes (Golovatch & Enghoff, 1994) gen. et comb. nov. Three new species are described from Thailand: N. erecta Srisonchai, Enghoff & Panha gen. et sp. nov. and N. gracilis Srisonchai, Enghoff & Panha gen. et sp. nov. from Kanchanaburi Province, and N. spatula Srisonchai, Enghoff & Panha gen. et sp. nov. from Tak Province. All new species are endemic to western Thailand and all are restricted to limestone habitats. Complete illustrations of external morphological characters, an identification key, and a distribution map are provided.
New species and records of Sericini from the Indian subcontinent (Coleoptera, Scarabaeidae) II
(2019)
The current paper presents new locality records, including first state records for Mizoram and Nagaland, of 50 species of Sericini (Coleoptera: Scarabaeidae: Melolonthinae) from the Indian subcontinent. Nine new species are described herein: Maladera naveeni sp. nov., M. sujitrae sp. nov., M. thirthahalliensis sp. nov., M. viraktamathi sp. nov., Neoserica (s. lat.) reuteri sp. nov., Oxyserica goertzae sp. nov., Selaserica hosanagarana sp. nov., Serica (s. str.) eberlei sp. nov. and S. (s. str.) tashigaonensis sp. nov.
A new millipede species of the genus Sechelleptus Mauriès, 1980 is described and illustrated from Mayotte Island, Indian Ocean. This new species, S. arborivagus sp. nov., found on trees, looks particularly similar to the sympatric S. variabilis VandenSpiegel & Golovatch, 2007, but is much larger and has a very different ecological behavior. Phylogenetic analyses based on a concatenated dataset of the COI and 16S rRNA genes and including nine species of Spirostreptidae (including Sechelleptus, Doratogonus Attems, 1914, Bicoxidens Attems, 1928 and Spirostreptus Brandt, 1833), strongly support the monophyly of Sechelleptus. Despite the similarity of their genitalia, the molecular analyses also reveal a clear-cut genetic divergence between S. arborivagus sp. nov. and S. variabilis (22.55% for COI and 6.63% for 16SrRNA) and further suggest the presence of a higher diversity within the genus Sechelleptus on Mayotte.
The Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Fulgoromorpha, Dictyopharidae, Dictyopharinae, Orthopagini) is revised. Four species are included: C. jocosa (Gerstaecker, 1895) (the type species, with confirmed records from Cameroon, Equatorial Guinea and Gabon), C. lindbergae sp. nov. (described from Sierra Leone), C. rhinoceros sp. nov. (described from Togo) and C. simplex Melichar, 1912 (so far only known from Equatorial Guinea, Bioko island). Lectotypes are designated for C. jocosa and C. simplex and both species are redescribed including habitus photographs and detailed illustrations of the male and female genitalia which are published for the first time. A key for identification of the species of Centromeriana is provided. As far as known, the genus is endemic to the (Guineo-)Congolian region of western Africa.