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Inverse modeling of circular lattices via orbit response measurements in the presence of degeneracy
(2023)
The number and location of beam position monitors (BPMs) and steerers with respect to the quadrupoles in a circular lattice can lead to degeneracy in the context of fitting linear optics and extracting lattice information from measured closed orbits. Furthermore, the measurement uncertainties due to the imperfection of BPMs and steerers can be propagated by the fitting process in ways that prohibit the successful extraction of discrepancies between lattice elements in the real machine and their description in the corresponding model. We systematically studied the influence of the placement of BPMs and steerers on the reconstruction of linear optics and corresponding lattice information. The derivative of orbit response coefficients with respect to the quadrupole strengths, the Jacobian, is derived as an analytical formula. This analytical version of the Jacobian is used to further derive the theoretical limitations of fitting linear optics from closed orbits in terms of the placement of BPMs and steerers. It is further demonstrated that when evaluating the Jacobian during the fitting procedure, the analytical version can be used in place of the conventional finite-difference computation. This allows for greatly improved efficiency when computing the Jacobian during each iteration of the fitting procedure. The approach is tested with large-scale simulations and the findings are verified by measurement data taken on SIS18 synchrotron at GSI Helmholtz Centre for Heavy Ion Research. The presented methods are of general nature and can be applied to other accelerator lattices as well. The fitting procedure by using the analytical Jacobian is tested in conjunction with various methods for mitigating quasidegeneracy and the results agree with those obtained by using the conventional Jacobian via finite-difference approximation.
Elliptic flow of heavy-flavor decay electrons in Au+Au collisions at √sNN = 27 and 54.4 GeV at RHIC
(2023)
We report on new measurements of elliptic flow (v2) of electrons from heavy-flavor hadron decays at mid-rapidity (|y|<0.8) in Au+Au collisions at sNN−−−√ = 27 and 54.4 GeV from the STAR experiment. Heavy-flavor decay electrons (eHF) in Au+Au collisions at sNN−−−√ = 54.4 GeV exhibit a non-zero v2 in the transverse momentum (pT) region of pT< 2 GeV/c with the magnitude comparable to that at sNN−−−√=200 GeV. The measured eHF v2 at 54.4 GeV is also consistent with the expectation of their parent charm hadron v2 following number-of-constituent-quark scaling as other light and strange flavor hadrons at this energy. These suggest that charm quarks gain significant collectivity through the evolution of the QCD medium and may reach local thermal equilibrium in Au+Au collisions at sNN−−−√=54.4 GeV. The measured eHF v2 in Au+Au collisions at sNN−−−√= 27 GeV is consistent with zero within large uncertainties. The energy dependence of v2 for different flavor particles (π,ϕ,D0/eHF) shows an indication of quark mass hierarchy in reaching thermalization in high-energy nuclear collisions.
The family of cubic noncentrosymmetric 3-4-3 compounds has become a fertile ground for the discovery of novel correlated metallic and insulating phases. Here, we report the synthesis of a new heavy fermion compound, Ce3Bi4Ni3. It is an isoelectronic analog of the prototypical Kondo insulator Ce3Bi4Pt3 and of the recently discovered Weyl-Kondo semimetal Ce3Bi4Pd3. In contrast to the volume-preserving Pt-Pd substitution, structural and chemical analyses reveal a positive chemical pressure effect in Ce3Bi4Ni3 relative to its heavier counterparts. Based on the results of electrical resistivity, Hall effect, magnetic susceptibility, and specific heat measurements, we identify an energy gap of 65-70 meV, about 8 times larger than that in Ce3Bi4Pt3 and about 45 times larger than that of the Kondo-insulating background hosting the Weyl nodes in Ce3Bi4Pd3. We show that this gap as well as other physical properties do not evolve monotonically with increasing atomic number, i.e., in the sequence Ce3Bi4Ni3-Ce3Bi4Pd3-Ce3Bi4Pt3, but instead with increasing partial electronic density of states of the d orbitals at the Fermi energy. To understand under which condition topological states form in these materials is a topic for future studies.
