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Abstract. More than 1300 specimens of Eucnemidae collected from Heredia Province in Costa Rica during the 1990s Arthropods of La Selva (ALAS) survey were studied from 2018 through 2022. One new genus of false click beetle, Absensiugum Otto, Muona and Córdoba-Alfaro, is described. Nematodes teres Horn, from the Nearctic and Caribbean regions, is transferred to this new genus to form Absensiugum teres, new combination. Sixteen new species of false click beetle (Coleoptera: Eucnemidae) are described from Costa Rica. These new species are: Adelothyreus brevis, Adelothyreus costaricensis, Adelothyreus totus, Quirsfeldia stethonoides, Lacus pectinatus, Maelodrus costaricensis, Onichodon confluentus, Onichodon rufus, Isarthrus striatus, Absensiugum brunneum, Dromaeolus americanus, Dromaeolus brunneus, Dromaeolus herediensis, Dromaeolus holdridgei, Deltometopus bicolor and Nematodes apicalis. Three additional records outside of the Heredia Province from the Osa Peninsula and Panama for Lacus pectinatus are included in this study. Identification keys are provided for species of Adelothyreus Chevrolat, Onichodon Newman, Dromaeolus Kiesenwetter, Deltometopus Bonvouloir and Nematodes Berthold in Costa Rica. Diagnostic differences are briefly noted for each species within the Neotropical region. A list of Eucnemidae from Heredia Province is provided.
ZooBank registration. urn:lsid:zoobank.org:pub:C1D5B819-A964-4679-B090-84CDBBC59D6A
Thomsonista Nearns and Nascimento, a new genus of Onciderini Thomson, 1860 (Coleoptera: Cerambycidae: Lamiinae), is described and illustrated. Seven new species of Onciderini are described and illustrated: Hesychotypa antonkozlovi from Ecuador; Hesychotypa danilevskyi from Panama; Lingafelteria pandolfii from Brazil; Oncideres antonkozlovi and Oncideres erwini from Peru; Oncideres johnmarvini from Costa Rica and Panama; and Thomsonista antonkozlovi from Colombia.
A new Brazilian leafhopper genus of the subfamily Neocoelidiinae is proposed: Takiyaella gen. nov. based on T. sexguttata (Chiamolera & Cavichioli, 2003) gen. et comb. nov. and five new species: T. anomala gen. et sp. nov., T. coelhomarquesae gen. et sp. nov., and T. daniela gen. et sp. nov. from the state of Rio de Janeiro; T. cavichiolii gen. et sp. nov. from the state of Paraná, and T. mejdalanii gen. et sp. nov. from the state of Minas Gerais. A distribution map and identification key to Takiyaella species are provided.
Here I describe a new genus, Iviephengus gen. nov., based on a single species, Iviephengus ferreirai gen. et sp. nov., from Peru. This new genus is characterized by the following combination of characters: interantennal distance close to 3 × the antennal socket length; antenna 12-segmented, IV to XI each with two long symmetrical compressed and apically slightly enlarged branches; labrum fused to frontoclypeus; mandibles short, obliquely crossed, each with a notch on the external margin to fit the other mandible and without extra teeth; maxillary palpi 4-segmented, last segment digitiform; labial palpi 2-segmented; posterior tentorial pit consisting of a single small fossa; wing with radial cell closed and transverse, vein r4 interrupted, r3 absent; first tarsomere of pro- and mesotarsus with a ventral comb covering the posterior half of the tarsomere; claws simple, without any teeth; aedeagus with paramere symmetrical, apex unevenly round, toothed inward, with short and sparse bristles. I provide a key to Mastinocerinae genera with 12-segmented antennae and the first pro- and mesotarsomere with ventral combs. I also provide illustrations for the diagnostic features for this new genus. Finally, I discuss the presence and function of some modifications in the mandible and the sternite VIII in Phengodidae and other Coleopteran families.
