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The present study aims to resolve the taxonomic confusion involving several taxa within Mycetophagidae Leach, 1815, originating from the introduction of the genus Atritomus Reitter, 1877, and then by its subsequent controversial interpretation. A detailed overview of the taxonomic and nomenclatural history of the taxa previously linked to Atritomus is provided. The authors propose the introduction of Stereophilus Biscaccianti, Audisio & Esser gen. nov. for Atritomus filicornis Reitter, 1887, and the restoration of Entoxylon Ancey, 1869 at the genus rank, together with some rectifications regarding the authorship and the date of publication of both Entoxylon and its type species, E. abeillei Ancey, 1869. Moreover, the Ethiopian species Atritomus vicinus Grouvelle, 1908 is herein transferred to the genus Typhaeola Ganglbauer, 1899 based on the examination of the holotype. The following new combinations are proposed: Entoxylon baudii (Seidlitz, 1889) comb. nov. (from Esarcus Reiche, 1864), Entoxylon besucheti (Dajoz, 1964) comb. nov. (from Esarcus subg. Entoxylon), Entoxylon franzi (Dajoz, 1964) comb. nov. (from Esarcus subg. Entoxylon), Entoxylon inexpectatus (Dajoz, 1964) comb. nov. (from Esarcus subg. Entoxylon), Entoxylon martini (Reitter, 1887) comb. nov. (from Esarcus), Stereophilus filicornis (Reitter, 1887) gen. et comb. nov. (from Atritomus), Typhaeola vicina (Grouvelle, 1908) comb. nov. (from Atritomus).
Umbyquyra gen. nov., a new Theraphosinae genus with stridulatory bristles on the palpal trocanther of pedipalp trochanter and first leg, is proposed. The genus differs from the other genera with stridulatory bristles on the same segments, Acanthoscurria Ausserer, 1871, Cyrtopholis Simon, 1892, Longilyra Gabriel, 2014 and Nesipelma Schmidt & Kovarik, 1996, by having a palpal bulb with a very short and acuminate embolus and four short keels; separated tibial apophysis; and female spermathecae resembling those of Cyrtopholis, with two seminal receptacles with elongated ducts emerging from a common area. Cyrtopholis palmarum Schiapelli & Gerschman, 1945 and C. schmidti Rudloff, 1996 from Brazil and Acanthoscurria acuminata Schmidt & Tesmoingt in Schmidt, 2005 from Bolivia are transferred to the new genus. The female of Umbyquyra palmarum (Schiapelli & Gerschman, 1945) gen. et comb. nov. and the male of U. schmidti (Rudloff, 1996) gen. et comb. nov. are described for the first time. Cyrtopholis zorodes Mello-Leitão, 1923 is considered a junior synonym of Acanthoscurria gomesiana Mello-Leitão, 1923 and Cyrtopholis meridionalis (Keyserling, 1891) is considered a nomen dubium. Eight new species from Brazil are described: Umbyquyra paranaiba gen. et sp. nov., U. cuiaba gen. et sp. nov., U. araguaia gen. et sp. nov., U. sapezal gen. et sp. nov., U. belterra gen. et sp. nov., U. caxiuana gen. et sp. nov., U. tucurui gen. et sp. nov. and U. tapajos gen. et sp. nov. Data and maps on the geographic distribution are provided.
Two new genera and species of tiger beetles from Baltic amber (Coleoptera: Carabidae: Cicindelinae)
(2017)
Two fossil tiger beetle species (Coleoptera, Carabidae, Cicindelinae) are described from Eocene Baltic amber using light microscopic and X-ray microscopic techniques. Both species are considered representatives of the subtribe Iresina Rivalier, 1971 due to the shared combination of character states: glabrous head, six labral and four suborbital setae, and glabrous pronotum. Palaeopronyssiformia groehni Wiesner, Will, and Schmidt, new genus, new species, is characterized by a glabrous and furrowed head with six labral setae, large eyes, presence of two supraorbital setae on each side, mandibles with two teeth of the incisor region, and a glabrous and furrowed pronotum. Palaeoiresina cassolai Wiesner, Will, and Schmidt, new genus, new species, is characterized by a unicolored, undentated labrum, mandibles with two teeth of the incisor region, glabrous head with six labral setae, two clypeal setae, two supraorbital setae on each side, and a glabrous pronotum, mesepisternum, mesepimeron, and metepisternum. The species described here represent the only known tiger beetle fossils preserved in Baltic amber.
Synchroidae Lacordaire, 1859 is a taxonomically and biologically poorly known group. In the present paper, diagnostic characters used to separate genera are analysed and the phylogenetic relationships within this family are preliminarily investigated. Results suggest that the characteristic Synchroa pangu Hsiao, Li, Liu & Pang, 2016 can be removed to establish a new genus, Thescelosynchroa gen. nov. The new combination, T. pangu (Hsiao, Li, Liu & Pang) gen. et comb. nov., is proposed. The definitions of Synchroa Newman, 1838 and Synchroina Fairmaire, 1898 are revised. Moreover, morphological analysis and character comparison also suggest that the familial placement of Mallodrya subaenea Horn, 1888 is questionable. Six species are re-examined and rediagnosed: Synchroa chinensis Nikitsky, 1999, S. elongatula Nikitsky, 1999, S. formosana Hsiao, 2015, S. melanotoides Lewis, 1895, S. punctata Newman, 1838 and Synchroina tenuipennis Fairmaire, 1898. The male of S. chinensis and the female of S. formosana are described for the first time. Synchroa elongatula and Synchroina tenuipennis are newly recorded from Laos and Indonesia, respectively. We also hypothesize that the Eastern Asian-North American disjunction of Synchroa could be connected to a Mid-Late Tertiary migration of plants via the Bering Land Bridge.
