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Ants of the Tetramorium solidum group occur in Africa, with the vast majority of species endemic to the arid regions of southern Africa. The first revision of the genus was published more than 30 years ago and ant surveys have since considerably expanded the number of specimens available for study. The revision of this group reveals five new species, expanding the total number to 19. Almost all the species in this group occur in the southern parts of the Afrotropical region, with the exception of T. setuliferum Emery, 1895 and T. rothschildi (Forel, 1907). These two species have broad distributions within African grasslands and savannas, with T. setuliferum occurring in southern Africa and T. rothschildi in East Africa and the Sahel. Five new species are described in this revision: T. aisha sp. nov., T. brigitteae sp. nov., T. duncani sp. nov., T. lerouxi sp. nov. and T. margueriteae sp. nov. An illustrated key is presented and descriptions of new species are provided, supported by montage images and distribution maps.
Homollea Arènes (Rubiaceae, subfamily Ixoroideae, tribe Pavetteae) is a genus of shrubs and small trees endemic to western and northern Madagascar. The genus comprises five species occurring in dry deciduous forest, often in limestone areas. The five species are narrow endemics and their conservation status is either Endangered (4 species) or Critically Endangered (1 species). Homollea is characterized by few-flowered, pseudo-axillary, pedunculate inflorescences, well-developed calyces with the lobes much longer than the tube, laterally flattened seeds with a shallow, elongated to linear hilum and entire endosperm, ovules arising from the upper margin of the placenta, and, pollen grains with supratectal elements in the shape of microgemmae. Until now, three species were known and their descriptions are amended. Two further species, H. furtiva De Block sp. nov. and H. septentrionalis De Block sp. nov., are described as new for science. The five species are dealt with in detail: descriptions, distribution maps, conservation assessments, illustrations, lists of exsiccatae and an identification key are given.
Literature about Mesitiinae Kieffer, 1914 has not been treated extensively from a taxonomic viewpoint in comparison with other subfamilies in Bethylidae Latreille, 1802. Our research on species of Metrionotus Móczár, 1970, Clytrovorus Nagy, 1972 and Sulcomesitius Móczár, 1970 revealed a new hypopygium shape pattern, namely a 'star-shaped' hypopygium, which is characteristic of a new genus, Astromesitius gen. nov., with two new species Astromesitius thionyi gen. et sp. nov. and Astromesitius olavoi gen. et sp. nov. The descriptions of both new species are based on male specimens collected in Thailand and the United Arab Emirates. Astromesitius quatei (Móczár, 1977) gen. et comb. nov. is designated as type species for the new genus, which is erected for a total of seven species. The main diagnostic characteristics are the head longer than wide; a clypeus with a median lobe quadrate; an antenna with distinct long setae, with pedicel caliciform, and with flagellomeres long and caliciform; pronotum and anteromesoscutum with longitudinal sulcus indistinct or absent; metapectal-propodeal complex with posterior projection hardly distinct or absent; hypopygium star-shaped; genitalia with aedeagus slender and fusiform.
Although Gastrotricha have previously been recorded in bottom sediments of greenhouses as well as in micro-reservoirs of Bromeliaceae, palm houses provide a more specific microhabitat for these animals that often originate from different regions of the world. This paper presents an investigation of gastrotrich assemblages associated with aquatic plants. Eight species of the epiphytic chaetonotids were found in the Jubilee Greenhouse of the Botanical Garden in Kraków (Poland), including three species new to science, Chaetonotus (Chaetonotus) invitatus sp. nov., C. (Hystricochaetonotus) horridus sp. nov. and C. (H.) inaequabilis sp. nov. Two other species are new to Poland, C. (C.) paucisquamatus Kisielewski, 1991 and C. (Zonochaeta) cestacanthus Balsamo, 1990, both recorded here for the first time outside their terra typica. These observations confirm that greenhouses and palm houses provide many ecological niches and favourable conditions for the development of a number of unintentionally introduced species which cannot be found outside their original climate conditions.
Thirty-six species of various thecate hydroids occur in two recent, deep-water collections from off New Caledonia. Of these, nine are new, namely Solenoscyphus subtilis Galea, sp. nov., Hincksella immersa Galea, sp. nov., Synthecium rectangulatum Galea, sp. nov., Diphasia alternata Galea, sp. nov., Dynamena opposita Galea, sp. nov., Hydrallmania clavaformis Galea, sp. nov., Symplectoscyphus acutustriatus Galea, sp. nov., Symplectoscyphus elongatulus Galea, sp. nov. and Zygophylax niger Galea, sp. nov. The male and female gonothecae of Caledoniana decussata Galea, 2015, the female gonothecae of Caledoniana microgona Galea, 2015, as well as the gonothecae of both sexes of Solenoscyphus striatus Galea, 2015 are described for the first time. The systematic position of the genera Solenoscyphus Galea, 2015 and Caledoniana Galea, 2015 is discussed on both morphological and molecular grounds, and both are confidently placed within the family Staurothecidae Maronna et al., 2016. In light of the molecular data, the genera Billardia Totton, 1930 and Dictyocladium Allman, 1888 are assigned to the families Syntheciidae Marktanner-Turneretscher, 1890 and Symplectoscyphidae Maronna et al., 2016, respectively. The previously undescribed gonothecae of Hincksella neocaledonica Galea, 2015, and the male gonothecae of Sertularella tronconica Galea, 2016, were found. Thyroscyphus scorpioides Vervoort, 1993, a peculiar hydroid with putative stem nematothecae, is redescribed and assigned to the new genus Tuberocaulus Galea, gen. nov. Noteworthy new records from the study area are: Tasmanaria edentula (Bale, 1924), Hincksella sibogae Billard, 1918, Dictyocladium reticulatum (Kirchenpauer, 1884), Salacia sinuosa (Bale, 1888) and Billardia hyalina Vervoort & Watson, 2003. Most species are illustrated to facilitate their identification, and the morphology of the new ones is compared to that of their related congeners.