A new feature for the separation of Trochosa spinipalpis and T. terricola males (Araneae, Lycosidae)
(2006)
A new feature on the tip of the palp, which enables the separation of male Trochosa spinipalpis (F. O. P.-Cambridge, 1895) from T. terricola Thorell, 1856, is described. T. terricola exhibits a hairless strip on the tip of the palp, while T. spinipalpis lacks this feature and has long hairs on the whole palp.
We describe a new species of Cyrtodactylus on the basis of two specimens collected from Ta Kou Nature Reserve, Binh Thuan Province, southern Vietnam. Cyrtodactylus chungi sp. nov. is distinguished from the remaining Indochinese bent-toed geckos by a combination of the following characters: relatively small body size (SVL up to 68.5 mm); a continuous neckband; 5 or 6 irregular transverse dorsal bands; 11 or 12 bands on original tail; keeled tubercles present on dorsum, posterior limbs and tail; 17 or 18 irregular dorsal tubercle rows; 30 or 31 ventral scale rows; ventrolateral skin folds indistinct; an angular series of seven precloacal pores in male and six pitted, enlarged precloacal scales in female, each series separated by a diastema of undifferentiated scales from 4–6 enlarged, poreless femoral scales; median subcaudals slightly enlarged; 17–20 subdigital lamellae under the fourth toe. Based on molecular analyses of the fragment of mitochondrial gene cytochrome c oxidase subunit I (COI), the new species is recovered as the sister taxon to Cyrtodactylus cattienensis s. str. with a genetic divergence of more than 9%. In phylogenetic analyses, the new species is recovered as a member of the Cyrtodactylus irregularis species group.
A new cypridopsine genus, Cyprettadopsis gen. nov., described here, is principally characterized by the reduced caudal ramus, the strongly serrated claw G2 of the antenna (A2), the A2 subquadrate terminal segment, the undivided penultimate segment of the second thoracopod (T2), the morphology of the third thoracopod bearing a distinctly separated terminal segment, the complete septa on the posteroventral margin and the incomplete septa on the anterior margin of both valves. Based on a combination of these characters, a new tribe, Cyprettadopsini trib. nov., is created in the subfamily Cypridopsinae Kaufmann, 1900 to accommodate this new genus, and one new species, Cyprettadopsis sutura gen. et sp. nov., is described as the type species. Apart from the above generic characters, the following features are also typical of the new species: the tiny needlepoint-like pores along the anterior and ventral margins of both valves, the remarkably large β-seta on the mandibular palp and the considerably short d2 seta on the T2. The presence of marginal septa in the new genus is a distinctive character and constitutes the first record of this feature within Cypridopsinae. The taxonomically relevant characters in the new taxon and related taxa are briefly discussed.
A new cryptic species, Aleochara (Xenochara) castaneimarmotae Klimaszewski, Webster, and Brunke, new species (Coleoptera: Staphylinidae: Aleocharinae), associated with Marmota monax (Linnaeus) burrows and caves in eastern North America, is described and illustrated. A key to Canadian species of subgenus Xenochara Mulsant and Rey and revised distributions of the taxonomically difficult fumata species group are provided. Aleochara quadrata Sharp is recorded from Washington and Oregon for the first time.
