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Twelve new species are assigned to the genus Otitoma Jousseaume, 1898 in the family Pseudomelatomidae Morrison, 1966 and herein described: O. hadra sp. nov., O. neocaledonica sp. nov., O. rubiginostoma sp. nov and O. tropispira sp. nov. from New Caledonia; O. boucheti sp. nov., O. nereidum sp. nov. and O. sororcula sp. nov. from the Fiji Islands; O. xantholineata sp. nov. from the Solomon to the Fiji Islands; O. crassivaricosa sp. nov. from Fiji to Hiva Oa Island (Marquesas Archipelago); O. philpoppei sp. nov. from the Philippines but also reported from the Fiji Islands; O. elegans sp. nov. from the Fiji Islands and O. philippinensis sp. nov. from the Philippines. New data on O. carnicolor (Hervier, 1896) are provided. Otitoma mitra (Kilburn, 1986), from Southern Mozambique, is here considered a synonym of O. cyclophora (Deshayes, 1863). Drillia batjanensis Schepman, 1913, previously assigned to the genus Maoritomella Powell, 1942 in the family Borsoniidae Bellardi, 1875, is here assigned to the genus Otitoma. Photographs of the holotype of Drillia batjanensis are provided for the first time. In addition, color photographs of the type specimens of the following species are provided: Drillia kwandangensis Schepman, 1913, D. timorensis Schepman, 1913 and Mitrellatoma mitra Kilburn, 1986.
Contributions to the knowledge of the mite genus Stigmaeus Koch, 1836 (Acari: Stigmaeidae) of Turkey
(2017)
Based on the mite specimens collected within the scope of a study on Erzincan (Turkey) mite biodiversity, two species of the genus Stigmaeus are described and illustrated here: S. bifurcus sp. nov. as new to science and S. miandoabiensis Bagheri & Zarei, 2012 as a new record for Turkey. Some morphological abnormalities in the new species are noted. The deutonymph of S. miandoabiensis is described for the first time in this study. Discovery of this stage from soil and litter under Pinus sylvestris in Turkey adds more data to our knowledge of the species.
The genus Oshaibahus El-Sonbati & Wilson gen. nov. is described with the type species Platymetopius zizyphi Bergevin, 1922. Two new species, O. kadiae El-Sonbati & Wilson gen. et sp. nov. from Sudan and O. linnavuorii El-Sonbati & Wilson gen. et sp. nov. from Iraq are described, and a new combination, Oshaibahus zizyphi (Bergevin, 1922) gen. et comb. nov. is proposed. The genus Masiripius Dlabola, 1981 (type species: Mahalana lugubris Distant, 1918) is redefined. An illustrated key to genera of the subtribe Opsiina and a key to species of Oshaibahus gen. nov. are presented to facilitate identification.
One of the main challenges to adequately conserve the African fish fauna is to improve our so far unsatisfactory taxonomic knowledge of important portions of the ichthyofauna. In the present study, we attempted to unravel the taxonomic diversity of some species of Enteromius Cope, 1867, a problematic African fish genus, recently collected in the north-eastern part of the Congo basin. We used an integrative approach, combining DNA barcodes and morphological analyses. For one of the species complexes found, the E. miolepis/eutaenia species complex, we evaluated taxonomic diversity over a larger geographic scale within the Congo drainage system. Although initial literature-based species identifications allowed us to assign all examined specimens to four tentative species, DNA barcodes indicated the presence of 23 distinct mitochondrial lineages. The majority of these lineages appeared endemic to particular rivers, and in most rivers multiple lineages occur in sympatry. Subsequent exploratory morphometric analyses indicated that almost all these lineages are morphologically distinguishable and that they may therefore represent undescribed species. As only a part of the Congo basin and a subset of the species diversity within Enteromius were examined, it appears that the species richness of Enteromius in the Congo basin is severely underestimated.
The inventory of biodiversity in the Comoros archipelago is still in progress. We propose here to contribute to this effort by assessing the diversity of the Hymenophyllaceae family, which forms one of the most diverse and emblematic fern communities in rainforests of the region. Data were extracted from floras, literature, recent collects and observations, and from an exhaustive investigation of the Paris herbarium collection. We also completed an online knowledge database including a computeraided identification (CAI) tool by using the Xper platform. Our checklist recognizes 21 taxa involving 1 endemic and 7 species newly reported for the archipelago. The taxonomy and discrimination of each taxon is discussed and a dichotomous key is provided. We also defined 80 morphological characters and their corresponding states into the Xper database and CAI that could be easily enriched for neighbouring areas and additional taxa.