The Swedish species of Ophion Fabricius, 1798 are revised. More than 4800 specimens and relevant type material were studied; 234 sampled specimens produced COI sequences. The study recognises 41 species, 18 of which are described as new to science, mainly from Fennoscandian material: Ophion angularis Johansson & Cederberg sp. nov., Ophion arenarius Johansson sp. nov., Ophion autumnalis Johansson sp. nov., Ophion borealis Johansson sp. nov., Ophion broadi Johansson sp. nov., Ophion brocki Johansson sp. nov., Ophion confusus Johansson sp. nov., Ophion ellenae Johansson sp. nov., Ophion inclinans Johansson sp. nov., Ophion kallanderi Johansson sp. nov., Ophion matti Johansson sp. nov., Ophion norei Johansson sp. nov., Ophion paraparvulus Johansson sp. nov., Ophion paukkuneni Johansson sp. nov., Ophion splendens Johansson sp. nov., Ophion sylvestris Johansson sp. nov., Ophion tenuicornis Johansson sp. nov. and Ophion vardali Johansson sp. nov. Barcoding analysis also indicated the possible presence of at least three additional, partly cryptic species, but these cannot be separated morphologically with certainty at this point. Ophion costatus Ratzeburg, 1848 and Ophion artemisiae Boie, 1855 are interpreted and defined. Ophion slaviceki Kriechbaumer, 1892 is excluded from synonymy with Ophion luteus Linnaeus, 1758 stat. rev. Ophion polyguttator (Thunberg, 1824) stat. rev. and Ophion variegatus Rudow, 1883 stat. rev. are excluded from synonymy with O. obscuratus Fabricius, 1798. Ophion variegatus is redescribed and a neotype is designated. Ophion albistylus Szépligeti, 1905 (syn. nov.) is synonymized with Ophion pteridis Kriechbaumer, 1879 and Ophion frontalis Strobl, 1904 (syn. nov.) is synonymized with Ophion areolaris Brauns, 1889 syn. nov. Eleven species are reported from Sweden for the first time: Ophion artemisiae, Ophion crassicornis Brock, 1982, Ophion costatus, Ophion dispar Brauns, 1895, Ophion forticornis Morley, 1915, Ophion kevoensis Jussila, 1965, Ophion ocellaris Ulbricht, 1926, Ophion perkinsi Brock, 1982, Ophion subarcticus Hellén, 1926, Ophion variegatus Rudow, 1883 and Ophion wuestneii Kriechbaumer, 1892. The study shows that a number of species that previously have been treated as highly variable taxa, actually consist of several valid species that are separable using morphological characters. An illustrated key for the determination of the Swedish Ophion species is provided.
We revise the genus Attemsostreptus Verhoeff, 1941 based on type material of the type species, A. costatus Verhoeff, 1941, synonymise A. orobius (Kraus 1958) with A. costatus and describe a second species of the genus, A. reflexus sp. nov., collected from Kimboza Forest Reserve in Tanzania, and discuss the dubious tribe Trachystreptini.
Alpheus macrocheles (Hailstone, 1835), a species originally described from the northeastern Atlantic, has been reported from Brazil based on material from the north and northeast coasts and Espírito Santo. However, a thorough morphological comparison between Brazilian material reported as A. macrocheles and eastern Atlantic material of A. macrocheles revealed consistent differences, suggesting that the Brazilian specimens belong to an undescribed species. Alpheus ramosportoae sp. nov. is therefore now described based on material from Amapá to Pernambuco, Brazil. Morphological differences between the new species and A. macrocheles s. str. were supported by the clear divergence of 16S rRNA gene sequences (18% of genetic distance), separating the species in two distinct clades. Differences in the color pattern also were observed and illustrated.
During recent investigations on the terrestrial invertebrates of the tropical rainforest on Martinique Island (Pitons du Carbet), specimens of a new species of the terrestrial amphipod genus Cerrorchestia Lindeman, 1990, C. taboukeli sp. nov., were collected by means of different quantitative and non-quantitative methods (hand collection and Tullgren extraction) in the forest floor. The new species can be easily distinguished from the only other species of the genus, C. hyloraina Lindeman, 1990, by gnathopod 2 (carpus short, palm longer than wide), pereopod 4 dactylus with a denticulate patch, pereopod 5 basis ovate with a deep posterodistal lobe reaching the distal end of the ischium, pleopod 3 ramus with more than six articles. Cerrorchestia tabouleki sp. nov. is the first forest-hopper discovered in the Lesser Antilles, raising the question of island colonization by terrestrial amphipods. Ecological data and a key to terrestrial Talitridae of Central America and the Caribbean islands are provided.
The Asian species of the subfamily Thaumastodinae Champion, 1924 are reviewed. Seven new species are described: Acontosceles borneensis sp. nov., Pseudeucinetus papuanus sp. nov., Mexico ogasawaraensis sp. nov., M. baliensis sp. nov., M. papuanus sp. nov., M. palauensis sp. nov. and M. borneensis sp. nov. The genus Babalimnichus Satô, 1994 is treated as a junior synonym of the genus Mexico Spilman, 1972, and three known species of the genus Babalimnichus are transferred to Mexico, viz. M. taiwanus (Satô, 1994) comb. nov., M. masamii (Satô, 1994) comb. nov. and M. splendens (Hernando & Ribera, 2003) comb. nov. Additional specimen data are shown, and new distributional records are as follows: Acontosceles chujoi Yoshitomi & Satô, 2005 from Vietnam; A. zetteli Pütz, 2008 from Laos; Pseudeucinetus javanicus Yoshitomi & Putra, 2010 from Lombok Island; Mexico taiwanus (Satô, 1994) comb. nov. from Lutao, Lanhsu and the Yonaguni-jima Islands; and M. masamii (Satô, 1994) comb. nov. from Kume-jima. A species list of the subfamily Thaumastodinae is given, with ZooBank LSIDs. The phylogenetic relationships of the thaumastodine genera are discussed.
The world species of Netomocera Bouček, 1954 (Hymenoptera Linnaeus, 1758: Pteromalidae Dalman, 1820), excluding those from the Oriental region, are revised. The Oriental species are excluded because their types could not be examined, the species limits could not be reliably assessed based on original descriptions and available Oriental material was scarce. Eighteen species, including 11 species described as new, are recognized: N. africana Hedqvist, 1971; N. alboscapus Hedqvist, 1971; N. amethysta sp. nov.; N. celebensis sp. nov.; N. cyanocephala sp. nov.; N. desaegeri sp. nov.; N. formiciformis sp. nov.; N. gloriosa sp. nov.; N. irregularis sp. nov.; N. masneri sp. nov.; N. merida sp. nov.; N. meridionalis sp. nov.; N. nearctica Yoshimoto, 1977; N. ramakrishnai Sureshan, 2010; N. rufa Hedqvist, 1971; N. sedlaceki Bouček, 1988; N. setifera Bouček, 1954; N. virgata sp. nov. The female brachypterous form of N. nearctica and the male of N. alboscapus are described for the first time. A key to both sexes is provided, as well as diagnoses, descriptions and illustrations for all treated species. The genus is reported for the first time in the Neotropical region. For several species, new distributional records are also given.