The lichen-forming genus Pertusaria under its current circumscription is polyphyletic and its phylogenetic affiliations are uncertain. Here we study the species of the genera Pertusaria and Varicellaria which containlecanoric acid as major constituent, have disciform apothecia, strongly amyloid asci, non-amyloid hymenial gel, 1-2-spored asci, and 1- or 2-celled ascospores with thick, 1-layered walls. We infer phylogenetic relationships using maximum likelihood and Bayesian analyses based on four molecular loci (mtSSU, nuLSU rDNA, and the protein-coding, nuclear RPB1 and MCM7 genes). Our results show that the lecanoric acid-containing species form a well-supported, monophyletic group, which is only distantly related to Pertusaria s.str. The phylogenetic position of this clade is unclear, but placement in Pertusaria s.str. is rejected using alternative hypothesis testing. The circumscription of the genus Varicellaria is enlarged to also include species with non-septate ascospores. Seven species are accepted in the genus: Varicellaria culbersonii (Vězda) Schmitt & Lumbsch, comb. nov., Varicellaria hemisphaerica (Flörke) Schmitt & Lumbsch, comb. nov., Varicellaria kasandjeffii (Szatala) Schmitt & Lumbsch, comb. nov., Varicellaria lactea (L.) Schmitt & Lumbsch, comb. nov., Varicellaria philippina (Vain.) Schmitt & Lumbsch, comb. nov., Varicellaria rhodocarpa (Körb.) Th. Fr., and Varicellaria velata (Turner) Schmitt & Lumbsch, comb. nov. A key to the species of Varicellaria is provided.
A new planthopper genus, Aodingus Chen & Li gen. nov. and three new species (A. hainanensis Chen & Li gen. et sp. nov., A. obscurus Chen & Li gen. et sp. nov. and A. cuongi Chen & Li gen. et sp. nov.) are described from China and Vietnam. The new genus is superficially similar to Procidelphax Bartlett, 2009 in general appearance in that the body is strongly dorsoventrally flattened. Distinctive features of the new taxon include broadly compressed body with vertex broad, apical margin broadly rounded, middle part concave, median and submedian carinae absent, frons broad and large, wider at base than at apex, forewing broad and long, aedeagus tubular, curved ventrally. A diagnosis for all species, illustrations and an identification key of new genus are provided. A key to the Chinese genera of Tropidocephalini is also provided.
Bureschia serbica sp. nov., is described from a cave in the Stara Planina Mt in Serbia as a second species in the genus. It is a troglobite amphibious species, predominantly aquatic like the nominal species of the genus, Bureschia bulgarica. The new species is adapted to amphibious life in running groundwater, unlike the nominal one that lives in subterranean lakes and ponds. Body structure changes, most noticeable the significant changes in the structure of the mouthparts, as a result of preadaptation in aquatic and amphibious Synocheta, are discussed.
Lodevoisadia coheni gen. et sp. nov. is described as the ninth species of ‘Grylloblattodea’ from the middle Permian of the Salagou Formation, near Lodève town (France). It is currently not reasonable to place this species into a specific family, even though it seems to share most characters with the small family Tunguskapteridae. The lack of phylogenetic analysis and the current poor delineation of the majority of the grylloblattodean families (lacking synapomorphies) render any attribution of new taxa to a particular family often uncertain.
The “trachystreptoform” species of Spirostreptidae, i.e., species which would formerly have been ascribed to the tribe Trachystreptini, from the Udzungwa Mountains are (re)described, including one new genus and five new species: Attemsostreptus reflexus Akkari & Enghoff, 2019, A. cataractae Enghoff sp. nov., A. leptoptilos Enghoff sp. nov., A. julostriatus Enghoff sp. nov., Lophostreptus tersus (Cook, 1896) (= L. ptilostreptoides Carl, 1909 syn. nov.), L. magombera Enghoff sp. nov., and Udzungwastreptus marianae Enghoff gen. et sp. nov. The type material of Lophostreptus regularis Attems, 1909 (= L. tersus) is discussed. The discussion includes paragraphs on the classification and the Udzungwa fauna of Spirostreptidae, on grouping of the Udzungwa trachystreptoform species in relation to altitude, and on the possibly recent immigration of A. reflexus and L. tersus into the Udzungwa Mts.
Three new species of the family Pyrgodesmidae Silvestri, 1896 from the Udzungwa Mountains are described: Cryptocorypha geminiramus sp. nov., Cryptocorypha cactifer sp. nov., and Cryptocorypha exovo sp. nov. Several additional, unidentifiable pyrgodesmids from the Udzungwa Mountains are recorded. Elythesmus enghoffi Hoffman, 1978 (Cryptodesmidae Karsch, 1880), is recorded from the East Usambara (type locality), West Usambara, Uluguru, Nguru (Kanga) and Udzungwa Mountains, and its variation in size and body ring numbers is analyzed. The surface sculpture of pro- and metazonites is discussed.