Scandinavian members of the acrocarpous moss genus Oncophorus were revised after field observations had suggested unrecognized diversity. Based on molecular (nuclear: internal transcribed spacers 1 and 2, ITS; plastid: trnGUCC G2 intron, trnG, rps4 gene + trnS-rps4 spacer, rps4) and morphological evidence, four morphologically distinguishable species are recognized, Oncophorus elongatus (I.Hagen) Hedenäs, O. integerrimus Hedenäs sp. nov. (syn. O. virens var. elongatus Limpr.), O. virens (Hedw.) Brid., and O. wahlenbergii Brid. (O. sardous Herzog, syn. nov.). Oncophorus elongatus was earlier recognized, but much of its variation was hidden within O. wahlenbergii. Its circumscription is here expanded to include plants with long leaves having mostly denticulate or sharply denticulate upper margins and with long and narrow marginal cells in the basal portion of the sheathing leaf lamina. The new species O. integerrimus sp. nov. differs from O. virens in having more loosely incurved leaves and entire or almost entire upper leaf margins. Besides these characters, the species in the respective pairs differ in quantitative features of the leaf lamina cells. Several cryptic entities were found, in several cases as molecularly distinct as some of the morphologically recognizable species, and phylogeographic structure is present within O. elongatus and O. virens.
Ichneumonopsis Hardy,1973, a genus of oriental fruit flies, is revised and two new species, I. hancocki sp. nov. (from Peninsular Malaysia) and I. taiwanensis sp. nov. (from Taiwan), are described. A key to the three species of Ichneumonopsis is presented. In northern Thailand larvae of I. burmensis Hardy, 1973 develop in bamboo shoots of Pseudoxytenanthera albociliata (Munro) Nguyen and Dendrocalamus strictus (Roxburgh) Nees (Poaceae), not Melocalamus compactiflorus as previously reported. The recently discovered association of I. burmensis with bamboo substantiates our previous assumption assigning Ichneumonopsis to the primarily bamboo-inhabiting tribe Gastrozonini. Hence, we synonymize Ichneumonopsidini under Gastrozonini (syn. nov.).
The Middle East biting midges of the tribes Palpomyiini (20 species in three genera) and Sphaeromiini s. lat. (six species in five genera) are reviewed. Three new species are described and illustrated: Bezzia libanensis Alwin & Szadziewski sp. nov., B. sharjahi Alwin & Szadziewski sp. nov. and Palpomyia freidbergi Alwin & Szadziewski sp. nov. Bezzia aegyptia Kieffer, 1925 is recognized as a new junior synonym of B. albicornis (Meigen, 1818) (syn. nov.) and B. omanensis Boorman & van Harten, 2002, is recognized as a junior synonym of B. (Sivabezzia) pachypyga Remm, 1974 (syn. nov.). Keys to the genera and species of the tribes Palpomyiini and Sphaeromiini of the Middle East are also provided.
The Chimarra lehibemavo species-group, new and endemic to Madagascar (Trichoptera, Philopotamidae)
(2017)
The Chimarra lehibemavo group is described to include thirteen new species: Chimarra lehibemavo sp. nov., C. cebegepi sp. nov., C. fenoevo sp. nov., C. forcellinii sp. nov., C. fotobohitra sp. nov., C. gattolliati sp. nov., C. gensonae sp. nov., C. jejyorum sp. nov., C. hamatra sp. nov., C. makiorum sp. nov., C. moramanga sp. nov., C. saha sp. nov. and C. tamara sp. nov. The adults are easily recognizable by their large size, yellow colour and the structure of the male genitalia. The membranous tergum IX and the absence of the mesal lobe of tergum X are observed in other lineages, but the strong asymmetrical deformation of the phallotheca is apomorphic. The group is monophyletic with unknown affinities, but a preliminary phylogenetic placement is suggested following genetic analysis of two specimens. With one exception, the species have restricted geographical distributions in Madagascar and inhabit rivers in eastern pristine rainforests.
To date, six species of the Australian endemic millipede genus Boreohesperus have been recognized: all have highly localized distributions, consistent with being short-range endemic species, and all are from the Cape Range and Pilbara region of Western Australia. In this paper, we describe three new species, B. alcyonis sp. nov., B. psittacinus sp. nov., and B. vascellus sp. nov., each from a different island in the Kimberley region of north-western Australia.
Saurichthyidae is a speciose group of fishes, ranging from the Late Permian to the Middle Jurassic. Early Jurassic saurichthyids are usually considered morphologically less disparate and taxonomically less diverse than their Triassic counterparts, consisting of only two valid species. These were historically differentiated almost entirely based on cranial ratios, and both had stratigraphic ranges spanning the Early Jurassic. Here, we revise the Early Jurassic saurichthyid fishes of Europe based on restudy of the type material of Saurorhynchus brevirostris (Woodward, 1895) and S. acutus (Agassiz, 1844). We identify four species based on cranial osteology: S. acutus, S. brevirostris, S. anningae sp. nov., and S. hauffi sp. nov. Saurorhynchus brevirostris and S. anningae sp. nov. are known from the Sinemurian of England only, whereas S. acutus and S. hauffi sp. nov. share a broad European distribution during the Toarcian, from southwestern Germany to England. Saurorhynchus anningae sp. nov. and S. brevirostris, in particular, show disparity in such ecomorphologically important traits as cranial fineness, and tooth and jaw morphology. In contrast, S. acutus and S. hauffi sp. nov. show much lower levels of disparity, differing from each other in nostril morphology, dermal ornamentation, and position of the lateral extrascapular. The new species do not increase diversity estimates for Saurorhynchus in either the Sinemurian or Toarcian interval.