First records of Elachistinae are given from Thailand. Ten species of Elachistinae are reported, eight of which are described as new: Urodeta longa Sruoga & Kaila sp. nov., Elachista buszkoi Sruoga & Kaila sp. nov., E. oryx Sruoga & Kaila sp. nov., E. pellineni Sruoga & Kaila sp. nov., E. capricornis Sruoga & Kaila sp. nov., E. phichaiensis Sruoga & Kaila sp. nov., E. loeiensis Sruoga & Kaila sp. nov. and E. siamensis Sruoga & Kaila sp. nov. The new species are diagnosed and illustrated with photographs of the adults and genitalia. One species remains unidentified to species level.
A total of nine families of Trichoptera were identified from material collected in Malaise and light traps in the western part of Nyungwe National Park, southwestern Rwanda, late October 2018. Included in the material was an undescribed species of Pisuliidae which is described herein as Silvatares laetae Ngirinshuti & Johanson sp. nov. The new species adds to the six Pisuliidae species previously recorded for the East African region, five endemic to Tanzania and one to Uganda. This study portrays the first results of an ongoing survey on the Trichoptera fauna of Rwanda.
This paper is part of a project of studying benthic diatom biodiversity on marine coastal regions of Sweden with focus on rare and less known species. Two new species of Cocconeis Ehrenb. are described from Vrångö, a small island in the west coast of Sweden. Both species were found as epiphytic on the green alga Ulva intestinalis L. Cocconeis magnoareolata Al-Handal, Riaux-Gob., R.Jahn & A.K.Wulff sp. nov. is a small species not exceeding 9 μm in length and characterized by having large subquadrangular areolae on the sternum valve. Cocconeis vrangoensis Al-Handal & Riaux-Gob. sp. nov. appears similar to some taxa of the 'Cocconeis scutellum complex', but differs by its stria density on both valves and variable features of the areola and valvocopula ultrastructure. Detailed descriptions based on light and electron microscopy examination, a comparison with closely related taxa, as well as a description of the habitat of both species are here presented.
Four new species of Brueelia Kéler, 1936 are described and illustrated. All of them parasitize African endemic host species in the families Passeridae, Ploceidae, and Estrildidae (Passeriformes). They are: Brueelia pofadderensis sp. nov. ex Passer melanurus damarensis Reichenow, 1902 and P. m. vicinus Clancey, 1958; B. semiscalaris sp. nov. ex Granatina granatina (Linnaeus, 1758); B. sima sp. nov. ex Malimbus nitens (Gray, 1831); B. terpsichore sp. nov. ex Euplectes jacksoni (Sharpe, 1891) and E. progne delamerei (Shelley, 1903). In addition, Brueelia bicurvata (Piaget, 1880) is redescribed and reillustrated from non-type material. A summary of all published records of lice in the Brueelia complex from Africa since 1980 is provided. We also estimate the unknown diversity of African species of Brueelia based on an index of host specificity calculated for each host family independently. The unknown diversity is estimated to be over 1000 species of Brueelia from African hosts, compared to the < 50 species in this genus currently recorded from Africa.
This paper is based on three collections of Ptiliidae from Sarawak totalling more than 2000 specimens made by staff and affiliates of the Natural History Museum, London, between March–July 1978. One new genus Niptella gen. nov with its type species Niptella gutta gen. et sp. nov. and 24 new species are described and figured: Sindosium collinsi sp. nov., Bambara hammondi sp. nov., Bambara subtortuosa sp. nov., Bambara tortuosa sp. nov., Erro brookei sp. nov., Cissidium globulum sp. nov., Cissidium longum sp. nov., Cissidium marshallae sp. nov., Cissidium pauxillum sp. nov., Cissidium subfoveolatum sp. nov., Cissidium triangulum sp. nov., Discheramocephalus nigritus sp. nov., Kuschelidium sarawakense sp. nov., Ptinella alisonae sp. nov., Acrotrichis acuta sp. nov., Acrotrichis belli sp. nov., Acrotrichis bidens sp. nov., Acrotrichis geiseri sp. nov., Acrotrichis globosa sp. nov., Acrotrichis hanskii sp. nov., Acrotrichis muluensis sp. nov., Acrotrichis plaga sp. nov., Storicricha resticula sp. nov., Storicricha umbella sp. nov. New records and information are provided for Dipentium latum Darby, 2019; Ptiliola semitaria Darby, 2018; Baeocrara minima Darby, 2019; Acrotrichis agricola Darby, 2019; Acrotrichis britteni Johnson, 1969; Acrotrichis bubalis Darby, 2019; Acrotrichis cognata (Matthews, 1877) and Acrotrichis cursitans (Nietner, 1856).
We present a review of Dendrocephalus (Dendrocephalinus) with an updated diagnosis for the subgenus and a key to all known species. We provide new records of Dendrocephalus alachua, which was previously supposed to be extinct, and we describe a new species, Dendrocephalus proeliator sp. nov., which is separated from all other species based on the form of the male frontal appendage. Dendrocephalus proeliator sp. nov. appears to be morphologically intermediate between D. alachua and D. lithaca. In addition, we provide conservation assessments for all four species in the subgenus, according to IUCN Red List standards. We also report for two species the first known examples of direct male-male agonistic behaviour and competition for access to areas frequented by receptive females.