In relativistic heavy-ion collisions, a global spin polarization, PH, of Λ and Λ¯ hyperons along the direction of the system angular momentum was discovered and measured across a broad range of collision energies and demonstrated a trend of increasing PH with decreasing sNN−−−√. A splitting between Λ and Λ¯ polarization may be possible due to their different magnetic moments in a late-stage magnetic field sustained by the quark-gluon plasma which is formed in the collision. The results presented in this study find no significant splitting at the collision energies of sNN−−−√=19.6 and 27 GeV in the RHIC Beam Energy Scan Phase II using the STAR detector, with an upper limit of PΛ¯−PΛ<0.24% and PΛ¯−PΛ<0.35%, respectively, at a 95% confidence level. We derive an upper limit on the naïve extraction of the late-stage magnetic field of B<9.4⋅1012 T and B<1.4⋅1013 T at sNN−−−√=19.6 and 27 GeV, respectively, although more thorough derivations are needed. Differential measurements of PH were performed with respect to collision centrality, transverse momentum, and rapidity. With our current acceptance of |y|<1 and uncertainties, we observe no dependence on transverse momentum and rapidity in this analysis. These results challenge multiple existing model calculations following a variety of different assumptions which have each predicted a strong dependence on rapidity in this collision-energy range.
In relativistic heavy-ion collisions, a global spin polarization, PH, of Λ and Λ¯ hyperons along the direction of the system angular momentum was discovered and measured across a broad range of collision energies and demonstrated a trend of increasing PH with decreasing sNN−−−√. A splitting between Λ and Λ¯ polarization may be possible due to their different magnetic moments in a late-stage magnetic field sustained by the quark-gluon plasma which is formed in the collision. The results presented in this study find no significant splitting at the collision energies of sNN−−−√=19.6 and 27 GeV in the RHIC Beam Energy Scan Phase II using the STAR detector, with an upper limit of PΛ¯−PΛ<0.24% and PΛ¯−PΛ<0.35%, respectively, at a 95% confidence level. We derive an upper limit on the naïve extraction of the late-stage magnetic field of B<9.4⋅1012 T and B<1.4⋅1013 T at sNN−−−√=19.6 and 27 GeV, respectively, although more thorough derivations are needed. Differential measurements of PH were performed with respect to collision centrality, transverse momentum, and rapidity. With our current acceptance of |y|<1 and uncertainties, we observe no dependence on transverse momentum and rapidity in this analysis. These results challenge multiple existing model calculations following a variety of different assumptions which have each predicted a strong dependence on rapidity in this collision-energy range.
In relativistic heavy-ion collisions, a global spin polarization, PH, of Λ and Λ¯ hyperons along the direction of the system angular momentum was discovered and measured across a broad range of collision energies and demonstrated a trend of increasing PH with decreasing sNN−−−√. A splitting between Λ and Λ¯ polarization may be possible due to their different magnetic moments in a late-stage magnetic field sustained by the quark-gluon plasma which is formed in the collision. The results presented in this study find no significant splitting at the collision energies of sNN−−−√=19.6 and 27 GeV in the RHIC Beam Energy Scan Phase II using the STAR detector, with an upper limit of PΛ¯−PΛ<0.24% and PΛ¯−PΛ<0.35%, respectively, at a 95% confidence level. We derive an upper limit on the naïve extraction of the late-stage magnetic field of B<9.4⋅1012 T and B<1.4⋅1013 T at sNN−−−√=19.6 and 27 GeV, respectively, although more thorough derivations are needed. Differential measurements of PH were performed with respect to collision centrality, transverse momentum, and rapidity. With our current acceptance of |y|<1 and uncertainties, we observe no dependence on transverse momentum and rapidity in this analysis. These results challenge multiple existing model calculations following a variety of different assumptions which have each predicted a strong dependence on rapidity in this collision-energy range.
Evidence for the singly Cabibbo suppressed decay Λ+c→pπ0 is reported for the first time with a statistical significance of 3.7σ based on 6.0 fb−1 of e+e− collision data collected at center-of-mass energies between 4.600 and 4.843 GeV with the BESIII detector at the BEPCII collider. The absolute branching fraction of Λ+c→pπ0 is measured to be (1.56+0.72−0.58±0.20)×10−4. Combining with the branching fraction of Λ+c→nπ+, (6.6±1.2±0.4)×10−4, the ratio of the branching fractions Λ+c→nπ+ and Λ+c→pπ0 is calculated to be 4.2+2.2−1.9; this is an important input for the understanding of the decay mechanisms of charmed baryons. In addition, the absolute branching fraction of Λ+c→pη is measured to be (1.63±0.31stat±0.11syst)×10−3, which is consistent with previous measurements.