The Egyptian fauna of mealybugs (Hemiptera: Coccoidea: Pseudococcidae) is reviewed and an illustrated key to the 30 genera and 54 species is provided. Phenacoccus madeirensis Green is reported for the first time in Egypt. A new genus, Ezzatacoccus Evans and Abd-Rabou, is described and illustrated with Amonostherium arabicum Ezzat, 1960 designated as its type species. Octococcus salicicola Priesner and Hosny, 1935 is reinstated as a valid taxon and transferred to Misericoccus Ferris, new combination. Ripersia cressae Hall is transferred to Maconellicoccus Ezzat, new combination and Planococcus lindingeri (Bodenheimer) is transferred back to Formicococcus Takahashi, revised status.
ZooBank registration. urn:lsid:zoobank.org:pub:8CA7B000-E8D4-463D-95B0-431BA0A7BA57
The evolution and interrelationships of carnivorous squamates (mosasaurs, snakes, monitor lizards, Gila Monsters) are a contentious part of reptile systematics and go to the heart of conflict between morphological and molecular data in inferring evolutionary history. One of the best-preserved fossils in this motley grouping is “Saniwa” feisti Stritzke, 1983, represented by complete skeletons from the early-middle Eocene of Messel, Germany. We re-describe it on the basis of superficial examination, stereoradiography, and high-resolution X-ray computed tomography of new and published specimens. The scalation of the lizard is unique, consisting of small, keeled scales on the head (including a row of enlarged medial supraorbitals) and large, rhomboidal, keeled scales (invested by osteoderms) that covered the rest of the body. Two paired longitudinal rows of enlarged scales ran down the neck. The head was laterally compressed and box-shaped due to the presence of a strong canthal-temporal ridge; the limbs and tail were very long. Notable osteological features include: a toothed, strap-like vomer; septomaxilla with a long posterior process; palpebral with a long posterolateral process; a lacrimal boss and a single lacrimal foramen; a well-developed cultriform process of the parabasisphenoid; two hypoglossal (XII) foramina in addition to the vagus; a lack of resorption pits for replacement teeth; and possibly the presence of more than one wave of developing replacement teeth per locus. There are no osteological modifications suggestive of an intramandibular hinge, but postmortem displacement of the angular-prearticular-surangular complex in multiple specimens suggests that there might have been some degree of mobility in the lower jaw based on soft-tissue modifications. Using phylogenetic analyses on a data-set comprising 473 morphological characters and 46 DNA loci, we infer that a monophyletic Palaeovaranidae Georgalis, 2017, including Eosaniwa Haubold, 1977, lies on the stem of Varanidae Merrem, 1820, basal to various Cretaceous Mongolian taxa. We transfer feisti to the new genus Paranecrosaurus n. gen. Analysis of gut contents reveals only the second known specimen of the cryptozoic lizard Cryptolacerta hassiaca Müller, Hipsley, Head, Kardjilov, Hilger, Wuttke & Reisz, 2011, confirming a diet that was at least partly carnivorous; the preservation of the teeth of C. hassiaca suggests that the gastric physiology of Paranecrosaurus feisti (Stritzke, 1983) n. comb. had high acidity but low enzyme activity. Based on the foregoing and linear discriminant function analysis, we reconstruct P. feisti n. comb., as a powerful, widely roaming, faunivorous-carnivorous stem monitor lizard with a sensitive snout. If the molecular phylogeny of anguimorphs is correct, then many of the features shared by Helodermatidae Gray, 1837 and Varanidae must have arisen convergently, partly associated with diet. In that case, a reconciliation of morphological and molecular data would require the discovery of equally primitive fossils on the helodermatid stem.
The adult stage of Helioandesia tarregai gen. et sp. nov. (Lepidoptera: Yponomeutoidea: Heliodinidae) is described and illustrated from the arid western slopes of the Andes of northern Chile. The larvae of H. tarregai gen. et sp. nov. feed as leaf skeletonizers on Mirabilis acuta (Reiche) Heimerl (Nyctaginaceae). The mostly gray forewing of H. tarregai gen. et sp. nov., ornamented with strongly bulging metallic spots, resembles that of the representatives of the mainly Nearctic Lithariapteryx Chambers, 1876. However, the latter lacks CuP in the forewing, has a single bristle in the female frenulum, and lacks a well-developed cornutus. Helioandesia gen. nov. clustered as sister to Neoheliodines Hsu, 2004 in a cladistic analysis, although no synapomorphies were found for this cluster, while Lithariapteryx was sister to Helioandesia gen. nov. + Neoheliodines based on two synapomorphies. The genetic distance between a DNA barcode sequence of H. tarregai gen. et sp. nov. and representatives of other genera of Heliodinidae Heinemann, 1877 was 9.0–12.5% (K2P), and a maximum likelihood analysis based on this molecular marker confirmed the placement of H. tarregai gen. et sp. nov. as a member of this micromoth family. This contribution represents the first confirmed record of Heliodinidae for Chile.
Sundapyrochroa, a new genus of pyrochroine Pyrochroidae, is described from three Sunda Shelf species most recently assigned to Pseudopyrochroa Pic: Sundapyrochroa atricolor (Pic), comb. nov., recorded from Peninsular Malaysia, Sundapyrochroa nigripennis (Pic), comb. nov., recorded from Peninsular Malaysia, and Sundapyrochroa sumatrensis (Pic), comb. nov., recorded from Borneo (Malaysia: Sabah) and Sumatra (Indonesia). Schizotus rotundicollis Pic, syn. nov., is proposed as a junior synonym of S. sumatrensis. No evidence could be found to support subspecifi c categories for S. nigripennis, thus, Pseudopyrochroa nigripennis notaticeps Pic, syn. nov., is proposed as a junior synonym of S. nigripennis. Gross anatomy of the cranium (female, male), antennae (female, male), and genitalia (male) are both diagnostic and enigmatic, suggesting no clear relationships with other pyrochroine genera.