Three new species of Thinophilus Wahlberg, 1844 are described from marine sites on the Andaman Sea coasts in peninsular Thailand: T. inaequalis Samoh & Grootaert sp. nov. belonging to the 'Thinophilus simplex-group' sensu Grootaert, 2018, T. plektron Samoh & Grootaert sp. nov., an unplaced species characterised by a long flattened apical spur on the hind tibia, and T. subapicalis Samoh & Grootaert sp. nov. belonging to the 'Thinophilus spinatus-group' sensu Grootaert, 2018. New records are given for following species occurring in peninsular Thailand: T. apicatus Grootaert, 2018, T. chaetulosus Grootaert, 2018, T. clavatus Zhu, Yang & Masunaga, 2006, T. evenhuisi Grootaert, 2018, T. lenachanae Grootaert, 2018, T. longicilia Evenhuis & Grootaert, 2002, T. nigrilineatus Grootaert, 2018, T. simplex Grootaert, 2018, T. superbus Grootaert, 2018, and T. yeoi Grootaert, 2018.
The gigas species group of the subgenus Canthidium (Neocanthidium) is defi ned and described. This species group is composed of three described species [C. gigas Balthasar, 1939, Brazilian Atlantic Forest, including intrusions into Cerrado, C. bokermanni (Martínez et al., 1964), Chaco and western Cerrado in Brazil, Bolivia, Paraguay, and Argentina, and C. kelleri (Martínez et al., 1964), Brazilian Cerrado and neighbouring open areas] and three new species: Canthidium stofeli sp. nov. from the western and southern regions of the Brazilian Amazon, Canthidium feeri sp. nov. from French Guiana, and Canthidium ayri sp. nov. from the Brazilian Atlantic Forest. We present descriptions and redescriptions, illustrations, an identifi cation key and comments on the distributions of the species of the gigas group.
In the present paper three empidoid fly species new to science are described from the island Sal of the Cabo Verde archipelago. Aphrosylus salensis sp. nov., a marine dolichopodid fly, is found in the splash zone of a rocky sea shore. Crossopalpus salensis sp. nov. (Hybotidae) is recorded from two different sebkhas. Crossopalpus complicatus sp. nov. is reported from an irrigated wasteland. A key to the species of these two genera occurring on Cape Verde is provided. The presence of the morphospecies Tachytrechus tessellatus (Macquart, 1842), a presumed Old World tropical species, is confirmed.
Eleven species of Ctenopelma Holmgren, 1857 are reported from China. Five species are new to science: C. labiatum Sheng, Sun & Li sp. nov., from Ningxia Hui Autonomous Region, C. lii Sheng, Sun & Li sp. nov., from Liaoning province, C. rufofasciatum Sheng, Sun & Li sp. nov., reared from Cephalcia lariciphila (Wachtl, 1898) from Beijing, C. pineatum Sheng, Sun & Li sp. nov., reared from Acantholyda posticalis (Matsumura, 1912) and Cephalcia lariciphila from Beijing and Henan, Shanxi, Shan’xi, Liaoning, Jilin and Heilongjiang provinces, and C. spiraculare Sheng, Sun & Li sp. nov., reared from Cephalcia lariciphila from Henan and Shanxi provinces. One species, C. nigrum Holmgren, 1857, reared from Cephalcia lariciphila in Beijing, is a new record for China. Ctenopelma tomentosum (Desvignes, 1856) was reared from Neurotoma sibirica Gussakovskij, 1935 (new host record) in Liaoning province. A key to species of Ctenopelma known in China is provided.
Description of Hemicordulia tuiwawai sp. nov. from Kadavu Island, Fiji (Odonata: Corduliidae)
(2019)
Hemicordulia tuiwawai sp. nov. (Odonata: Corduliidae) is described and diagnosed based on material collected from Kadavu Island, Fiji; holotype: Wainitayuki River about 750 m above Baidamudamu village, -19.0916, 178.1038; 37 m a.s.l., 06 June 2016, M. Marinov leg. This species is distinguished from its congeners in the field by the contrasting colouration – dark green metallic body with bright yellow spots on the synthorax and base of the abdomen. This pattern is comparable to H. pacifica Selys, 1871. However, Hemicordulia tuiwawai sp. nov. can be recognised by the larger size and unique shape of the caudal appendages and genital hamule (in males) and vulvar scale (in females).
New species and records of Sericini from the Indian subcontinent (Coleoptera, Scarabaeidae) II
(2019)
The current paper presents new locality records, including first state records for Mizoram and Nagaland, of 50 species of Sericini (Coleoptera: Scarabaeidae: Melolonthinae) from the Indian subcontinent. Nine new species are described herein: Maladera naveeni sp. nov., M. sujitrae sp. nov., M. thirthahalliensis sp. nov., M. viraktamathi sp. nov., Neoserica (s. lat.) reuteri sp. nov., Oxyserica goertzae sp. nov., Selaserica hosanagarana sp. nov., Serica (s. str.) eberlei sp. nov. and S. (s. str.) tashigaonensis sp. nov.
Ampharete oculicirrata sp. nov. (Annelida: Ampharetidae) is described from samples collected by the Joint Nature Conservation Committee and Marine Scotland Science, in the West Shetland Shelf NCMPA in the NE Atlantic. This species is characterised by a very small body size, thin and slender paleae, twelve thoracic and eleven abdominal uncinigers, presence of eyes both in the prostomium and the pygidium, the latter provided with a pair of long lateral cirri. The external micro-morphology of the new taxon was studied using scanning electron microscopy and compared with species described or reported from the North Atlantic. Two complementary keys to all species of Ampharete in the area are also provided.
The Afrotropical (including Malagasy Subregion) species of the genus Asobara Foerster, 1863, are revised. In addition to the redescribed 15 known species, 25 new species are described and illustrated, viz., Asobara abyssiniensis Peris-Felipo, sp. nov., A. caboverdensis van Achterberg, sp. nov., A. carinata Peris-Felipo, sp. nov., A. cracentis van Achterberg, sp. nov., A. elongitarsis van Achterberg, sp. nov., A. fletcheri Peris-Felipo, sp. nov., A. harrinsmithensis Peris-Felipo, sp. nov., A. kawandensis Peris-Felipo, sp. nov., A. kibalensis van Achterberg, sp. nov., A. laticlypeata van Achterberg, sp. nov., A. mediana van Achterberg, sp. nov., A. mellicephalata van Achterberg, sp. nov., A. natalensis Peris-Felipo, sp. nov., A. notleyi Peris-Felipo, sp. nov., A. robusta van Achterberg, sp. nov., A. sarae Peris-Felipo, sp. nov., A. somersetensis Peris-Felipo, sp. nov., A. stubbsi Peris-Felipo, sp. nov., A. taylori Peris-Felipo, sp. nov., A. vanalpheni van Achterberg, sp. nov., A. vanharteni van Achterberg, sp. nov., A. victoriana Peris-Felipo, sp. nov., A. zaprionae van Achterberg, sp. nov., A. zimbabwana Peris-Felipo, sp. nov., A. zululana Peris-Felipo, sp. nov. Moreover, the following new combination is suggested: Asobara pulchricornis (Szépligeti, 1911) comb. nov. A key to all Afrotropical (including Malagasy) species is provided for the first time.