A large, newly collected material of the genus Aquattuor from the Udzungwa Mts, Tanzania, has been studied. Two new species are described, viz., A. mollilobus sp. nov. from the Udzungwa Mts and A. nguruensis sp. nov. from the Nguru Mts, Tanzania. Aquattuor claudiahempae Enghoff & Frederiksen, 2015, is recorded as new for the Udzungwa fauna. Characters from the first pair of male legs are illustrated for all Aquattuor species and are shown to be useful for species discrimination. A new gonopodal difference between A. submajor Enghoff, 2015 and A. udzungwensis Enghoff, 2015 is described and illustrated. Intrageneric relationships and distribution patterns are discussed. A distribution map and a key to all Aquattuor species are presented.
Damacornu gen. nov. (type species: D. transversum gen. et sp. nov.), Geotypodon papei sp. nov. and Spinotarsus fortehamatus sp. nov. are described, and Helicochetus dimidiatus (Peters, 1855), H. mutaba Kraus, 1960 and Hoffmanides dissutus (Hoffman, 1963) are recorded from the Udzungwa Mts, Tanzania. A complete overview of the 39 odontopygid species now known from the Udzungwa Mts is given, including notes on endemism, biogeographical relationships and altitudinal distribution patterns.
Three new genera of Odontopygidae are described, all based on new species from the Udzungwa mountains, Tanzania, and all monotypic: Casuariverpa gen. nov. (type species: C. scarpa gen. et sp. nov.), Yia gen. nov. (type species: Y. geminispina gen. et sp. nov.), and Utiliverpa gen. nov. (type species: U. decapsulatrix gen. et sp. nov.). Similarities and differences between the new genera and other genera are discussed.
Two species of the genus Prionopetalum Attems, 1909, are recorded from the Udzungwa Mountains: P. asperginis sp. nov. and P. kraepelini (Attems, 1896). Prionopetalum stuhlmanni Attems, 1914, is synonymized under P. kraepelini. Odontopyge fasciata Attems, 1896, is transferred from Prionopetalum to Aquattuor Frederiksen, 2013, and new illustrations are given. A new illustrated key to species of Prionopetalum is provided.
The new genus Geotypodon gen. nov. is described. It includes two species from the Udzungwa Mountains: G. millemanus gen. et sp. nov. (type species) and G. submontanus gen. et sp. nov., one species from nearby Iringa: G. iringensis gen. et sp. nov., and 18 previously described species hitherto incorrectly assigned to Odontopyge Brandt, 1841.
The genus Aquattuor Frederiksen, 2013 is revised. A. denticulatus Frederiksen, 2013 (type species) from the East Usambara Mts, Tanzania, is redescribed, and six new species are described: A. claudiahempae sp. nov. from Mt. Kilimanjaro, Tanzania, and five species from the Udzungwa Mts, Tanzania: A. longipala Enghoff sp. nov., A. major Enghoff sp. nov., A. stereosathe Enghoff sp. nov., A. submajor Enghoff sp. nov., and A. udzungwensis Enghoff sp. nov.