In this study, the new genus Spanglerelmis Polizei & Bispo is described; Microcylloepus ochus Hinton, 1940 is synonymized with Microcylloepus femoralis Hinton, 1940, and transferred to the new genus; and two new species, S. xiririca gen. et sp. nov. and S. timburi gen. et sp. nov. are described. The new genus can be characterized by the combination of the following characters: 1) pronotum without transverse, longitudinal or oblique impressions, sulci or gibbosities on disc; 2) elytra with a carina on interval III and two sublateral carinae on intervals V and VI; 3) mesoventrite with sides strongly raised; and 4) femora with an oblique belt of tomentum dorsally and a transverse belt ventrally. The specimens of the two new species were collected mainly in riffles of unimpacted streams in the Atlantic Forest in São Paulo State, Brazil. Scanning electron microscope images, an identification key for the genus and habitat notes are also presented.
Seven new euptychiine (Lepidoptera: Nymphalidae: Satyrinae) taxa are described and named herein, namely Harjesia argentata Nakahara, Zacca and Lamas, n. sp., Orotaygetis Nakahara and Zacca, n. gen., O. surui Nakahara, Zacca and Lamas, n. sp., Euptychoides sanmarcos Nakahara and Lamas, n. sp., Pseudeuptychia cuzquenya Nakahara and Lamas, n. sp., P. languida austrina Nakahara and Lamas, n. ssp., and Godartiana astronesthes Lamas and Nakahara, n. sp. A revisional note is provided for Harjesia Forster, 1964 and Pseudeuptychia Forster, 1964, and as a result, Taygetis vrazi Kheil, 1896 is removed from Harjesia and a new taxonomic arrangement, Pseudodebis vrazi n. comb., is proposed based on both morphology and molecular data.
The weevil genus Cryptolarynx Van Schalkwyk, 1966 is endemic to the Northern and Western Cape provinces of South Africa. The two previously known species of the genus, C. vitis (Marshall, 1957) and C. estriatus (Marshall, 1957), have an aberrant globular body and head shape, which has made it difficult to place the genus into the classification systems of the Curculionoidea. This paper presents the description of 21 new species of Cryptolarynx from South Africa (C. subglaber Haran sp. nov., C. squamulatus Haran sp. nov., C. muellerae Haran sp. nov., C. hirtulus Haran sp. nov., C. robustus Haran sp. nov., C. namaquanus Haran sp. nov., C. carinatus Haran sp. nov., C. variabilis Haran sp. nov., C. pyrophilus Haran sp. nov., C. pilipes Haran sp. nov., C. armatus Haran sp. nov., C. falciformis Haran sp. nov., C. oberprieleri Haran sp. nov., C. spinicornis Haran sp. nov., C. cederbergensis Haran sp. nov., C. homaroides Haran sp. nov., C. marshalli Haran sp. nov., C. endroedyi Haran sp. nov., C. oberlanderi Haran sp. nov., C. san Haran sp. nov., and C. luteipennis Haran sp. nov.) and of one new genus and species, Hadrocryptolarynx major Haran gen. et sp. nov., also from South Africa. A redescription of the genus Cryptolarynx is provided to incorporate the characters of the new species. The plant genus Oxalis (Oxalidaceae) is recorded as larval host for several species of Cryptolarynx and for Hadrocryptolarynx Haran gen. nov., as their larvae develop in the subterranean bulbs of members of the genus, and the egg, larva and pupa of C. variabilis are described. The characters of the Cryptolarynx larva confirm that Cryptolaryngini are an early-diverging group of Curculionidae, with a placement among taxa currently classified in the subfamily Brachycerinae sensu lato, and although their exact taxonomic position remains unresolved, some larval characters, and also pupal ones, suggest a close relationship between Cryptolaryngini and Stenopelmus Schoenherr. Potential use of species of Cryptolarynx in the biological control of weedy South African species of Oxalis is discussed.
This paper revises the genus Ganelius Benesh, which is endemic to Madagascar, in the stag beetle tribe Figulini Burmeister (Coleoptera: Lucanidae: Lucaninae). Species in the genus are striking in their highly sexually dimorphic mandibles, a character that rarely occurs in the tribe. The genus was previously comprised of four species, but three of these names were problematic or questionably applied. A lectotype is designated for G. madagascariensis Laporte, a neotype is designated for Ganelius oberndorferi Nonfried, and Nigidius passaliformis Benesh is newly synonymized with G. oberndorferi. Two new species, G. gnamptus Paulsen and G. zombi Paulsen, are described from western Madagascar. The identity of the overlooked Ganelius nageli (Kriesche) is fixed through a neotype designation, and the species is moved to the new genus Agnelius, which is distinguished from Ganelius by a lack of sexual dimorphism, serrate protibiae, and a more flattened body.
Chondrocyclus Ancey, 1898 is a genus of nine species of African operculate land snails restricted to indigenous forest and mesic thicket. Worn specimens (i.e., without a periostracum or operculum), on which some species descriptions and records were based, appear to be indistinguishable morphologically. A comprehensive revision of Chondrocyclus s.l. is provided here based on comparative morphological examinations of the shell, protoconch, periostracum, operculum, radula and penis, and on mitochondrial genes cytochrome c oxidase subunit I and 16S rRNA. Two genus-level lineages are recognised, Chondrocyclus s.s. and Afrocyclus gen. nov. Revised species descriptions are given for seven species. Two species, C. meredithae Bruggen, 1983 and C. chirindae Bruggen, 1986 both from north of South Africa, are removed from Chondrocyclus. Twelve new species are described: C. herberti sp. nov., C. silvicolus sp. nov., C. amathole sp. nov., C. pondoensis sp. nov., C. devilliersi sp. nov., C. pulcherrimus sp. nov., C. cooperae sp. nov., C. langebergensis sp. nov., C. kevincolei sp. nov., A. oxygala gen. et sp. nov., A. potteri gen. et sp. nov. and A. bhaca gen. et sp. nov. This is the first detailed systematic revision of an Afrotropical cyclophorid group to include morphological and molecular data. This study complements research on other taxa of low-vagility forest-dwelling habitat specialists by providing comparative distribution data for an independent, widespread group. Such evidence is urgently needed for conservation of South Africa’s threatened forest biome.