The differential cross section for Z0 production, measured as a function of the boson's transverse momentum (pT), provides important constraints on the evolution of the transverse momentum dependent parton distribution functions (TMDs). The transverse single spin asymmetry (TSSA) of the Z0 is sensitive to one of the polarized TMDs, the Sivers function, which is predicted to have the opposite sign in p+p →W/Z+X from that which enters in semi-inclusive deep inelastic scattering. In this Letter, the STAR Collaboration reports the first measurement of the Z0/γ∗ differential cross section as a function of its pT in p+p collisions at a center-of-mass energy of 510 GeV, together with the Z0/γ∗ total cross section. We also report the measurement of Z0/γ∗ TSSA in transversely polarized p+p collisions at 510 GeV.
Using an 𝑒+𝑒− collision data sample of (27.08±0.14)×108 𝜓(3686) events collected by the BESIII detector, we report the first observation of 𝜒𝑐𝐽→Ω−¯Ω+ (𝐽=0, 1, 2) decays with significances of 5.6𝜎, 6.4𝜎, and 18𝜎, respectively, where the 𝜒𝑐𝐽 mesons are produced in the radiative 𝜓(3686) decays. The branching fractions are determined to be ℬ(𝜒𝑐0→Ω−¯Ω+) = (3.51±0.54±0.29)×10−5, ℬ(𝜒𝑐1→Ω−¯Ω+)=(1.49±0.23±0.10)×10−5, and ℬ(𝜒𝑐2→Ω−¯Ω+)=(4.52±0.24±0.18)×10−5, where the first and second uncertainties are statistical and systematic, respectively.
Elliptic flow of heavy-flavor decay electrons in Au+Au collisions at √sNN = 27 and 54.4 GeV at RHIC
(2023)
We report on new measurements of elliptic flow (v2) of electrons from heavy-flavor hadron decays at mid-rapidity (|y|<0.8) in Au+Au collisions at sNN−−−√ = 27 and 54.4\,GeV from the STAR experiment. Heavy-flavor decay electrons (eHF) in Au+Au collisions at sNN−−−√ = 54.4\,GeV exhibit a non-zero v2 in the transverse momentum (pT) region of pT< 2\,GeV/c with the magnitude comparable to that at sNN−−−√=200 \,GeV. The measured eHF v2 at 54.4\,GeV is also consistent with the expectation of their parent charm hadron v2 following number-of-constituent-quark scaling as other light and strange flavor hadrons at this energy. These suggest that charm quarks gain significant collectivity through the evolution of the QCD medium and may reach local thermal equilibrium in Au+Au collisions at sNN−−−√=54.4\,GeV. The measured eHF v2 in Au+Au collisions at sNN−−−√= 27\,GeV is consistent with zero within large uncertainties. The energy dependence of v2 for different flavor particles (π,ϕ,D0/eHF) shows an indication of quark mass hierarchy in reaching thermalization in high-energy nuclear collisions.
Flow coefficients (v2 and v3) are measured in high-multiplicity p+Au, d+Au, and 3He+Au collisions at a center-of-mass energy of √sNN = 200 GeV using the STAR detector. The measurements are conducted using two-particle correlations with a pseudorapidity requirement of |η|< 0.9 and a pair gap of |Δη|>1.0. The primary focus of this paper is on the analysis procedures and methods employed, especially the subtraction of non-flow contributions. Four well-established non-flow subtraction methods are applied to determine vn, and their validity is verified using the HIJING event generator. The vn values are compared across the three collision systems at similar multiplicities, which allows for cancellation of final state effects and isolation of the impact of the initial geometry. While the v2 values display differences among these collision systems, the v3 values are largely similar, consistent with the expectations of subnucleon fluctuations in the initial geometry. The ordering of vn differs quantitatively from previous measurements obtained using two-particle correlations with a larger rapidity gap; this difference could be partially attributed to the effects of flow decorrelations in the rapidity direction.