A new species of vermetid gastropod belonging to the genus Novastoa Finlay, 1926, N. rapaitiensis sp. nov., is described from French Polynesia and the Great Barrier Reef, based on morpho-anatomical and molecular data, increasing the recognized extant diversity of this genus from five to six species. The new species is characterized by the largest operculum in the genus to date, with a conspicuous spindle-shaped mammilla that readily distinguishes this species from its congeners. Based on available data, members of the studied populations of N. rapaitiensis sp. nov. are interpreted as a single species, although slight morphological and color differences exist between localities separated by up to 7000 km. The larval development of N. rapaitiensis sp. nov. is inferred to be direct, raising new questions about genetic connectivity and dispersal trajectories across a vast geographic range. Additional molecular data may prove helpful in refining our current knowledge on the morphological variability within this species and verifying the degree of cryptic diversity in this genus.
Examination of leucothoid amphipods of the Red Sea has revealed seven species not previously reported from this location. Leucothoe minoculis sp. nov., Leucothoe pansa sp. nov., Leucothoe reimeri sp. nov., and Paranamixis sommelieri sp. nov. are described and the range of Leucothoe predenticulata Ledoyer, 1978, L. acutilobata Ledoyer, 1978 and L. squalidens Ledoyer, 1978 is expanded to include the Red Sea. Clarification of reports of L. acanthopus Schellenberg, 1928 and L. bannwarthi (Schellenberg, 1928) is provided and Leucothoe alani sp. nov. is described from outside the Red Sea.
We describe a new minute species of the genus Pristimantis, P. boucephalus sp. nov., from the Yanachaga-Chemillén National Park, Región Pasco, Peru. The description is based on a freshly collected male specimen found at 2950 m a.s.l. in a cloud forest and four previously unidentified museum specimens consisting of two adult males, one subadult female and a juvenile from the Yanachaga-Chemillén National Park. The new species is mainly characterized by a snout–vent length of 13.4–14.5 mm in adult males (n = 3), and 12.5 mm in the only known subadult female, and is compared morphologically and genetically with other taxonomically and biogeographically relevant species of Pristimantis. The new species is characterized by its small size, disproportionally large head with short snout, absence of a tympanic annulus and membrane, and reddish-copper iris. Phylogenetically it belongs to a speciose clade, an as yet unnamed species group, comprising both montane (Andes, Guiana Shield) and lowland (Amazon) taxa from the northern part of South America. The new species is genetically close to the sympatric P. cruciocularis. Species of Pristimantis occurring in the Cordillera Yanachaga region in the Andes of central Peru are members of six divergent phylogenetic lineages.
The genus Koiulus gen. nov. and its type-species, Koiulus interruptus gen. et sp. nov., are described from the Russian Far East. The new genus is compared with other genera of Mongoliulidae, in particular with Ussuriiulus Golovatch, 1980, also from the Russian Far East, with which it shares the absence of ozopores from individual body rings distributed along the body, a condition so far otherwise unknown in the superorder Juliformia. A synoptic table of genera and a list of species of Mongoliulidae are presented.
The species richness of the aquatic (non-marine) gastropod family Hydrobiidae Stimpson, 1865 reaches its peak in the European region partly because other areas, such as North Africa, remain to be extensively surveyed. Most of the hydrobiid species described in North Africa have been ascribed to the subfamilies Pseudamnicolinae, Hydrobiinae and the genus Mercuria. Little is known about the presence of other hydrobiid subfamilies. This study examines several specimens of gastropods collected from two springs in Tunisia. Based on a comprehensive literature review and rigorous anatomical and molecular comparisons with known species from North Africa, Europe and Asia Minor of similar morphology, we here describe a new genus, Bullaregia gen. nov., and new species Bullaregia tunisiensis gen. et sp. nov. Although the shell shape of this Tunisian species resembles that of European hydrobiid genera such as Belgrandiella, it differs in other anatomical structures (i.e., penis with a glandular strap-like lobe, and two seminal receptacles, SR1 pedunculate, SR2 sessile and less developed). Using maximum likelihood and Bayesian approaches, phylogenetic relationships inferred from mtCOI sequences point to an independent evolutionary lineage for this new taxon outside Belgrandiella and other known hydrobiid subfamilies.
Eight new species of marine dolichopodid flies from southern Thailand belonging to the genus Thinophilus Wahlberg, 1844 are described and illustrated: Thinophilus boonrotpongi sp. nov., T. langkawensis sp. nov., T. minutus sp. nov., T. parmatoides sp. nov., T. parvulus sp. nov., T. spinatus sp. nov., T. spinatoides sp. nov. and T. variabilis sp. nov. A key is provided to the species of the Thai-Malay Peninsula.
The land snail genus Amphidromus Albers, 1850 and its subgenus Syndromus Pilsbry, 1900 (family Camaenidae) in Laos are revised. Fourteen species from various habitats of deciduous forest and limestone areas throughout Laos are confirmed. The classification is revised and discussed, based on material examined from Laos, Cambodia, Vietnam and Thailand and after comparison with the type specimens in many museums. Two new species, A. (Amphidromus) syndromoideus Inkhavilay & Panha sp. nov., from central Laos, and A. (Syndromus) xiengkhaungensis Inkhavilay & Panha sp. nov., from northern Laos, are described and discussed.