Twenty new species of the millipede genus Chaleponcus Attems, 1914, are described from the Udzungwa Mountains: C. netus sp. nov., C. quasimodo sp. nov., C. malleolus sp. nov., C. scopus sp. nov., C. nikolajscharffi sp. nov., C. mwanihanensis sp. nov., C. basiliscus sp. nov., C. krai sp. nov., C. nectarinia sp. nov., C. circumvallatus sp. nov., C. ibis sp. nov., C. vandenspiegeli sp. nov., C. vilici sp. nov., C. teres sp. nov., C. hamerae sp. nov., C. termini sp. nov., C. gracilior sp. nov., C. mwabvui sp. nov., C. howelli sp. nov. and C. tintin sp. nov. Together with C. dabagaensis Kraus, 1958, they constitute the Chaleponcus dabagaensis-group, well characterized by apparently apomorphic gonopodal characters, presumably monophyletic, and the first example of a major radiation within the Udzungwas. All species are restricted to altitudes >1390 m, all but one were found in only one, rarely two forest reserves, and the vast majority of specimens were collected in montane forest. Chaleponcus gracilior sp. nov. was collected in four forest reserves, often in secondary habitats where other species were only exceptionally found. Co-occurrence of multiple species, inter-specific differences in body size and unusual tarsal setation of a few species tentatively suggest adaptive radiation.
A morphological and molecular review of the genus Goniurosaurus, including an identification key
(2021)
The genus Goniurosaurus (tiger geckos) currently consists of 23 species distributed in China, Japan and Vietnam. Several species complexes and recent discoveries of cryptic species pose challenges to the species identification, which is crucial to effectively implement the recent listing of the species from China and Vietnam in CITES Appendix II and the species from Japan in CITES Appendix III. Based on the results of our field work in northern Vietnam and data compiled from literature, we herein provide a taxonomic review of the genus Goniurosaurus. Our phylogenetic analyses showed that all recorded populations of tiger geckos from Vietnam, which were found to be monophyletic with low intra-specific genetic divergences, are assigned to one of the four species: G. catbaensis, G. huuliensis, G. lichtenfelderi or G. luii. Both genetic and morphological analyses confirm that the species from China and Vietnam can be split into three major groups. Based on the newly collected data, we provide an extended morphological description of the Vietnamese species. In addition, we provide an identification key for all Goniurosaurus species from China, Japan and Vietnam in order to assist authorities in the enforcement of the recent CITES listing.
Although extensively studied by different authors over the past 150 years, the taxonomy of Canthon Hoffmannsegg, 1817 and allied genera (which are here informally referred to as 'Canthon sensu lato') still remains problematic. With the aim of resolving some of the questions surrounding these taxa, the present work reviews the taxonomy of one of them, the genus Sylvicanthon Halffter & Martínez, 1977. As defined here, Sylvicanthon is distributed mainly throughout the vast areas of tropical rainforests in the Neotropical region and includes 15 species divided into two groups: the enkerlini group, with a single species, S. enkerlini (Martínez et al., 1964) comb. nov., and the candezei group, with five subgroups: the candezei subgroup, with S. candezei (Harold, 1869), S. genieri sp. nov. and S. foveiventris (Schmidt, 1920); the aequinoctialis subgroup, with S. aequinoctialis (Harold, 1868) comb. nov. and S. proseni (Martínez, 1949) stat. et comb. nov.; the bridarollii subgroup, with S. bridarollii (Martínez, 1949), S. seag sp. nov., S. edmondsi sp. nov. and S. attenboroughi sp. nov.; the furvus subgroup, with S. furvus (Schmidt, 1920), S. monnei sp. nov., S. mayri sp. nov. and S. obscurus (Schmidt, 1920); and the securus subgroup, with a single species, S. securus (Schmidt, 1920) comb. nov. Three species originally included in Sylvicanthon are here (re)transferred to Canthon: Canthon xanthopus Blanchard, 1846 and C. machadoi (Martínez & Pereira, 1967) comb. nov., as well as C. cobosi (Pereira & Martínez, 1960) stat. et comb. nov., which had been previously in synonymy under C. xanthopus. Descriptions, redescriptions, illustrations and comparative tables on the external morphology (including the genital capsule) of the genus and its species are presented, as well as a detailed discussion on their biogeography, comparative morphology, hypotheses on their phylogenetic relationships, data on natural history and a detailed historical revision of the classification of 'Canthon sensu lato'. Finally, we also discuss the socalled 'species problem' (i.e., the definition of the scientific term 'species') and its consequences to dung beetle taxonomy and favour the solution offered by the Biological Species Concept.