Neopanorpa van der Weele, 1909 is the second largest genus in Panorpidae, and over 170 Oriental species have been reported hitherto. In this paper, we describe two new species: Neopanorpa luojishana sp. nov. and Neopanorpa xingmini sp. nov. from Southwest China. We also report the first discovery of the male of Neopanorpa ocellaris (Navás, 1908) from Guangxi and Guizhou, China, a species recorded from Sikkim previously. Neopanorpa brevivalvae Chou & Wang, 1988 is synonymized with Neopanorpa lungtaushana Cheng, 1957. In addition, Neopanorpa furcula nom. nov. is proposed for Neopanorpa furcata Zhou, 2005, a junior homonym preoccupied by Neopanorpa furcata (Hardwicke, 1825). The biogeographical implications of some Neopanorpa species are discussed in brief.
The genus Elaeidobius Kuschel, 1952 (Curculionidae, Curculioniae, Derelomini) is an Afrotropical genus associated with the male inflorescences of the oil palm Elaeis guineensis Jacq. The activity of species in this genus is critical for pollen transportation and for the fruit set of this economically important palm. In this study, the genus Elaeidobius was revised using an integrative taxonomic approach, combining traditional taxonomic treatment of species and an analysis of sequences of mitochondrial genes (COI and COII). A total of eight species is now recognized: five now formally included within it [E. bilineatus Hustache, 1924) comb. nov., E. kamerunicus (Faust, 1898) comb. nov., E. plagiatus (Fåhraeus, 1844) comb. nov., E. singularis (Faust, 1898) comb. nov., E. subvittatus Faust, 1898)], one transferred here from the genus Prosoestus to the genus Elaeidobius, E. spatulifer (Marshall, 1950) comb. nov., and two newly described species (E. pilimargo Haran & Kuschel sp. nov., E. piliventris Haran & Kuschel sp. nov.). The following new synonymies are proposed: Prosoestus armatus Voss, 1956 = E. bilineatus (Hustache, 1924) comb. nov. and Derelomus uelensis Hustache = E. singularis (Faust, 1898). An illustrated key to the species is provided with photographs of the adult habitus and male genitalia.
This study reports on 25 species of hydroids occurring in the collections gathered during KANACONO and KANADEEP expeditions carried out in the SE of New Caledonia in 2016, and off the western coast of the island in 2017, respectively. Of these, 19 have not been dealt with in earlier reports on these collections. Two new genera and four new species are described, viz, Actinopluma mirifica Galea gen. et sp. nov., provisionally assigned to the family Kirchenpaueriidae Stechow, 1921, Schizoplumularia helicoidalis sp. nov., belonging to the Plumulariidae McCrady, 1859, and Corhiza patula Galea sp. nov. and Thamnopteros uniserius Galea gen. et sp. nov., both placed in the family Halopterididae Millard, 1962. The gonotheca and the medusoid gonophore of Plumularia contraria Ansín Agís et al., 2014 are described for the first time, allowing a genus transfer to Dentitheca Stechow, 1919, as D. contraria comb. nov. Plumularia conjuncta Billard, 1913, known earlier from a minute portion of colony, is redescribed based on a complete, though infertile, specimen. Similarly, complete specimens corresponding to the hydroid previously referred to as Antennella megatheca Ansín Agís et al., 2009 are documented, allowing a provisional reallocation to Corhiza Millard, 1962 and a description of its so far unknown gonothecae. Fertile material assignable to the poorly-known Monostaechas fisheri Nutting, 1905 allows the recognition of this hydroid as a valid species, distinct from M. quadridens (McCrady, 1859). Most taxa are illustrated to validate the reliability of their identifications. Finally, phylogenetic reconstructions of the families Aglaopheniidae, Plumulariidae, and Halopterididae, based on the 16S rRNA, allowed a first genetic characterization of some of the species dealt with in this work.
Sixteen aglaopheniid hydroids occurring in two recent, deep water collections from off New Caledonia are reported upon, of which 8 species are new, namely Cladocarpus asymmetricus sp. nov., C. partitus sp. nov., C. pennatus sp. nov., Lytocarpia fragilis sp. nov., L. pilosa sp. nov., L. pseudoctenata sp. nov., L. subtilis sp. nov. and Macrorhynchia spiralis sp. nov., the latter producing medusoid gonophores. Although not occurring in the present collections, brief notes on M. disjuncta (Pictet, 1893), including the first description of its gonosome, are provided to support its specific separation from M. phoenicea (Busk, 1852).
A new species of praying mantis, Vates phoenix sp. nov. (Mantidae, Vatinae), is described from localities within Rio de Janeiro and São Paulo states, Brazil. This is the first record of Vates from the Atlantic Rainforest biome. The new species is unique among its congeners in having, among other features, strongly reduced cuticular projections above the lateral ocelli, structures otherwise well developed and produced in all other species of Vates. Remarks on the natural history and biogeography of Vates, in relation to this new finding, are further discussed.
A new genus and species, Richerius marqueti gen. et sp. nov., of a crab of the family Hymenosomatidae MacLeay, 1838 are described from the inland waters of New Caledonia based on several specimens collected in two streams at altitudes of 180 m and 500 m, respectively. Richerius marqueti gen. et sp. nov. was compared to the other freshwater species known in New Caledonia, Odiomaris pilosus (A. Milne-Edwards, 1873), and to species of Amarinus Lucas, 1980, a genus comprising many freshwater species in New Zealand, Australia, Indonesia, the Philippines, and Papua New Guinea, but never recorded in New Caledonia. The barcode fragment of the COI mitochondrial gene was sequenced for seven specimens of R. marqueti gen. et sp. nov., and all sequences were deposited in GenBank. A brief and updated review of the New Caledonian marine and freshwater hymenosmatid fauna is provided.