Japanese species of the genus Intybia are revised taxonomically, with the examination of the endophallic structure. Eight species, including one new species Intybia donan sp. nov. from Yonagunijima, are recognized. All species are described or redescribed with a key and figures. The endophallic structure contains one primary sclerite (gonoporal piece), three secondary sclerites (ligula, semigonoporal piece, and spinous plate) in some species, and a membranous basal area densely covered with many spines (spinous area). Based on the structures of the endophallus, the Japanese members of the genus are divided into two species groups (the histrio and pelegrini groups). The pelegrini species group is furthermore subdivided into three subgroups (subgroups 1–3). New distributional records are as follows: I. histrio from Hachijô-jima and Tanega-shima; I. niponica from Sakhalin and I. takaraensis from Tokuno-shima and Amami-Ôshima.
With 280 accepted species, the genus Riccardia S.F.Gray (Aneuraceae) is one of the most speciose genera of simple thalloid liverworts. The current classification of this genus is based on morphological and limited-sampling molecular studies. Very few molecular data are available and a comprehensive view of evolutionary relationships within the genus is still lacking. A phylogeny focusing on relationships within the large genus Riccardia has not been conducted. Here we propose the first worldwide molecular phylogeny of the genus Riccardia, based on Bayesian inference and parsimony ratchet analyses of sequences from three plastid regions (psbA-trnH, rps4, trnL-F). The results support the monophyly of Riccardia and a new monospecific genus, Afroriccardia Reeb & Gradst. gen. nov., is described based on molecular and morphological evidence. The results indicate that several currently recognized infrageneric divisions and a few species are not monophyletic, suggesting that further analyses are needed to arrive at a proper understanding of the phylogeny of the genus. Although evidence for an Andean clade was found, most of the species appear scattered in different clades without clear geographical segregation. Broader sampling and further analyses are necessary in order to improve our understanding of the phylogeny of this poorly known liverwort genus.
In Southeast Asia, bats of the genus Tylonycteris Peters, 1872 have traditionally been classified into two wide-ranging species, T. pachypus (Temminck, 1840) and T. robustula Thomas, 1915. Our comparative phylogeographic analyses based on two mitochondrial and seven nuclear genes, combined with our multivariate morphological analyses, show that these species actually represent cryptic species complexes that share a similar biogeographic history in three major regions, i.e., Sundaland, southern Indochina, and northern Indochina. Our molecular dating estimates suggest that Pleistocene climatic oscillations and sea level changes have repeatedly isolated ancestral populations of Tylonycteris spp. in distant bamboo forest refugia. The analyses indicate, however, that populations of the T. pachypus complex were less affected by forest fragmentation in mainland Southeast Asia than those of the T. robustula complex. Accordingly, we propose several taxonomic changes within the genus Tylonycteris: the species T. fulvida and T. malayana are revalidated, and a new species, T. tonkinensis Tu, Csorba, Ruedi & Hassanin sp. nov., endemic to northern Indochina, is described.
In this review (third part), several species within the Nassarius pauperus complex from the eastern Indian Ocean and western Pacific are treated, including a revised concept of Nassa paupera Gould, 1850, type species of the genus Reticunassa Iredale, 1936. In the most recent taxonomic revision, several species had been synonymized with Nassarius pauperus (Gould, 1850), despite distinctive differences among these species in shell morphology. We sequenced a fragment of the mitochondrial COI and the nuclear 28S genes of all available Nassarius pauperus complex species. Automatic barcode gap discovery and reciprocal monophyly were applied to propose species delimitation hypotheses and to support a new concept of the genus. Using morphological and molecular characters, Reticunassa is elevated to full genus rank. Six new species are described: Reticunassa visayaensis sp. nov., R. poppeorum sp. nov., R. annabolteae sp. nov., R. goliath sp. nov., R. intrudens sp. nov. and R. thailandensis sp. nov., while R. tringa (Souverbie, 1864) is recognized as a valid species.
The genus Brachypsectra LeConte, 1874 (Coleoptera: Brachypsectridae) in the Palaearctic Region
(2017)
We summarize all information regarding the genus Brachypsectra (Coleoptera: Brachypsectridae) in the Palaearctic Region. New material of B. kadleci Hájek, 2010 is reported from western and southern Iran and its intraspecific variability in coloration and morphology is described and figured. Brachypsectra jaechi sp. nov. from southern Turkey is described from males, and an unassociated Brachypsectra species from Cyprus is reported, including the descriptions of its female and larva. These findings are the first records of this genus as well as of the family Brachypsectridae for Turkey and Cyprus. Additionally, a diagnostic key to the adult males and larvae of all Brachypsectra species is updated.