Measurements of exclusive J/ψ, ψ(2s), and electron-positron (e+e−) pair photoproduction in Au+Au ultra-peripheral collisions are reported by the STAR experiment at √sNN=200 GeV. We report several first measurements at the Relativistic Heavy-Ion Collider, which are i) J/ψ photoproduction with large momentum transfer up to 2.2 (GeV/c)2, ii) coherent J/ψ photoproduction associated with neutron emissions from nuclear breakup, iii) the rapidity dependence of incoherent J/ψ photoproduction, iv) the ψ(2s) photoproduction cross section at mid-rapidity, and v) e+e− pair photoproduction up to high invariant mass of 6 GeV/c2. For measurement ii), the coherent J/ψ total cross section of γ∗+Au→J/ψ+Au as a function of the center-of-mass energy Wγ∗N has been obtained without photon energy ambiguities. The data are quantitatively compared with the Monte Carlo models STARlight, Sartre, BeAGLE, and theoretical calculations of gluon saturation with color glass condensate, nuclear shadowing with leading twist approximation, Quantum Electrodynamics, and the Next-to-Leading Order perturbative QCD. At the photon-nucleon center-of-mass energy of 25.0 GeV, the coherent and incoherent J/ψ cross sections of Au nuclei are found to be 71±10% and 36±7%, respectively, of that of free protons. These data provide an important experimental constraint for nuclear parton distribution functions and a unique opportunity to advance the understanding of the nuclear modification effect at the top RHIC energy.
For the search of the chiral magnetic effect (CME), STAR previously presented the results from isobar collisions (9644Ru+9644Ru, 9640Zr+9640Zr) obtained through a blind analysis. The ratio of results in Ru+Ru to Zr+Zr collisions for the CME-sensitive charge-dependent azimuthal correlator (Δγ), normalized by elliptic anisotropy (v2), was observed to be close to but systematically larger than the inverse multiplicity ratio. The background baseline for the isobar ratio, Y=(Δγ/v2)Ru(Δγ/v2)Zr, is naively expected to be (1/N)Ru(1/N)Zr; however, genuine two- and three-particle correlations are expected to alter it. We estimate the contributions to Y from those correlations, utilizing both the isobar data and HIJING simulations. After including those contributions, we arrive at a final background baseline for Y, which is consistent with the isobar data. We extract an upper limit for the CME fraction in the Δγ measurement of approximately 10% at a 95% confidence level on in isobar collisions at sNN−−−√=200 GeV.
We measure triangular flow relative to the reaction plane at 3 GeV center-of-mass energy in Au+Au collisions at RHIC. A significant v3 signal is observed for protons, whose magnitude increases for higher rapidity, higher transverse momentum, and more peripheral collisions. The triangular flow is essentially rapidity-odd with a rapidity slope at mid-rapidity, dv3/dy|(y=0), opposite in sign compared to the slope for directed flow. No significant v3 signal is observed for charged pions and kaons. Comparisons with models suggest that a mean field potential is required to describe these results, and that the triangular shape of the participant nucleons is the result of stopping and nuclear geometry.
We report results on an elastic cross section measurement in proton-proton collisions at a center-of-mass energy s√=510 GeV, obtained with the Roman Pot setup of the STAR experiment at the Relativistic Heavy Ion Collider (RHIC). The elastic differential cross section is measured in the four-momentum transfer squared range 0.23≤−t≤0.67 GeV2. We find that a constant slope B does not fit the data in the aforementioned t range, and we obtain a much better fit using a second-order polynomial for B(t). The t dependence of B is determined using six subintervals of t in the STAR measured t range, and is in good agreement with the phenomenological models. The measured elastic differential cross section dσ/dt agrees well with the results obtained at s√=546 GeV for proton--antiproton collisions by the UA4 experiment. We also determine that the integrated elastic cross section within the STAR t-range is σfidel=462.1±0.9(stat.)±1.1(syst.)±11.6(scale) μb.