During a survey of the moss-inhabiting diatom flora of the sub-Antarctic Campbell Island, located in the southern Pacific Ocean, several unknown centric diatoms were observed that could not be identified using the currently available literature. Detailed light and scanning electron microscopical observations and comparisons with the characters of several species of Melosira, Angusticopula, Ferocia and Arcanodiscus worldwide indicated that five of them should be described as new to science: Angusticopula cosmica Goeyers & Van de Vijver sp. nov., Arcanodiscus crawfordianus Goeyers & Van de Vijver sp. nov., A. indistinctus Goeyers & Van de Vijver sp. nov., A. saundersianus Goeyers & Van de Vijver sp. nov. and Ferocia houkiana Goeyers & Van de Vijver sp. nov. A sixth species, Angusticopula chilensis, was illustrated for the first time using SEM and as a result is considered to differ sufficiently from A. dickiei to warrant epitypification as A. chilensis. The new species were first described in genera that formerly were included within the genus Melosira. All six species are morphologically characterized and compared with similar species within their respective genera. Their presence and distribution on Campbell Island are discussed based on the observations made in the available samples.
A new genus and six new species of the family Gomphodesmidae from the Udzungwa Mts are described, including Pogoro alopias Rosenmejer & Enghoff sp. nov., Pogoro siren Rosenmejer & Enghoff sp. nov., Pogoropsis prolixopes Rosenmejer & Enghoff gen. et sp. nov., Emphysemastix frampt Olsen & Enghoff sp. nov., Agrophogonus hamulus Olsen & Enghoff sp. nov., and Agrophogonus pusillokiellandi Olsen & Enghoff sp. nov. Emphysemastix dracarys Olsen & Enghoff sp. nov. from Iringa city is described. Descriptive notes are given for Pogoro scharffi Hoffman, 2005, and Agrophogonus mwanihana Hoffmann, 2005. General gomphodesmid gonopod morphology is described and illustrated. A key to Udzungwa gomphodesmids is presented, as well as revised keys to all species of Emphysemastix and Agrophogonus. All gomphodesmid species from the Udzungwa Mts are mapped.
A new fossil diatom species, Tertiarius minutulus sp. nov., is described from a sediment sequence DEEP-5045-1 of Lake Ohrid. The species is characterized by small valves (3.0–8.0 μm) with a round shape, a marginal area with radially arranged costae and a central uneven area with scattered areolae. Externally, the alveoli are occluded by cribra perforated by irregularly arranged pores. Internally, the alveoli are simple and areolae are occluded with domed cribra. The marginal fultoportulae are situated on costae close to the valve margin, located on every 5th or 7th thick internal costa. One to three fultoportulae are present on the valve face, each surrounded by two to three satellite pores. One rimoportula is present, positioned on a costa at the valve face / mantle junction. The species is compared with morphologically similar taxa and a detailed differential diagnosis is provided. Tertiarius minutulus sp. nov. is known only as a fossil taxon from Lake Ohrid, with a first occurrence during the early stages of lake basin development.
A new jumping spider genus, Manzuma gen. nov. (Salticidae Blackwall, 1841), is described, type species is Manzuma nigritibia (Caporiacco, 1941). Aelurillus reconditus Wesołowska & van Harten, 1994 is synonymized with Rafalus nigritibiis (Caporiacco, 1941). Four new combinations are proposed: M. jocquei gen. et comb. nov. (ex Aelurillus), M. kenyaensis gen. et comb. nov. (ex Langelurillus), M. lympha gen. et comb. nov. (ex Rafalus) and M. nigritibia gen. et comb. nov. (ex Rafalus). Three species, M. botswana gen. et sp. nov. (♂♀, Botswana and Republic of South Africa), M. petroae gen. et sp. nov. (♂♀, Republic of South Africa) and M. tanzanica gen. et sp. nov. (♂, Tanzania), are described. The male of M. kenyaensis gen. et comb. nov. and female of M. lympha gen. et comb. nov. are described for the first time. A new aelurilline synapomorphy is proposed. Identification key for males is provided.
This paper examines the described species of Cissidium and adds 77 new ones based on a study of material in the Manchester Museum, UK, the Natural History Museum, London, UK and in the possession of Dr Yoshihiro Sawada, Japan. The new taxa are: Cissidium advena sp. nov., C. amazonicum sp. nov., C. amieuense sp. nov., C. angulatum sp. nov., C. angusi sp. nov., C. apterum sp. nov., C. arcuatum sp. nov., C. aristophanousi sp. nov., C. besucheti sp. nov., C. bifoveolatum sp. nov., C. bilineatum sp. nov., C. bomjesus sp. nov., C. brachypterum sp. nov., C. ceylonicum sp. nov., C. clareae sp. nov., C. cryptophagoides sp. nov., C. davaoense sp. nov., C. deanei sp. nov., C. delicatum sp. nov., C. dewhurst sp. nov., C. dividuum sp. nov., C. dybasi sp. nov., C. eciton sp. nov., C. ecuadoriense sp. nov., C. faustum sp. nov., C. flavum sp. nov., C. franzi sp. nov., C. fraternum sp. nov., C. glabratum sp. nov., C. greensladei sp. nov., C. houailou sp. nov., C. ibicarense sp. nov., C. impressum sp. nov., C. inexspectatum sp. nov., C. insulare sp. nov., C. kolombangaricum sp. nov., C. lamington sp. nov., C. lawrencei sp. nov., C. loebli sp. nov., C. logunovi sp. nov., C. losbanos sp. nov., C. lucidulum sp. nov., C. luctuosum sp. nov., C. mahleri sp. nov., C. mindanao sp. nov., C. misellum sp. nov., C. modestum sp. nov., C. montanum sp. nov., C. murphyi sp. nov., C. mussardi sp. nov., C. noumeae sp. nov., C. obscenum sp. nov., C. obscurum sp. nov., C. obsoletum sp. nov., C. opacum sp. nov., C. orami sp. nov., C. peruviense sp. nov., C. pilosellum sp. nov., C. pinense sp. nov., C. plaumanni sp. nov., C. puncticolle sp. nov., C. riparium sp. nov., C. robustum sp. nov., C. rogeri sp. nov., C. seditiosum sp. nov., C. semicalvum sp. nov., C. similatum sp. nov., C. steeli sp. nov., C. subangulatum sp. nov., C. subfaustum sp. nov., C. sueae sp. nov., C. trangse sp. nov., C. upiense sp. nov., C. visendum sp. nov., C. waginicum sp. nov., C. werneri sp. nov. and C. yoruba sp. nov. The division of the genus is discussed and five species groups are established. The majority of the new species are the first records of Cissidium for the 28 countries represented.