The first fossil wedge-shaped beetle (Coleoptera, Rhipiphoridae) from the middle Jurassic of China
(2017)
A new species of Ripiphoridae Gemminger & Harold, 1870, Archaeoripiphorus nuwa gen. et sp. nov., is described and illustrated from a well-preserved impression fossil from the Middle Jurassic Jiulongshan Formation collected at Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China, representing the oldest documented occurrence of the Ripiphoridae described from the Mesozoic era. It shares several characters belonging to two basal ripiphorid subfamilies (Pelecotominae and Ptilophorinae), but it cannot be attributed to either of them and is herein placed as Subfamily incertae sedis. An overall similarity between Archaeoripiphorus gen. nov. and Recent Pelecotominae and the occurrence of wood-boring beetles in the same Formation implies a similar parasitoid host preference in xylophagous beetles for A. nuwa gen. et sp. nov., putting a spotlight on a potential host-parasitoid relationship in the Mesozoic.
The Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Fulgoromorpha, Dictyopharidae, Dictyopharinae, Orthopagini) is revised. Four species are included: C. jocosa (Gerstaecker, 1895) (the type species, with confirmed records from Cameroon, Equatorial Guinea and Gabon), C. lindbergae sp. nov. (described from Sierra Leone), C. rhinoceros sp. nov. (described from Togo) and C. simplex Melichar, 1912 (so far only known from Equatorial Guinea, Bioko island). Lectotypes are designated for C. jocosa and C. simplex and both species are redescribed including habitus photographs and detailed illustrations of the male and female genitalia which are published for the first time. A key for identification of the species of Centromeriana is provided. As far as known, the genus is endemic to the (Guineo-)Congolian region of western Africa.
A new genus is erected within the Cetoniini to describe a newly discovered species with characters shared between Heteroclita Burmeister, 1842, Ichnestoma Gory & Percheron, 1833 and Meridioclita Krikken, 1982. Neoclita pringlei gen. et sp. nov. exhibits a simple clypeal structure without specialized armour, along with hypertrophic and hairy tarsal segments as well as a fully winged female. The new species also exhibits an aedeagal structure closest to Meridioclita, with dorsal lobes of parameres substantially narrower than the ventral ones. The species appears to be restricted to high altitudes in the southwestern peri-Drakensberg area of the Eastern Cape Province, South Africa. Similarly to other mountain relicts known from the southern African region, adults emerge only after major rainfall events during the late spring to early summer season and do not show any evidence of feeding. It appears that flying activity may be temporarily interrupted following soil desiccation, to resume promptly after the next rainfall.
The world’s smallest Begonia, Begonia elachista Moonlight & Tebbitt sp. nov., is described and illustrated from a limestone outcrop in the Amazonian lowlands of Pasco Region, Peru. It is placed within the newly described, monotypic Begonia sect. Microtuberosa Moonlight & Tebbitt sect. nov. and the phylogenetic affinities of the section are examined. Begonia elachista sp. nov. is considered Critically Endangered under the International Union for the Conservation of Nature (IUCN) criteria.
Six species of Anastatus Motschulsky, 1859 (Hymenoptera: Eupelmidae) were described from China in Chinese by J.K. Sheng and coauthors in 1997 and 1998: A. dexingensis, A. flavipes, A. fulloi, A. huangi, A. meilingensis and A. shichengensis. This represents almost half the species of Anastatus recorded from China, but no keys were given to differentiate the species and the original descriptions included only simple line drawings to illustrate the species. Because recognition of these species is critical prior to clarifying the Anastatus fauna of China and of the eastern Palaearctic and Oriental regions, we have redescribed the six species in detail in English based on original type material, illustrating the species through macrophotography of type material and providing a key to differentiate females of the species.
The genus Oxidus Cook, 1911 is revised to contain five species, O. avia (Verhoeff, 1937), O. gigas (Attems, 1953), O. gracilis (C.L. Koch, 1847), O. riukiaria (Verhoeff, 1940), and “species inquirenda” O. obtusus (Takakuwa, 1942). A cosmopolitan species, O. gracilis, is widely found in temperate and sub-tropical regions over the world, but other species have limited distribution in restricted regions, e.g., O. gigas in northern Vietnam, O. riukiaria and O. avia in the Ryukyu Islands (Japan). Four species, O. gracilis, O. riukiaria, O. avia and O. gigas, are confirmed as different from each other in gonopod characters, coloration and body size. The status of the last species, O. obtusus, is still doubtful and requires examination of further fresh material. The phylogenetic relationships among species of Oxidus is analyzed using two fragments of the mitochondrial genes COI (Cytochrome c Oxidase subunit I) and 16S rRNA. Three species of Oxidus are clearly separated from each other; O. gigas and O. gracilis form a monophyletic sister group with O. riukiaria. The genus Oxidus is also monophyletic and more closely related to the genus Tylopus Jeekel, 1968 than to the genera Sellanucheza Enghoff, Golovatch & Nguyen, 2004 or Kronopolites Attems, 1914. In addition, an identification key to species of Oxidus is provided.
Two new species of hangingflies, Terrobittacus rostratus sp. nov. and Terrobittacus angustus sp. nov., are described and illustrated from Yunnan, southwestern China, increasing the species number of Terrobittacus Tan & Hua, 2009 to six. Terrobittacus rostratus sp. nov. differs from its congeners by wings devoid of markings and epandrial appendages slightly longer than half the length of the gonocoxites. Terrobittacus angustus sp. nov. can be recognized by wing markings along R5 distally. A key to species of the genus is updated to include the two new species.