A taxonomic review was conducted of the type material of Quedius rove beetles (including Indoquedius, previously a subgenus) described by Otto Scheerpeltz from the 1934 Swedish expedition by René Malaise to Kambaiti, Myanmar. The specimens were mistakenly thought to be lost or compromised during the Second World War, and so the corresponding available names were not considered in the extensive taxonomic study of Quedius from the Himalayan Region and mainland China that has followed. This has resulted in the following synonyms: Indoquedius nonparallelus Zhao & Zhou, 2010 syn. nov. = I. malaisei (Scheerpeltz, 1965); I. baliyo Smetana, 1988 syn. nov. = I. micantiventris (Scheerpeltz, 1965); I. sanguinipennis (Scheerpeltz, 1965) syn. nov., I. bicoloris Smetana, 2014 syn. nov. = I. parallelicollis (Scheerpeltz, 1965); Quedius cornutus Cai et al., 2015 syn. nov. = Q. rutilipennis Scheerpeltz, 1965; Q. sundar Smetana, 1988 syn. nov., Q. hecato Smetana, 2012 syn. nov. = Q. semilaeviventris Scheerpeltz, 1965; Q. kambaitiensis Scheerpeltz, 1965 syn. nov. = Q. muscicola Cameron, 1932. The species collected by Malaise were treated within the most recent phylogenetic context, resulting in Malaisdius gen. nov., M. ruficeps (Scheerpeltz) comb. nov., and new or revised morphological concepts for the Apicicornis and Masasatoi species groups of Quedius (Microsaurus), and the Muscicola group of Quedius (Raphirus). Malaisdius smetanai gen. et sp. nov. is described from Nepal.
Redescription of some species of Bryozoa described by J. Jullien and L. Calvet in the NE Atlantic
(2015)
Five species described by Jullien and/or Calvet from the NE Atlantic are redescribed and stabilized by typification: Hippothoa amoena, Schizoporella confusa, S. jullieni, S. ovum and Smittia guernei. Three new synonymies are established: Schizoporella jullieni with Hippothoa amoena, S. ovum with Escharina alderi, and Escharella pseudopunctata with Smittia guernei. A new trypostegid genus, Pulpeirina gen. nov., is erected for H. amoena. Three new combinations are introduced.
Pyroghatsiana, a new genus of pyrochroine Pyrochroidae is described from the Southern Ghats of the Indian continental southern tip. The only known specimen is a female, Pyroghatsiana madurensis (Pic), new combination, originally placed in Dendroides Latreille, and subsequently transferred to Pseudodendroides Blair. Several striking differences including the dorsal interocular width between the compound eyes, shape and length of the third antennal segment, and shape of the pronotum preclude placement of Pyroghatsiana in either Dendroides, Pseudodendroides, or any other existing pyrochroine genus.
A new genus with a new species of soft-winged flower beetle, Pectotibialis paghmanensis Tshernyshev gen. et sp. nov. are described from Afghanistan. The new genus can be distinguished from the congeners of the tribe Apalochrini by the dark pectination in the apices of tibiae in both sexes, and the anterior tibiae which are hollowed at distal half, flattened and rounded femora, bituberculate basal parts of head and pronotum, two lamellate metathoracic appendages, tarsal comb above second tarsomere of anterior legs, and serrate antennae in the male. Based on the metathoracic appendages and comb in anterior legs would attribute this new species to the new genus Dromanthomorphus Pic, 1921, but all the other above-mentioned characters define its independent status and the designation of a new genus; Pectotibialis Tshernyshev gen. nov. The external appearance, special male characters and genitalia of the type species of the new genus are illustrated, and a distribution map is provided. A key to the Apalochrus-section of the tribe Apalochrini is provided.
After our taxonomic revision of Ootheca Chevrolat, 1837, and the description of Oothecoides Kortenhaus & Wagner, 2011 and Ootibia Kortenhaus & Wagner, 2012, it became clear that a further four galerucine species, closely related to the above named taxa, form a distinct monophyletic group, that constitutes a new genus, Oosagitta gen. nov. with O. anningae sp. nov., O. geescheae sp. nov., O. melanopicta sp. nov. and O. thomasi sp. nov.. Exosoma angolensis Laboissière, 1939, the type species of the new genus, and Ergana minuta Laboissière, 1937 are newly transferred to Oosagitta gen. nov. All species of Oosagitta gen. nov. are characterized by a broad body and pronotum, a more or less convex dorsum and short legs, and as such are most similar to the other above named genera. The antennae of Oosagitta gen. nov. are distinctly longer than those of Ootheca, Oothecoides and Ootibia. Genital structures of the males allow a reliable identifi cation of the genus. (Re-) descriptions are given for all species, including semi-schematic illustrations depicting the habitus outline, shape of the basal antennomeres and the median lobe. Photographs of the name-bearing types and distribution maps are provided.
Three subterranean leptodirine leiodid taxa, viz., Bozidaria Ćurčić & Pavićević gen. nov., Bozidaria serbooccidentalis Ćurčić & Pavićević gen. et sp. nov. and Proleonhardella (Proleonhardella) tarensis Ćurčić & Pavićević sp. nov., are described and diagnosed. Bozidaria Ćurčić & Pavićević gen. nov. belongs to the phyletic series of “Leonhardella”. The new beetle taxa differ from their closest relatives in numerous morphological characters. They most likely belong to phyletic lineages of Pliocene age. The new leiodid taxa are endemic to the Dinaric mountain chain of western Serbia. Keys to the leptodirine leiodid genera of the phyletic series of “Leonhardella” and to the taxa of the genus Proleonhardella Jeannel, 1910 are included.