Eight new species of Ecuadorian Ptiliidae are described and figured: Nossidium harrietae sp. nov., Ptenidium robustum sp. nov., P. tauriforme sp. nov., P. triangulum sp. nov., Notoptenidium brunneum sp. nov., Americoptilium nigrescens sp. nov., Discheramocephalus nigerrimus sp. nov. and D. striatus sp. nov. The insects were collected by J. McClarin, 2017–2018, in the Napo Province to the southeast of the capital Quito.
Micropterigidae is a family of Lepidoptera characterized by plesiomorphic morphological characters. Presently, this family consists of more than 22 genera and is distributed throughout all biogeographic realms. Vietomartyria Hashimoto & Mey, 2000 presently consists of six species mainly from South China. Two new species of the genus, V. wuyunjiena sp. nov. and V. maoershana sp. nov., are described herein from China; adult genitalia and wing venation are illustrated. Some biological accounts of the new species are also provided. A key to all described Vietomartyria species is given.
Three species of the genus Cryptochetum Rondani, 1875 from China are described and figured as new to science: C. euthyiproboscise sp. nov., C. glochidiatusum sp. nov., and C. longilingum sp. nov. An identification key to the known species of Cryptochetum from China is presented. The type specimens of the new species are deposited in the Henan Agricultural University.
This paper describes a novel species of the genus Heterophoxus Shoemaker, 1925 from Brazil. The material examined was collected during the Mini Biological Trawl Project, off southeast and southern Brazil's coast off the states of Rio de Janeiro, Santa Catarina, and Rio Grande do Sul, ranging in depth from 25 to 128 meters. The new species, H. shoemakeri sp. nov., is similar to H. videns K.H. Barnard, 1930; however, the former species can be distinguished by: pereopod 7 basis posterior margin serrate with acute teeth, merus posterior margin not produced; epimeral plate 3 posteroventral corner with a longer acute projection, posterior margin almost straight; uropod 3 outer ramus article 2 clearly notable in both sexes. For Brazilian waters, this is the second record of the genus and the first species described with type material from the country. Besides that, we provide an identification key for the world species of Heterophoxus and a distribution map for the new species.
A large, newly collected material of the genus Aquattuor from the Udzungwa Mts, Tanzania, has been studied. Two new species are described, viz., A. mollilobus sp. nov. from the Udzungwa Mts and A. nguruensis sp. nov. from the Nguru Mts, Tanzania. Aquattuor claudiahempae Enghoff & Frederiksen, 2015, is recorded as new for the Udzungwa fauna. Characters from the first pair of male legs are illustrated for all Aquattuor species and are shown to be useful for species discrimination. A new gonopodal difference between A. submajor Enghoff, 2015 and A. udzungwensis Enghoff, 2015 is described and illustrated. Intrageneric relationships and distribution patterns are discussed. A distribution map and a key to all Aquattuor species are presented.
New data of Torrenticolidae Piersig, 1902 are presented in this paper from Wuyishan National Nature Reserve, P. R. China. Three new species, Torrenticola suptilisrostrum Gu & Guo sp. nov., Torrenticola spinextension Gu & Guo sp. nov. and Torrenticola wuyiensis Gu & Guo sp. nov. are described and illustrated in detail, and one new record for Chinese fauna is given, Torrenticola dentifera Wiles, 1991, which was originally described from Malaysia. In addition, an updated key is provided to all species of Torrenticolidae in China.
A new species, Cryptomonas uralensis Martynenko, Gusev, Kulizin & Guseva sp. nov., is described from western slopes of the Ural Mountains (Russia) based on morphological and molecular data. Phylogenetic relationships inferred from nuclear SSU and LSU rDNA sequences show that the new species forms a clade with C. tetrapyrenoidosa Skuja emend. Hoef-Emden & Melkonian. Comparison of secondary structures of nuclear rDNA ITS2, including analysis of Compensatory Base Changes (CBC), confirms the separation between C. uralensis sp. nov. and C. tetrapyrenoidosa. Cell morphology and sizes of C. uralensis sp. nov. are very similar to C. tetrapyrenoidosa and C. pyrenoidifera, and C. uralensis sp. nov. may thus represent a species that can only be reliably identified using molecular data.
As a result of the integrative (morphology and DNA barcodes) revision of the Taiwanese species of the rove beetle genus Quedius belonging to the abnormalis group, in the subgenus Microsaurus, three valid hypogean species are reported for the fauna of this island: Quedius masuzoi Watanabe, 1989, Q. nishikawai Watanabe, 1991 and Q. adilus sp. nov. A new synonymy, Quedius masuzoi = Quedius chiangi Smetana, 1995 syn. nov., is established along with providing additional records, a larval description and bionomic information for this species. Finally, the biogeographic origin of the abnormalis group in Taiwan is discussed.
Neoxorides Clément, 1938 is a small genus of ichneumonid wasps that develop on wood-boring beetles. We here revise the Western Palaearctic species of the genus, with a special focus on Sweden. A new species, N. striatus Johansson sp. nov. is described. Neoxorides opacus (Kokujev, 1903) stat. rev. is excluded from the synonymy with N. nitens (Gravenhorst, 1829) and reinstated as a valid species. The species is redescribed from European material. Neoxorides kissi (Ulbricht, 1911) is confirmed as a junior synonym of N. opacus. A neotype for N. opacus is designated. New characters to distinguish N. varipes (Holmgren, 1860), N. collaris (Gravenhorst, 1829) and N. montanus Oehlke, 1966 are presented. Almost all of the species treated are supported by DNA-barcoding. An illustrated key to the species occurring in the Western Palaearctic is provided, a prerequisite for making the genus accessible for future ecological and conservation studies.