In this work we present a revision of the genus Ommatoiulus Latzel, 1884 in Portugal. Based on recently collected material and older museum samples, including type specimens, we describe six new species to science, viz. Ommatoiulus alacygni sp. nov., O. camurus sp. nov., O. denticulatus sp. nov., O. litoralis sp. nov., O. staglae sp. nov. and O. stellaris sp. nov. The species O. alacygni sp. nov., O. denticulatus sp. nov. and O. staglae sp. nov. described from the Algarve are outstanding by their extremely reduced mesomerital process. The species O. porathi (Verhoeff, 1893) and O. andalusius (Attems, 1927) are recorded and redescribed for the first time after their original description. The finding of O. andalusius – originally described from Andalusia in Spain – constitutes a new record for Portugal together with two species, viz. O. fuentei (Brolemann, 1920) and O. martensi Mauriès, 1969. The taxonomic status of several species is revised. Thus Archiulus (Schistocoxitus) cingulatus Attems, 1927 is here considered as a junior synonym of Ommatoiulus lusitanus (Verhoeff, 1895) while Schizophyllum cervinum Verhoeff, 1910 is synonymized with Ommatoiulus moreleti (Lucas, 1860). An identification key to all hitherto known Portuguese species of Ommatoiulus is presented as well as a distribution map illustrating the various species occurrences in the country.
The genus Austrotinodes contains 55 species, largely distributed in the Neotropics, with a few species occurring in the Australian Region. In Brazil, 10 species have been recorded, mainly in the southern and southeastern regions. Herein, we describe 13 new species from Brazil, all of them named in memory of great Brazilian scientists: Austrotinodes absaberi sp. nov., A. adolfolutzi sp. nov., A. berthalutzae sp. nov., A. chagasi sp. nov., A. costalimai sp. nov., A. cruzi sp. nov., A. donagrazielae sp. nov., A. gusmaoi sp. nov., A. lattesi sp. nov., A. lenti sp. nov., A. santosdumonti sp. nov., A. vanzolinii sp. nov. and A. vitalbrazili sp. nov. Additionally, we give new distribution records for A. amazonensis Flint & Denning, 1989, A. longispinum Thomson & Holzenthal, 2010, A. paraguayensis Flint, 1983 and A. taquaralis Thomson & Holzenthal, 2010. We also provide new diagnoses and illustrations for A. amazonensis and A. paraguayensis, to facilitate identification of those species.
Nomenclatural changes are made in three previously described genera in the planthopper tribe Hemisphaeriini (Hemiptera: Fulgoromorpha: Issidae: Issinae), viz Gergithus Stål, 1870, Mongoliana Distant, 1909 and Hemisphaeroides Melichar, 1903. In addition, a new genus, Gnezdilovius gen. nov., with Gergithus lineatus Kato, 1933 as its type species, is described for 40 species formerly included in Gergithus, and the generic characteristics of the latter genus is revised. One new species, Gergithus frontilongus sp. nov. from China (Yunnan), is described and illustrated. One additional Gergithus species, previously misidentified as G. signatifrons Melichar, 1906 from Siberut Island, is mentioned and illustrated. Gergithus contusus Walker, 1851 is transferred to Mongoliana and Hemisphaerius atromaculatus Distant, 1916 and H. fuscoclypeatus Distant, 1916 are transferred to Hemisphaeroides. Checklists for all four genera are provided detailing the nomenclatural changes and a key to the 19 genera of Hemisphaeriini is provided. Morphological diversity and distribution of the genera are briefly discussed.
Two new species of the Rhyacophila nigrocephala species group, R. voluta sp. nov. and R. linguiformis sp. nov., are described, diagnosed and illustrated. Rhyacophila voluta sp. nov. is similar to R. pentagona Malicky & Sun, 2002 in male genitalia, but can be diagnosed by the inferior appendages of the same length as the complex of preanal appendages and dorsal lobe of segment IX, fused base of the basal segments of inferior appendages, and slightly incised distal margin of apical segment of inferior appendage. Rhyacophila linguiformis sp. nov. is similar to R. rima Sun & Yang, 1995 and R. esorima Mey, 1996 in male genitalia, but can be diagnosed by the progressively narrowed complex of preanal appendages and the dorsal lobe of segment X, the short anal sclerites, and the small gap between the upper and lower lobes of apical segment of inferior appendages. An updated checklist of 23 species recorded from China is presented.