The STAR Collaboration presents measurements of the semi-inclusive distribution of charged-particle jets recoiling from energetic direct-photon γdir and neutral-pion (π0) triggers in p+p and central Au+Au collisions at sNN−−−√=200 GeV over a broad kinematic range, for jet resolution parameters R=0.2 and 0.5. Medium-induced jet yield suppression is observed to be larger for R=0.2 than for 0.5, reflecting the angular range of jet energy redistribution due to quenching. The magnitude of suppression is similar for γdir- and π0-triggered data, which constrains the color-charge and path-length dependence of jet quenching. Theoretical model calculations incorporating jet quenching do not fully describe the measurements.
Cortical tracking of stimulus features (such as the envelope) is a crucial tractable neural mechanism, allowing us to investigate how we process continuous music. We here tested whether cortical and behavioural tracking of beat, typically related to rhythm processing, are modulated by pitch predictability. In two experiments (n=20, n=52), participants’ ability to tap along to the beat of musical sequences was measured for tonal (high pitch predictability) and atonal (low pitch predictability) music. In Experiment 1, we additionally measured participants’ EEG and analysed cortical tracking of the acoustic envelope and of pitch surprisal (using IDyOM). In both experiments, finger-tapping performance was better in the tonal than the atonal condition, indicating a positive effect of pitch predictability on behavioural rhythm processing. Neural data revealed that the acoustic envelope was tracked stronger while listening to atonal than tonal music, potentially reflecting listeners’ violated pitch expectations. Our findings show that cortical envelope tracking, beyond reflecting musical rhythm processing, is modulated by pitch predictability (as well as musical expertise and enjoyment). Stronger cortical surprisal tracking was linked to overall worse envelope tracking, and worse finger-tapping performance for atonal music. Specifically, the low pitch predictability in atonal music seems to draw attentional resources resulting in a reduced ability to follow the rhythm behaviourally. Overall, cortical envelope and surprisal tracking were differentially related to behaviour in tonal and atonal music, likely reflecting differential processing under conditions of high and low predictability. Taken together, our results show diverse effects of pitch predictability on musical rhythm processing.
In order to provide a reassessment of the Neotropical genus Pseudonannolene Silvestri, 1895, a cladistic analysis, biogeographic analysis, and taxonomic review were conducted in the present work. For the cladistic approach, 91 morphological characters were scored for 53 terminals as the ingroup and 10 as the outgroup. Three synapomorphies support the monophyly of the genus: presence of a longitudinal suture on the promentum, penial bases partially fused, and the internal branch of the gonopods surrounding the telopodite; and two homoplastic transformations: the lateral lobe of the collum densely striated and setae present up to the apical portion of the prefemoral process on the first leg-pair of males. The genus Pseudonannolene is recovered as sister-group of Epinannolene Brölemann, 1903 (Pseudonannoleninae). A total of 226 occurrence points were recorded for Pseudonannolene, with the majority of records from the Chacoan subregion, composed by Araucaria Forest, Atlantic, and Parana Forest provinces. The biogeographical searches using the Geographically explicit Event Model recovered two biogeographic reconstructions (cost of 79 000), with the vicariance events occurring more frequently in the deep clades, whereas sympatry and points of sympatry occurred in more inclusive clades. The first reconstruction recovered four vicariances, 13 sympatries, 4 points of sympatry, and 21 founder events, and the second reconstruction recovered four vicariances, 12–13 sympatries, 4–5 points of sympatry, and 21 founder events. The genus Pseudonannolene comprises 56 species, including 8 new species herein described: P. alata sp. nov., P. aurea sp. nov., P. bucculenta sp. nov., P. curvata sp. nov., P. granulata sp. nov., P. insularis sp. nov., P. morettii sp. nov., and P. nicolau sp. nov.; P. brevis Silvestri, 1902 and P. rugosetta Silvestri, 1897 are regarded as species inquirendae; a neotype of P. alegrensis Silvestri, 1897 is here proposed with male described for the first time. The following taxa are synonymized: P. canastra Gallo & Bichuette, 2020 and P. saguassu Iniesta & Ferreira, 2013 with P. ambuatinga Iniesta & Ferreira, 2013; P. marconii Iniesta & Ferreira, 2013 with P. longicornis (Porat, 1888); P. chaimowiczi Fontanetti, 1996, P. gogo Iniesta & Ferreira, 2013, P. rosineii Iniesta & Ferreira, 2014, P. taboa Iniesta & Ferreira, 2014, and P. longissima Iniesta & Ferreira, 2014 with P. microzoporus Mauriès, 1987; P. tricolor gracilis Brölemann, 1902 and P. tricolor rugosus Schubart, 1945 with P. tricolor Brölemann, 1902; P. auguralis Silvestri, 1902 with P. rocana Silvestri, 1902; and P. abbreviata Silvestri, 1902 with P. typica Silvestri, 1895. P. inops Brölemann, 1929 is proposed here as new status from P. bovei inops. A dichotomous identification key is presented to facilitate the species identification.