This paper reviews the genus Microtachycines and establishes a new genus, Megatachycines gen. nov. One new species and two new combinations of the genus Megatachycines are recorded, i.e., Megatachycines pentus gen. et sp. nov., Megatachycines elongatus (Qin, Liu & Li, 2017) comb. nov., and Megatachycines trispinosus (Qin & Li, 2020) comb. nov. Images illustrating the morphology of all species of the genera Microtachycines and Megatachycines are provided.
Many nomenclatural changes are implemented in the beetle families Georissidae, Histeridae, Hydraenidae, Hydrochidae, Hydrophilidae, Ptiliidae, Leiodidae and especially Staphylinidae, of the beetle series Staphyliniformia (Coleoptera), in preparation for making a world catalog of this group available online. Limited taxonomic changes are also made in the staphylinid subfamilies Osoriinae and Staphylininae.
At the level of family-group taxa, Article 29.4 of the current (1999) Zoological Code is reviewed and the original spellings of two tribal names, Nymphisterini Tishechkin (Histeridae) and Cryptonotopsisini Pace (Staphylinidae), are resurrected. The tribal name Stictocraniini Jakobson (Staphylinidae) is also resurrected as the valid name for its new synonym Fenderiini Scheerpeltz.
Changes at the genus-group level in Histeridae include placing Contipus Marseul as a new synonym of Hister Linnaeus due to the current placement of its validly designated type species C. subquadratus Marseul; proposal of Contipides Newton gen. nov. (type species Contipus digitatus Marseul) for the 10 species that had remained in Contipus of authors; and new designation of Idolia laevigata Lewis as type species of Idolia Lewis. In Ptiliidae, Rodwayia ovata Lea is newly designated as type species of Rodwayia Lea, and Throscidium germainii Matthews is newly designated as type species of Throscidium Matthews. In Staphylinidae, Paramichrotus Naomi is resurrected as a valid subgenus of Hesperosoma Scheerpeltz with Hemihesperosoma Hayashi placed as a new synonym of it; Sonoma corticina Casey is reaffi rmed as the type species of Sonoma Casey in place of Faronus tolulae LeConte; Stanosthetus Dejean is recognized as an available name and junior synonym of Euplectus Kirby; Taplandria Pace (type species T. guyanensis Pace) is recognized as a junior homonym and new synonym of Taplandria Pace (type species T. fl ava Pace); and Termitobiella Wasmann is resurrected as the valid name for the genus Felda Blackwelder. Replacement names for preoccupied generic or subgeneric names include in Histeridae Bellatricides Newton nom. nov. for Pachylister (Bellatrix) Mazur, junior homonym of Bellatrix Boie; and in Staphylinidae Foxiides Newton nom. nov. for Foxia Pace, junior homonym of Foxia Ashmead, and Xenasterides Newton nom. nov. for Xenaster Bierig, junior homonym of Xenaster Simonwitsch. Taxonomic changes at the generic level in Staphylinidae include proposal of Prolibia Newton gen. nov. (type species Lispinus californicus LeConte) for four Nearctic species recently placed in Clavilispinus Bernhauer; placement of Heterotrochinus Coiffait and its synonym Heterotrochus Coiffait as new synonyms of Eulibia Cameron; placement of the generic or subgeneric names Chapmaniella Bernhauer, Glenothorax Bierig, Euryolinus Bernhauer and Plesiolinus Bernhauer as new synonyms of Platydracus Thomson; and transfer of the subgenus Poikilodracus Scheerpeltz from Staphylinus Linnaeus to Platydracus. First reviser actions are used to select Georissites Ponomarenko (Georissidae) as the correct original spelling over the alternate original spelling Georyssites, and Kyrtusa Pace (Staphylinidae) as correct original spelling over Kirtusa.
Several hundred nomenclatural and taxonomic changes at the species group level are briefl y summarized here but are too numerous to list completely. Replacement names for preoccupied species or subspecies names in current use are proposed in Histeridae (3), Hydrochidae (1), Hydrophilidae (1), Leiodidae (2), Ptiliidae (3) and Staphylinidae (180); an additional staphylinid replacement name, Phloeopora nilgiriensis, is newly proposed by G. Paśnik. New or resurrected combinations are proposed for either nomenclatural or taxonomic reasons in the following genera (with indication of how many names in each genus): in Histeridae, Contipides Newton (10); in Staphylinidae, Abemus Mulsant and Rey (4), Allotrochus Fagel (6), Atheta Thomson (1), Cheilocolpus Solier (4), Eulibia Cameron (4), Foxiides Newton (1), Lispinus Erichson (3), Loncovilius Germain (2), Nacaeus Blackwelder (119), Naddia Fauvel (1), Neohypnus Coiffait and Sáiz (8), Neolosus Blackwelder (1), Ocypus Leach (2), Ontholestes Ganglbauer (1), Platydracus Thomson (59), Prolibia Newton (4) Termitobiella Wasmann (10), Thyreocephalus Guérin-Méneville (4), Xenasterides Newton (1), and Zeoleusis Steel (3). First reviser actions are used to resolve the correct original spellings (of two or more original spellings) of two species of Hydraena Kugelann (Hydraenidae) and 21 species of Staphylinidae. Changes in priority or availability of names are cited to establish the following names as valid over one or more new synonyms each: Acrotrichis rotundata (Haldeman) and Acrotrichis glabricollides Newton sp. nov. in Ptiliidae, Nemadiopsis franki Perreau in Leiodidae, and Gyrophaena nigra Kraatz, Heterothops fumigatus LeConte, Loncovilius germaini (Scheerpeltz), Philonthus upotovus Newton, sp. nov., Stenus fulviventris Rougemont, and nine species of Homalota Mannerheim in Staphylinidae. Finally, the species Eleusis lata Coiffait and Eleusis microlestiformis Coiffait are noted as not belonging to the genus Eleusis Laporte de Castelnau or to Staphylinidae, and are transferred without generic assignment to the subfamily Inopeplinae of the family Salpingidae.