Review of the clavatus group of the lanternfly genus Pyrops (Hemiptera: Fulgoromorpha: Fulgoridae)
(2017)
The clavatus group of Pyrops Spinola, 1839 is reviewed and redefined. The new combination Pyrops atroalbus (Distant, 1918) comb. nov. is proposed, as atroalbus is reinstated as a full species from status of subspecies of Pyrops watanabei (Matsumura, 1913). Pyrops nigripennis (Chou & Wang, 1985) and Pyrops clavatus mizunumai (Sato & Nagai, 1994) are proposed as junior synonyms of P. clavatus (Westwood, 1839). The Philippine species P. polillensis (Baker, 1925) is removed from the group and not attributed to any of the currently defined species groups. Hence, the clavatus group is restricted to continental Southeast Asia and Taiwan and contains three species: P. atroalbus comb. nov., P. clavatus and P. watanabei. A key to the species of the group and illustrations of the male genitalia are provided. The intraspecific colour variation in the group is discussed and illustrated. The genus Pyrops is removed from the subfamily Fulgorinae and not attributed to any of the currently defined subfamilies of Fulgoridae.
Two new species of the genus Gergithoides Schumacher, 1915 (Issinae, Hemisphaeriini), G. gnezdilovi sp. nov. from Bidoup-Nui Ba National Park in Central Vietnam and G. nui sp. nov. from Pia-Oac National Park in North Vietnam, are described. These are the only species of the genus formally recorded from Vietnam to date. Habitus, details and male genitalia are illustrated and a distribution map is provided. Four females representing three or four additional species, known from females only, are mentioned and illustrated. Taxonomic and biogeographical updates based on a thorough review of the literature are proposed and discussed for G. carinatifrons Schumacher, 1915, G. rugulosus (Melichar, 1906) and G. undulatus Wang & Che, 2003.
The genus Otraleus Günther, 1935 is recorded from the Philippines for the first time. Four new species, Otraleus bellemansae sp. nov., O. applai sp. nov., O. christianae sp. nov. and O. elizabethae sp. nov., are described from the highlands of Northwestern Luzon. The characters allowing separation from O. hypsimelathrus Günther, 1935 and O. labanrataensis Soew-Choen, 2016, are given. A new genus closely related to Otraleus, Capuyanus gen. nov., is described with a single species, C. magwilangi sp. nov., as type-species. An identification key and distribution maps are provided for all species.
Five new species of Bakerius Bondar (Hemiptera: Aleyrodidae: Aleurodicinae) are described and illustrated from the Americas and Vietnam based on the adult, nymph, and pupal stages: Bakerius asiaticus, Bakerius colombianus, Bakerius hondurensis, Bakerius leei and Bakerius peruvianus. The following six species: Bakerius attenuatus Bondar 1923, Bakerius calmoni Bondar 1928, Bakerius marmoratus (Hempel 1923), Bakerius phrygilanthi Bondar 1923, Bakerius sanguineus Bondar 1928, and Bakerius sublatus Bondar 1928 are re-described. An identification key to the New World genera of the subfamily Aleurodicinae, and a key to the adults and the puparia of Bakerius species are provided.
Distributional records of the 30 tiger beetle species and subspecies (Coleoptera: Carabidae: Cicindelinae) known for Brunei Darussalam are given together with habitus photos for 21 species. Neocollyris (Neocollyris) labiomaculata (Horn, 1892), Neocollyris (Neocollyris) emarginata (Dejean, 1825), Therates spectabilis fl avissimus Brouerius van Nidek, 1957, Heptodonta analis s. str. (Fabricius, 1801), Cosmodela velata (Bates, 1872), Lophyra (s. str.) fuliginosa (Dejean, 1826), Cylindera (Leptinomera) fi ligera (Bates, 1878), Myriochila (s. str.) specularis brevipennis (Horn, 1897), Abroscelis tenuipes araneipes (Schaum, 1863) and Callytron doriai (Horn, 1897) are reported for the fi rst time for the Sultanate.
Distributional records of the 177 tiger beetle species and subspecies known for Vietnam are given together with a key and habitus photos (Coleoptera: Carabidae: Cicindelinae). Tricondyla (Tricondyla) macrodera abruptesculpta Horn, 1925, Protocollyris festiva Naviaux, 2008, Neocollyris (Brachycollyris) purpureomaculata borea Naviaux, 1994, Neocollyris (Brachycollyris) torosa Naviaux, 2010, Neocollyris (Leptocollyris) rogeri Shook and Wu, 2006, Neocollyris (Leptocollyris) laosensis Naviaux, 1999, Neocollyris (Stenocollyris) rubens (Bates, 1878), Neocollyris (Pachycollyris) tricolor Naviaux, 1991, Neocollyris (Pachycollyris) panfi lovi Naviaux and Matalin, 2002, Therates rugulosus Horn, 1900, Therates bannapecolus Sawada and Wiesner, 1999, Calochroa interruptofasciata fl avolineata (Chaudoir, 1865), Cylindera (Ifasina) cyclobregma Acciavatti and Pearson, 1989, Cylindera (Ifasina) decolorata (Horn, 1907), Cylindera (Ifasina) humillima (Gestro, 1893) and Cylindera (Eugrapha) elisae elisae (Motschulsky, 1859) are reported for the fi rst time in the fauna of Vietnam. Neocollyris (Isocollyris) septentrionalis Naviaux, 1999, Neocollyris (Leptocollyris) linearis tenuicornis (Chaudoir, 1864), Neocollyris (Pachycollyris) aptera apicalis (Chaudoir, 1864), Neocollyris (Pachycollyris) contracta (Horn, 1905), Therates dimidiatus wallacei Thomson, 1857, Cylindera (Cylindera) pseudocylindriformis Horn, 1913, Cylindera (Ifasina) viridilabris (Chaudoir, 1852), Cylindera (Ifasina) psilica psilica (Bates, 1866), Cylindera (Eugrapha) venosa (Kollar, 1836) and Myriochila (Myriochila) undulata (Dejean, 1825) are excluded from the fauna of Vietnam