A new genus in the tribe Issini (Hemiptera: Fulgoromorpha: Issidae) is described from South China: Orbita Meng & Wang, gen. nov., its type species Orbita parallelodroma Meng & Wang, gen. et sp. nov. (China: Fujian) is described and illustrated. In addition, one new species of Neokodaiana Yang, N. minensis sp. nov. is described and illustrated from the same locality as the new genus. A key to the genera of Issini from China and neighbouring countries is provided.
A glance at the Odonata of the Cambodian coastal mountainous regions : end of dry season in 2010
(2010)
During a visit to the coastal regions of Cambodia within the Provinces of Koh Kong, Kampong Saom, Kampot and Kep within April 12-23, 2010, 59 species of Odonata were found. Of these, 51 were identified, 2 preliminarily identified (1 by exuviae, 1 to a known species with the valid name unresolved), 1 dubious (Aciagrion cf. pallidum), 3 not identified (just visual observations) and 2 seemingly undescribed, namely Burmagomphus sp. and Microgomphus sp. Fourteen species were recorded for the first time for this country, namely Neurobasis chinensis, Dysphaea gloriosa, Euphaea masoni, Onychargia atrocyana, Copera marginipes, Copera vittata, Prodasineura autumnalis Prodasineura verticalis sensu Asahina, 1983, Lathrecista asiatica, Orthetrum chrysis, Rhyothemis obsolescens, Tramea transmarina euryale, Zygonyx iris malayana, Zyxomma petiolatum; taking the current checklist for Cambodia up to 83 species. Some features of the collected specimens relevant to their taxonomy are discussed. Brief field notes are provided and Odonata records enumerated. The habitats and community of Odonata at the Cardamon Mts. foothills at the end of dry season are discussed.
Dual-task paradigms encompass a broad range of approaches to measure cognitive load in instructional settings. As a common characteristic, an additional task is implemented alongside a learning task to capture the individual’s unengaged cognitive capacities during the learning process. Measures to determine these capacities are, for instance, reaction times and interval errors on the additional task, while the performance on the learning task is to be maintained. Opposite to retrospectively applied subjective ratings, the continuous assessment within a dual-task paradigm allows to simultaneously monitor changes in the performance related to previously defined tasks. Following the Cognitive Load Theory, these changes in performance correspond to cognitive changes related to the establishment of permanently existing knowledge structures. Yet the current state of research indicates a clear lack of standardization of dual-task paradigms over study settings and task procedures. Typically, dual-task designs are adapted uniquely for each study, albeit with some similarities across different settings and task procedures. These similarities range from the type of modality to the frequency used for the additional task. This results in a lack of validity and comparability between studies due to arbitrarily chosen patterns of frequency without a sound scientific base, potentially confounding variables, or undecided adaptation potentials for future studies. In this paper, the lack of validity and comparability between dual-task settings will be presented, the current taxonomies compared and the future steps for a better standardization and implementation discussed.
New World species of the histerid beetle genus Xestipyge Marseul (Coleoptera: Histeridae: Dendrophilinae: Paromalini) are reviewed, and X. skelleyi, new species, is described. A dichotomous key and illustrations are provided as identification means for the now five included species from the Western Hemisphere.
Als Ergebnis eines aufwendigen, aber erfolgreichen Kreuzungsexperimentes von Acalles aeonii Wollaston 1864 x Acalles bodegensis Stüben 2000 wird folgende Synonymie endgültig bewiesen: Acalles aeonii Wollaston 1864 = Acalles bodegensis Stüben 2000 syn. nov. Zucht und Kreuzung der genannten Arten werden über zahlreiche Generationen hinweg an der Entwicklungspflanze Aeonium holochrysum W. & B. detailliert beschrieben und die Ergebnisse diskutiert. Mit einer ausführlichen Diskussion zur Notwendigkeit von Kreuzungsexperimenten in der taxonomisch-systematischen Forschung.