We report a new measurement of the production of electrons from open heavy-flavor hadron decays (HFEs) at mid-rapidity (|y|< 0.7) in Au+Au collisions at sNN−−−√=200 GeV. Invariant yields of HFEs are measured for the transverse momentum range of 3.5<pT<9 GeV/c in various configurations of the collision geometry. The HFE yields in head-on Au+Au collisions are suppressed by approximately a factor of 2 compared to that in p+p collisions scaled by the average number of binary collisions, indicating strong interactions between heavy quarks and the hot and dense medium created in heavy-ion collisions. Comparison of these results with models provides additional tests of theoretical calculations of heavy quark energy loss in the quark-gluon plasma.
Graphium chironides malayanum Eliot, 1982 was described as a taxon occurring sympatrically with G. bathycles bathycloides in Peninsular Malaysia. However, the validity of the subspecies has been questioned in a recent publication that was based on a study of DNA and morphology, implying that G. c. malayanum is a synonym of G. b. bathycloides and G. chironides is absent from the Peninsula. A re-examination of male wing morphology, genitalia and DNA shows that G. c. malayanum is a valid taxon distinguished from G. b. bathycloides by wider discal markings, a less falcate forewing, distinct differences in the arms of the harpe in the male genitalia and clearly divergent mtDNA COI genes. In the DNA analysis, G. c. malayanum formed a monophyletic clade closely related to G. chironides from China, and both were well-separated from the G. b. bathycloides clade. An examination of characters used in the previous study showed that the conclusions reached were due to misinterpretation of diagnostic characters, misidentification of specimens and the absence of G. c. malayanum among the specimens examined. When these characters were correctly interpreted, each specimen was readily assigned to the correct taxon. Diagnostic morphological characters are reclarified based on the current data.
Two new species of terrestrial isopods are described from iron ore caves in Brazil, within the Amazon biome, Circoniscus mendesi López-Orozco, Campos-Filho & Bichuette sp. nov. and C. xikrin López-Orozco, Campos-Filho & Carpio-Díaz sp. nov. (Scleropactidae). In addition, the knowledge of the distribution of Ctenorillo ferrarai Campos-Filho, Araujo & Taiti, 2014 (Armadillidae) is extended to Parauapebas, and Benthanoides tarzan Cardoso & Ferreira, 2023 to south area of the Campos Ferruginosos National Park, both in the State of Pará. Moreover, a distribution map and photographs of the species are given.
Three new species of Loxosceles Heinecken & Lowe, 1832 (Araneae, Sicariidae) from Brazilian caves
(2024)
Three new species of recluse spiders are described from Brazilian caves with both males and females. Loxosceles boqueirao Bertani & Gallão sp. nov. is found in the State of Bahia, in the Serra do Ramalho karst area, it belongs to the rufescens species group, and is closely related to L. cardosoi Bertani, von Schimonsky & Gallão, 2018 and L. carinhanha Bertani, von Schimonsky & Gallão, 2018 from the same karst area. Thus, there are now three species in the Serra do Ramalho karst area closely related, but noticeably distinct morphologically from other species of Loxosceles. Loxosceles planetaria Bertani & Gallão sp. nov. and L. bodoquena Bertani & Gallão sp. nov. are found in the State of Mato Grosso do Sul, in the Serra da Bodoquena karst area. They belong to the gaucho species group and are closely related to L. gaucho Gertsch, 1967. These are the first species of Loxosceles described from this karst area in the State of Mato Grosso do Sul. Loxosceles boqueirao sp. nov. and L. bodoquena sp. nov. bear some troglomorphisms and are, herein, proposed as troglobitic species, whereas L. planetaria sp. nov. is proposed as a troglophilic species. Brazil has now 22 described species of Loxosceles.