Sweat bees in the subgenus Lasioglossum (Dialictus) are one of the most diverse and abundant bee taxa, and a critically important component of bee biodiversity. Yet, the most basic taxonomic knowledge of these bees is lacking in many regions. As a step towards a better understanding of the L. (Dialictus) of the western Nearctic region, a revision of the ‘red-tailed’ L. (Dialictus) species was completed. Thirty-six species were revised, 20 of which are described as new, and two names are treated as junior subjective synonyms. Descriptions, figures, distribution maps, floral hosts, and keys to species for females and males are provided. The following 20 species are described as new: Lasioglossum (Dialictus) arenisaltans sp. nov., L. (D.) argammon sp. nov., L. (D.) austerum sp. nov., L. (D.) cactorum sp. nov., L. (D.) cembrilacus sp. nov., L. (D.) clastipedion sp. nov., L. (D.) clavicorne sp. nov., L. (D.) decorum sp. nov., L. (D.) festinum sp. nov., L. (D.) imbriumbrae sp. nov., L. (D.) julipile sp. nov., L. (D.) lilianae sp. nov., L. (D.) meteorum sp. nov., L. (D.) miltolepoides sp. nov., L. (D.) minckleyi sp. nov., L. (D.) perditum sp. nov., L. (D.) rufornatum sp. nov., L. (D. ) spivakae sp. nov., L. (D.) tessellatosum sp. nov., and L. (D.) torrens sp. nov. Previously unknown males of L. (D.) clematisellum (Cockerell, 1904), L. (D.) droegei Gibbs, 2009, L. (D.) kunzei (Cockerell, 1898), and L. (D.) pallidellum (Ellis, 1914) are described and figured for the first time. Lasioglossum (Dialictus) clarissimum (Ellis, 1914) (= Halictus clarissimus Ellis, 1914) and L. (D.) perexiguum (Sandhouse, 1924) (= Halictus (Chloralictus) perexiguus Sandhouse, 1924) are new subjective junior synonyms of L. mesillense (Cockerell, 1898) (= Halictus nymphalis var. mesillensis Cockerell, 1898). A lectotype specimen is newly designated for L. mesillense, for which the location of the type material has not previously been known. The following five new records for Mexico are reported: L. clematisellum, L. droegei, L. eophilus (Ellis, 1914), L. kunzei, and L. pallidellum.
Species of the genus Coelostoma Brullé, 1935 belonging to the subgenera Coelostoma s. str. and Holocoelostoma Mouchamps, 1958 from the Indian subcontinent are revised. Six species of Coelostoma s. str. and two species of Holocoelostoma are recognized, of which two are described as new: C. (Coelostoma) lyratum sp. nov. (India: Maharashtra) and C. (Coelostoma) nostocinum sp. nov. (India: Maharashtra, Goa, Karnataka, Kerala). Coelostoma (Coelostoma) fallaciosum Orchymont, 1936 and C. (Coelostoma) vividum Orchymont, 1936 were recorded for the first time from India and Bangladesh, respectively. Lectotypes are designated for C. aeneolum Régimbart, 1903 and Hydrobius stultus Walker, 1858. The previously confusing situation with Coelostoma (Holocoelostoma) stultum (Walker, 1858) and C. (Holocoelostoma) bhutanicum Jayaswal, 1972 is clarified based on new material of both species from India. Coelostoma sulcatum Pu, 1963 from China is removed from the synonymy of C. stultum and considered as a likely synonym of C. bhutanicum, a status which needs to be confirmed by a detailed study of type specimens. All species are (re)described and illustrated. Diagnosis of the subgenera of Coelostoma are modified in order to accommodate the species of the Indian subcontinent, resulting in narrowing down the concept of Coelostoma s. str. and widening the concept of Lachnocoelostoma Mouchamps, 1958.
We describe a new species of the genus Zhangixalus Li, Jiang, Ren & Jiang, 2019 from Ha Giang Province, Vietnam based on morphological and molecular data. In the molecular phylogenetic analyses, the new species is nested in the Zhangixalus duboisi (Ohler, Marquis, Swan & Grosjean, 2000) group, where it is sister to Z. duboisi with a genetic distance of 2.51%. The new species, Zhangixalus franki sp. nov., differs from its congeners by a combination of the following morphological characters: size large, SVL 77.9–85.8 mm in males; finger webbing formula I1-1II0-1III½-0IV; dorsal surface of head and body green with dark brown spots; ventral surface grey or dark grey, white stripe along edge of jaw, insertion of limbs, and along lateral ridges of fore and hind limbs and flank, separating upper green part from lower grey part. The new species occurs in evergreen montane tropical forests at an elevation of ca 1300 m a.s.l. The new discovery brings the total number of known species in the genus Zhangixalus to 38 and the species number reported from Vietnam to nine.
Two new species of Grandidierella Coutière, 1904 from Singapore waters are described based on specimens collected during the Comprehensive Marine Biodiversity Survey (CMBS) between 2010–2015. Grandidierella pawaiensis sp. nov. differs from the rest in the genus by a combination of characters; (1) gnathopod 1 carpus proximoventral corner with apically acute process, (2) inner face of carpus of gnathopod 1 proximoventral corner with curved process, (3) proximal margin of dactylus of gnathopod 1 male with excavation, and (4) gnathopod 2 basis anterior margin crenulate. Grandidierella sungeicina sp. nov. on the other hand, is distinguishable from other related species of Grandidierella found in the tropical region by its paired mid-dorsal projections on pereonites 2 to 5, 1 ventrodistal triangular projection on pereonite 4, and gnathopod 2 ischium with 1 large trapezoid projection.
The first Oriental species of the genus Trichopsomyia Williston, 1888, Trichopsomyia pilosa sp. nov. (Java), has been discovered and is now described. This Oriental species of Trichopsomyia has several characters strongly differing from the other species within this genus. It is hypothesized that it forms a separate group within Trichopsomyia. One species, Trichopsomyia formiciphila Downes, Skevington & Thompson, 2017, from Australia, is similar to the Oriental species described here, and the group is named after this first described species, hence the formiciphila group. The characters for a future phylogenetic analysis are discussed. The character states of the pilosity of the katepisternum and the shape of the metasternum hitherto used in a phylogenetic analysis of Syrphidae Latreille, 1802 are discussed too.