Three new species of Monepidosis Mamaev, 1966, a Holarctic genus of Porricondylinae (Diptera, Cecidomyiidae), are described: M. heterocera sp. nov. from Sweden and Germany, M. scepteroides sp. nov. from Sweden, and M. shikokuensis sp. nov. from Japan. A new porricondyline genus, Antipodosis gen. nov., is introduced for eight new species from New Zealand, named A. australis gen. et sp. nov., A. elongata gen. et sp. nov., A. granvillensis gen. et sp. nov., A. pureora gen. et sp. nov., A. rakiura gen. et sp. nov., A. rotoiti gen. et sp. nov., A. rotoroa gen. et sp. nov., and A. waipapa gen. et sp. nov. Male genitalic morphology indicates that Monepidosis and Antipodosis gen. nov. are closely related, together forming the Monepidosis group of genera, which stands out from the other Porricondylini. Monepidosis spatulata Spungis, 2006, a species originally described from Latvia and Lithuania, is for the first time reported to occur in Sweden.
The new genus Neotrichaphodioides and the new species N. woytkowskii from Peru are described. Aphodius caracanus Balthasar, A. ecuadoriensis Petrovitz, A. forsterianus Balthasar, and A. volxemi Harold are redescribed and figured, and transferred into Neotrichaphodioides, all becoming new combinations. New synonymies of Aphodius martinsi Petrovitz with N. caracanus (Balthasar) and Aphodius squamifer Petrovitz with N. volxemi (Harold) are presented. The lectotype of A. volxemi is here designated.
A new genus is erected within the Cetoniini to describe a newly discovered species with characters shared between Heteroclita Burmeister, 1842, Ichnestoma Gory & Percheron, 1833 and Meridioclita Krikken, 1982. Neoclita pringlei gen. et sp. nov. exhibits a simple clypeal structure without specialized armour, along with hypertrophic and hairy tarsal segments as well as a fully winged female. The new species also exhibits an aedeagal structure closest to Meridioclita, with dorsal lobes of parameres substantially narrower than the ventral ones. The species appears to be restricted to high altitudes in the southwestern peri-Drakensberg area of the Eastern Cape Province, South Africa. Similarly to other mountain relicts known from the southern African region, adults emerge only after major rainfall events during the late spring to early summer season and do not show any evidence of feeding. It appears that flying activity may be temporarily interrupted following soil desiccation, to resume promptly after the next rainfall.
Kempfidris : a new genus of myrmicine ants from the Neotropical region (Hymenoptera: Formicidae)
(2014)
The new genus Kempfidris gen. nov. is described based on the workers of a single species, K. inusualis comb. nov., from Brazil, Ecuador, and Venezuela. Kempfidris inusualis comb. nov. was originally described by Fernández (2007) and provisionally placed in Monomorium awaiting a better understanding of the internal relationships in Myrmicinae. Kempfidris gen. nov. has a series of distinctive morphological characters including the mandibular configuration, vestibulate propodeal spiracle, propodeal carinae, and cylindrical micro-pegs on the posteromedian portion of abdominal tergum VI and anteromedian portion of abdominal tergum VII. This last trait appears to be autapomorphic for the genus.
Fungivorous gall midges of the subfamilies Lestremiinae, Micromyinae, Winnertziinae and Porricondylinae were largely neglected in previous inventories of the Diptera faunas of the Czech and Slovak Republics. A taxonomic-faunistic study focusing on these subfamilies identified a total of 80 species, of which 49 are new records for the Czech Republic and 33 are new records for Slovakia. Species that have never before been found in central Europe are Aprionus dalarnensis Mamaev, 1998, A. oligodactylus Jaschhof, 2009, A. pigmentalis Mamaev, 1998, Asynapta inflata Spungis, 1988, Camptomyia gigantea Spungis, 1989, Cassidoides fulviventris (Mamaev, 1964), Claspettomyia hamata (Felt, 1907), Dendrepidosis longipennis (Spungis, 1981), Dicerura dispersa Jaschhof, 2013, Divellepidosis lutescens (Spungis, 1981), D. pallescens (Panelius, 1965), D. vulgata Jaschhof, 2013, Ekmanomyia svecica Jaschhof, 2013, Holoneurus ciliatus Kieffer, 1896, Monepidosis pectinatoides Jaschhof, 2013, Neocolpodia gukasiani (Mamaev, 1990), Neurolyga acuminata Jaschhof, 2009, Neurolyga interrupta Jaschhof, 2009, Parepidosis planistylata Jaschhof, 2013, Peromyia bidentata Berest, 1988, Porricondyla errabunda Mamaev, 2001, P. microgona Jaschhof, 2013, P. tetraschistica Mamaev, 1988, Schistoneurus irregularis Mamaev, 1964, Spungisomyia fenestrata Jaschhof, 2013, S. media (Spungis, 1981), Tetraneuromyia lamellata Spungis, 1987, T. lenticularis (Spungis, 1987), and Winnertzia parvispina Jaschhof, 2013. A new genus including a single new species of Porricondylini is described and named Glossostyles perspicua Jaschhof & Sikora gen. et sp. nov. on the basis of specimens collected in the Czech Republic and Sweden. Adult morphology suggests that Glossostyles gen. nov. is a close relative of Claspettomyia Grover, 1964.