The remarkable sharpshooter Prodigiella silvanoi gen. et sp. nov. is described and illustrated (including the external form, color, male and female terminalia) from the Atlantic Forest of southern and southeastern Brazil (states of Paraná and Rio de Janeiro). The new genus can be distinguished from other Neotropical genera of the Cicadellini by a combination of various morphological features, including an asymmetrical aedeagus with a bifid shaft and peculiar basal and apical processes and ovipositor valvula II distinctly expanded beyond basal curvature, its dorsal margin with 35–40 teeth, and ventral margin without preapical prominence. A discussion comparing Prodigiella with superficially similar taxa of the genera Macugonalia Young, 1977, Ruppeliana Young, 1977, and Versigonalia Young, 1977 is provided. The discovery of this peculiar new genus indicates that much collecting work in the remaining parts of the Atlantic Forest is clearly and urgently needed.
Widely distributed phoretic blister beetles usually display unstructured geographic patterns of genetic diversity within species, resulting from recurrent long-dispersal events across their range. To investigate the extent of this pattern in the phoretic genus Lampromeloe, and particularly in western Mediterranean and European populations of L. variegatus, we performed: (1) a phylogeographic analysis based on fragments of the mitochondrial genes COI and 16S, and (2) a morphological revision based on qualitative and morphometric traits. Two allopatric lineages were recovered within L. variegatus, one distributed across Europe and the other in North Africa. These lineages are readily distinguishable based on morphological traits. We conclude that these lineages constitute two diagnosable evolutionary units and, consequently, we describe the North African populations as a new species, Lampromeloe pantherinus sp. nov. The new species is closely related to L. variegatus, from which it differs mainly by the elytral macrosculpture and by the shape of male genitalia. The elytral macrosculpture of L. pantherinus is somewhat similar to that of L. cavensis, a species widely distributed in the lowlands of the Mediterranean Basin. However, these two non-sister species can be distinguished by the morphology and macrosculpture of the pronotum and by the shape of the male genitalia.
The first sawfly from the Oligocene of Céreste (Southern France (Hymenoptera: Tenthredinidae)
(2024)
Luberotenthredo cerestensis gen. et sp. nov. is the first record of the sawfly family Tenthredinidae from the Oligocene of Céreste (Southern France). This taxon is described and illustrated based on a well-preserved specimen. This genus resembles the extant genus Perineura (subfamily Tenthredininae, tribe Perineurini) with which it shares forewing venation similarities and numerous morphological characters. This new taxon is the first fossil representative of the tribe Perineurini and can be used as a calibration point for future investigation of the diversification of the family Tenthredinidae.
The koinobiont endoparasitoid genus Triraphis Ruthe, 1855 (Rogadinae Foerster, 1863) is a group of braconid wasps that contains 74 species distributed along the Nearctic, Neotropical, Oriental and Palearctic regions. We amplified a fragment of the cytochrome oxidase subunit I (COI) for 19 specimens of Triraphis from the Chamela Biological Station (CBS), a region mainly composed of tropical dry forest near the Pacific coast of Jalisco, Mexico. Based on genetic distances among specimens of Triraphis from the CBS and all COI sequences of BINs assigned to Triraphis and Rogas Nees, 1819 available in the BOLDSYSTEMS database, we identified three clusters in the CBS that correspond with T. bradzlotnicki Sharkey, 2021, T. davidwahli Sharkey, 2021 and T. defectus Valerio, 2015, which were previously described from Costa Rica. Based on morphology, we identified individuals of T. fusciceps Cresson, 1869 and provided COI sequences of this species for the first time. Four genetic clusters of Triraphis correspond to four new species that are described here: T. kardia sp. nov., T. ocellatus sp. nov., T. divergens sp. nov. and T. luzabrilae sp. nov. Since T. bradzlotnicki and T. davidwahli were exclusively described with molecular data (COI), we morphologically described them based on Mexican specimens.