Two new genera and species of tiger beetles from Baltic amber (Coleoptera: Carabidae: Cicindelinae)
(2017)
Two fossil tiger beetle species (Coleoptera, Carabidae, Cicindelinae) are described from Eocene Baltic amber using light microscopic and X-ray microscopic techniques. Both species are considered representatives of the subtribe Iresina Rivalier, 1971 due to the shared combination of character states: glabrous head, six labral and four suborbital setae, and glabrous pronotum. Palaeopronyssiformia groehni Wiesner, Will, and Schmidt, new genus, new species, is characterized by a glabrous and furrowed head with six labral setae, large eyes, presence of two supraorbital setae on each side, mandibles with two teeth of the incisor region, and a glabrous and furrowed pronotum. Palaeoiresina cassolai Wiesner, Will, and Schmidt, new genus, new species, is characterized by a unicolored, undentated labrum, mandibles with two teeth of the incisor region, glabrous head with six labral setae, two clypeal setae, two supraorbital setae on each side, and a glabrous pronotum, mesepisternum, mesepimeron, and metepisternum. The species described here represent the only known tiger beetle fossils preserved in Baltic amber.
Weimarer Beiträge 63/2017
(2017)
Die Weimarer Beiträge sind eine Zeitschrift für Literaturwissenschaft, aktuelle ästhetische Theorie und Kulturwissenschaft. Zu Ihren Schwerpunkten gehören moderne Literatur im Rahmen anderer Künste und Medien, die Wechselbeziehungen von Literatur, philosophischer und ästhetischer Reflexion sowie die kritische Analyse der Gegenwartskultur.
The Antillean fauna of Mantispidae (Neuroptera) consists of six species in four genera, all in the subfamily Mantispinae. Two new species are here described: Leptomantispa antillesensis Hoffman, n. sp., is described from Puerto Rico, Anguilla, British Virgin Islands (Grand Camanoe, Guana, Tortola), United States Virgin Islands (St. Croix, St. John, St. Thomas), Dominica, and Guadeloupe, and Leptomantispa hispaniolaensis Hoffman, n. sp. is described from Hispaniola (Dominican Republic). Mantispilla zayasi Alayo is synonymized under Leptomantispa pulchella (Banks), new synonomy, and Mantispilla taina Alayo is synonymized under Zeugomantispa minuta (Fabricius), new synonomy. Climaciella cubana Enderlein is recorded for the fi rst time from Hispaniola (Dominican Republic). Dicromantispa sayi (Banks) is newly reported from the Cayman Islands (Cayman Brac and Little Cayman), and Hispaniola (Dominican Republic). Zeugomantispa minuta (Fabricius) is now recorded from the Bahamas (Andros) and Hispaniola (Dominican Republic).
From a moss sample collected in the Manabí Province in Ecuador, we extracted 96 specimens of a new species of eutardigrade. No eggs were found. In order to obtain eggs, an in vitro culture was prepared. In total, 136 specimens (including ten simplex), one exuvia and 44 eggs (including two with embryos) of the new species were obtained. In addition to the traditional taxonomic description with morphometrics, light and scanning microscopy imaging, we also provide nucleotide sequences of three nuclear (18S rRNA, 28S rRNA, ITS-2) and one mitochondrial (COI) DNA fragments of the new species. Macrobiotus polypiformis sp. nov. belongs to the hufelandi group and is most similar to Ma. paulinae Stec, Smolak, Kaczmarek & Michalczyk, 2015, but differs from it mainly by the lack of dorso-lateral patches of granulation on the cuticle, egg processes with longer and more numerous filaments and in some morphometric characters of both eggs and adults. Moreover, we provide a short discussion on the modifications/abnormalities of the claws in eutardigrades and the possible consequences on the taxonomic status of Mesobiotus armatus (Pilato & Binda, 1996), suggesting its consideration as species inquirenda (with uncertain taxonomic status).
Three new fossil bryozoan species, a ctenostome and two cheilostomes, are described and figured from Pleistocene strata of the Wanganui Basin, New Zealand. Buskia waiinuensis sp. nov., a soft-body ctenostome preserved as a mould bioimmuration, is the first fossil record of the genus from New Zealand. Microporella rusti sp. nov., which is notable for the lack of ooecia in the large suite of colonies available, is one of the most common bryozoans in the Nukumaru Limestone and Nukumaru Brown Sand shellbeds, forming large encrusting sheet-like colonies, but is uncommon in younger beds. Rare, small-sized colonies of Parkermavella columnaris sp. nov. were found as fossil in two Quaternary beds, the Nukumaru Limestone and the Upper Kai-Iwi Shellbed, and also alive on rocks from greater Cook Strait area.