The geographical range of the typically host-specific species of chewing lice (Phthiraptera) is often assumed to be similar to that of their hosts. We tested this assumption by reviewing the published records of twelve species of chewing lice parasitizing wild and domestic chicken, one of few bird species that occurs globally. We found that of the twelve species reviewed, eight appear to occur throughout the range of the host. This includes all the species considered to be native to wild chicken, except Oxylipeurus dentatus (Sugimoto, 1934). This species has only been reported from the native range of wild chicken in Southeast Asia and from parts of Central America and the Caribbean, where the host is introduced. Potentially, this discontinuous distribution is due to a low tolerance for dry environments, possibly exacerbated by competitive exclusion by Cuclotogaster heterographus (Nitzsch, 1866). Our examinations of O. dentatus also revealed that this species differs significantly from other species of Oxylipeurus in the male and female genitalia, head structure and chaetotaxy, and other morphological characters. We therefore here erect the monotypic genus Gallancyra gen. nov. for O. dentatus, and redescribe the type species.
We simultaneously considered morphology and molecular phylogeny to modify the generic classification of the ‘pyropterine clade’ (Lycidae, Erotinae, Dictyopterini). To place species previously included in Benibotarus Kôno, 1932 in reciprocally monophyletic genera, we propose Gomezzuritus gen. nov. with the type-species Dictyopterus alternatus Fairmaire, 1856. Further, we transfer Gomezzuritus alternatus (Fairmaire, 1856) comb. nov., G. longicornis (Reiche, 1878) comb. nov., and G. rubripes (Pic, 1897) comb. nov. from Benibotarus to Gomezzuritus gen. nov. The pyropterine clade contains five genera in the Palaearctic region: Pyropterus Mulsant, 1838, Gomezzuritus gen. nov., Helcophorus Fairmaire, 1891, Greenarus Kazantsev, 1995, and Benibotarus Kôno, 1932. The arrangement of longitudinal elytral costae proved misleading for consideration of relationships. Two genera in distant positions share only four primary costae (Pyropterus and Helcophorus), and three similarly distant genera share the shortened primary costa 3, resulting in three primary and four secondary longitudinal costae (Gomezzuritus, Greenarus, and Benibotarus). The larva of Gomezzuritus alternatus is described in detail, and it is compared with the larvae of other Dictyopterini, including the presumed larva of G. longicornis.
Sharon gen. nov. is here described to include Asaphes? amoenus Philippi, 1861 comb. nov. from Chile. A redescription of the species is based on the female holotype and material from different geographic locations. Candèze (1891) placed Asaphes amoenus and Parasaphes elegans in the suprageneric group Asaphites. We discuss differences between Sharon gen. nov. and Hemicrepidius Germar, 1839, where Asaphes amoenus was later placed by Blackwelder (1944). Based on morphological characters, Sharon gen. nov. appears to be related to Parasaphes Candèze, 1881, Wynarka Calder, 1986, and Tasmanelater Calder, 1996, all from Australia, suggesting Gondwanan relationships.
A new monospecific genus, Pseudostegopterus gen. nov., endemic to the northwestern region of South Africa, is erected. The type species is described as Pseudospegopterus melonthinoides sp. nov. and is currently known only from male specimens. A provisional dichotomic key of the African Trichiina genera is also provided, in order to facilitate the identification of male specimens to the genus level.
The chewing louse species Lipeurus megalops Piaget, 1880, is redescribed and illustrated. This species has previously been placed in the genus Oxylipeurus Mjöberg, 1910, but marked differences in preantennal structure, male and female genitalia, abdominal chaetotaxy, and structure of abdominal plates indicate that this species is not closely related to other species in this genus. We therefore erect a new genus, Calidolipeurus gen. nov. for this species. Calidolipeurus is presently monotypic, containing only Calidolipeurus megalops gen. et comb. nov. We also provide a preliminary key to the Oxylipeurus-complex.
A new flower fly genus (Diptera, Syrphidae), Biema Huo & Zhao gen. nov. from China is described based on two new species: Biema wanglangensis Huo & Zhao gen. et sp. nov. (designated as type-species) and Biema qilianensis Huo & Liu gen. et sp. nov. The new genus can easily be distinguished by the following morphological features: head, mesonotum and scutellum black; postpronotum without pile; metasternum not reduced, posterior margin shallowly concave; katepisternum only with ventral pile patches; alula narrow, as wide as basal width of cell c; male postabdomen conspicuously more swollen than other segments, surstylus and postgonite complex, phallus unsegmented. The results of our Bayesian inference and Maximum Likelihood analysis based on sequences of the Cytochrome c oxidase subunit I (COI) gene, as well as the species delimitation tests, support the separation of Biema Huo & Zhao gen. nov. from its related taxa. Additionally, an identification key to the genera of the tribes Bacchini and Melanostomini occurring in China is provided.
Two species of the nematode family Diplopeltidae are described from Skagerrak. The new genus Belgopeltula gen. nov. is proposed for Diplopeltula belgica Vincx & Gourbault, 1992 and is characterised by: amphidial fovea circular in female and double-loop-shaped in male; excretory pore located at the level of cephalic setae bases; oral opening on the dorsal side of the body; pharynx subdivided into strongly muscularised fusiform corpus and weakly muscularised narrow and long postcorpus; female didelphic with antidromously reflexed ovaries; supplements absent. Mudwigglus micramphidium sp. nov. is characterised by: a body of 0.6 mm long; cephalic sensilla 1.5 μm long; amphidial fovea loop-shaped, 8 μm long and 3.5 μm wide; gymnostom without cuticularised ring; tail elongate conoid, with subcylindrical distal part; terminal setae absent; spicules 15 μm long; gubernaculum present; two midventral precloacal setae. It is distinguished from M. macramphidium Leduc, 2013 in having shorter amphidial fovea, shorter spicules and presence of two precloacal setae. Redescription of Diplopeltis cylindricauda Allgén, 1932 is provided based on type material. Diplopeltula minuta Vitiello, 1972 is transferred to the genus Mudwigglus Leduc, 2013. Diplopeltis cylindricauda Allgén, 1932, Diplopeltula laminata Vitiello, 1972 and Diplopeltula cassidaignensis Vitiello, 1972 are transferred to the genus Pseudaraeolaimus Chitwood, 1951.
One individual referable to Calliopiidae G.O. Sars, 1893 was collected from a chemically reduced habitat, the hydrothermal vent systems in Okinawa Trough, and was identified as a new genus and species belonging to this family after a morphological examination. A formal description of this new species and a discussion of the relationship of the new genus within Calliopiidae are presented.
Rhytidaphora Reshchikov & Quicke gen. nov. (type species Rhytidaphora thailandica Reshchikov & Quicke gen. et sp. nov.) from Thailand is described and illustrated. It belongs to the tribe Euryproctini of the subfamily Ctenopelmatinae (Hymenoptera, Ichneumonidae) based on the absence of glymma and the subapical notch on the ovipositor. The new taxon differs from all other genera of Euryproctini by the occipital carina being broadly incomplete dorsally, the hypostomal carina joining the occipital carina shortly before the base of the mandible, distinctly pectinate tarsal claws, and immovably fused and strongly sculptured second and third metasomal tergites.
A new genus, Plesioxylion Liu & Beaver gen. nov., is described for Amintinus gambianus Borowski, 2018 from West Africa with a more detailed description and new records of both sexes. We also provide a key to the ten Afrotropical genera in the tribe Xyloperthini Lesne, 1921 as the baseline information for a future study.
Afrocampe gen. nov. is described for its only species, A. prinslooi gen. et sp. nov., from Eastern and Western Cape, South Africa. This new genus is recognized as a member of the subfamily Tetracampinae Förster, 1856 of Tetracampidae Förster, 1856 based on the possession of 5-segmented tarsi in females and 4-segmented tarsi in males, the short straight calcar, the mesoscutum with distinct notauli, the mesoscutellum with two pairs of setae, the reduced mesopleural suture and the short stigmal and long postmarginal veins of the fore wing. Afrocampe gen. nov. is characterized by a large mesosoma, a non-convex first gastral tergite, an evenly acute calcar, a 5-segmented antennal funicle, a head lacking occipital carina and facial grooves and a long fore wing with distinctly delimited speculum, a bare admarginal area with a distinct admarginal row of setae on the underside and with 3 setal tracks (hair rows) radiating from the apex of the stigmal vein. The combination of these characters suggests a special status of the new genus within the subfamily Tetracampinae. Moreover, Afrocampe gen. nov. bears some resemblance to the Australian tetracampine genus Niticampe Bouček, 1988. The position of the latter in Tetracampinae, as well as habitus features of the former, are discussed.
A new fossil ceratopogonid genus and species from Lower Cretaceous Lebanese amber, Baskintoconops maaloufi Pielowska-Ceranowska gen. et sp. nov., is described and illustrated. The studied material originates from a newly discovered amber site in the Lebanese village Baskinta at a locality dubbed Qanat Bakish. The described genus is typified by its wing venation pattern combining characters of genera Fossileptoconops and Jordanoconops belonging to the subfamily Leptoconopinae.
Serratichneumon Riedel & Sheng gen. nov. and Serratichneumon maculatus Sheng & Riedel gen. et sp. nov. belonging to the tribe Ichneumonini of subfamily Ichneumoninae (Hymenoptera: Ichneumonidae), collected in the Oriental Region (China, Vietnam, and Indonesia), are described and illustrated. The new genus is placed in Tereshkin’s key to the Palaearctic genera of the subtribe Amblytelina, and compared with similar genera, Hepiopelmus Wesmael, 1845, and Tricholabus Thomson, 1894.
A new genus and species, Richerius marqueti gen. et sp. nov., of a crab of the family Hymenosomatidae MacLeay, 1838 are described from the inland waters of New Caledonia based on several specimens collected in two streams at altitudes of 180 m and 500 m, respectively. Richerius marqueti gen. et sp. nov. was compared to the other freshwater species known in New Caledonia, Odiomaris pilosus (A. Milne-Edwards, 1873), and to species of Amarinus Lucas, 1980, a genus comprising many freshwater species in New Zealand, Australia, Indonesia, the Philippines, and Papua New Guinea, but never recorded in New Caledonia. The barcode fragment of the COI mitochondrial gene was sequenced for seven specimens of R. marqueti gen. et sp. nov., and all sequences were deposited in GenBank. A brief and updated review of the New Caledonian marine and freshwater hymenosmatid fauna is provided.
Metopiinae is a relatively large subfamily of Darwin wasps with a cosmopolitan distribution. Species of this subfamily can be distinguished by a convex, and mostly undivided face and clypeus. Among 113 species in 24 genera of the Oriental region, only 24 species in eight genera are reported from India. Here, we describe and illustrate a new genus, Soliga, based on collections made from the Western Ghats and Eastern Himalayas, two important biodiversity hotspots. The combination of absence of epicnemial carina on dorsolateral part of mesopleuron, absence of propodeal carinae and dorsal carinae of first metasomal tergite differentiates the new genus from other metopiine genera. An illustrated key to the Indo-Australian metopiine genera is provided along with comments on new generic placement and character variability of metopiines.
The “trachystreptoform” species of Spirostreptidae, i.e., species which would formerly have been ascribed to the tribe Trachystreptini, from the Udzungwa Mountains are (re)described, including one new genus and five new species: Attemsostreptus reflexus Akkari & Enghoff, 2019, A. cataractae Enghoff sp. nov., A. leptoptilos Enghoff sp. nov., A. julostriatus Enghoff sp. nov., Lophostreptus tersus (Cook, 1896) (= L. ptilostreptoides Carl, 1909 syn. nov.), L. magombera Enghoff sp. nov., and Udzungwastreptus marianae Enghoff gen. et sp. nov. The type material of Lophostreptus regularis Attems, 1909 (= L. tersus) is discussed. The discussion includes paragraphs on the classification and the Udzungwa fauna of Spirostreptidae, on grouping of the Udzungwa trachystreptoform species in relation to altitude, and on the possibly recent immigration of A. reflexus and L. tersus into the Udzungwa Mts.