Palaeosphryon menatensis gen. et sp. nov., first unambiguous representative of the longhorn beetle subfamily Prioninae from the Paleocene of Menat (France), is described and illustrated. The new fossil is placed into the tribe Prionini, showing some similarities with some species of the extant genera Osphryon (Papua New Guinea) and Titanus (Brazil, Colombia, Guianas, Ecuador, Peru), viz. in general body shape, antennomere 3 as long as first and second together but shorter than the length of fourth plus fifth, elongate elytra, and small spines on the lateral margin of the pronotum disposed in a relatively similar way as in Osphryon. Nevertheless, the exact affinities of the new fossil within the Prionini remain uncertain because of the lack of a recent phylogenetic analysis in which it could be integrated. This fossil beetle is exceptional for its very large size, with a body 70 mm long. Some other large longhorn beetles have been found in the same outcrop, and are awaiting description. The positions of the previously described Cerambycidae from Menat are also discussed. This exceptional fauna of Cerambycidae is in accordance with the current palaeoenvironmental reconstruction for the Menat Konservat-Lagerstätte, as a small maar lake surrounded by a warm and humid, probably evergreen forest.
A new family of Cyphophthalmi with disjunct Mediterranean distribution, Parasironidae fam. nov., is proposed. The new family comprises four genera and seven species. Cimmerosiro gen. nov., Tirrenosiro gen. nov. and Ebrosiro gen. nov. are described as new genera, and Tirrenosiro axeli gen. et sp. nov., Cimmerosiro krivolutskyi gen. et sp. nov., C. juberthiei gen. et sp. nov. and C. rhodiensis gen. et sp. nov. as new species. Parasironidae stands out by a set of characteristics that we recognize as predominantly plesiomorphic. These characteristics and the present distribution indicate the great age of the family, probably early Mesosoic. We attribute its origin to the western part of the Cimmerian terrane, and its current distribution and diversification of the major clades to geotectonic events during the Mesozoic. Additionally, a new sensory organ (sensilla) has been discovered in Cyphophthalmi. This organ is located on the pedipalp coxae and is believed to have a potential hygroreceptive function.
The monotypic banchine ichneumonid genus Shortia is rediscovered after 39 years, based on two new species from India, far away from the type locality in Australia: S. karumban Ranjith sp. nov. and S. manjapulli Ranjith sp. nov., collected from the Western Ghats, India. The generic concept of Shortia is revised. Both new species and the type species of the genus, S. siccula Gauld, 1984, are illustrated. A taxonomic key for the identification of species of Shortia is provided and the possible causes for the disjunct distribution are discussed.
The “trachystreptoform” species of Spirostreptidae, i.e., species which would formerly have been ascribed to the tribe Trachystreptini, from the Udzungwa Mountains are (re)described, including one new genus and five new species: Attemsostreptus reflexus Akkari & Enghoff, 2019, A. cataractae Enghoff sp. nov., A. leptoptilos Enghoff sp. nov., A. julostriatus Enghoff sp. nov., Lophostreptus tersus (Cook, 1896) (= L. ptilostreptoides Carl, 1909 syn. nov.), L. magombera Enghoff sp. nov., and Udzungwastreptus marianae Enghoff gen. et sp. nov. The type material of Lophostreptus regularis Attems, 1909 (= L. tersus) is discussed. The discussion includes paragraphs on the classification and the Udzungwa fauna of Spirostreptidae, on grouping of the Udzungwa trachystreptoform species in relation to altitude, and on the possibly recent immigration of A. reflexus and L. tersus into the Udzungwa Mts.
Known by many names, the Chinese banyan (Ficus microcarpa L.) is a monoecious plant species originating from south-eastern Asia which has been introduced as an ornamental in numerous areas outside its native range, including the Mediterranean. Like every species of Ficus, it is associated with a series of chalcid wasp species, known as fig wasps. These species are distributed in the families Agaonidae, Epichrysomallidae, Eurytomidae, Ormyridae and Pteromalidae. In this publication, we describe a new species of Ormyrus Westwood, 1832 (Hymenoptera, Chalcidoidea, Ormyridae), O. microcarpae Askew & Koutsoukos sp. nov., reared from figs of F. microcarpa collected from Greece and Cyprus. The new species is compared with O. lini and O. watshami. This species is likely to be a parasitoid of Meselatus bicolor Chen, 1999 (Hymenoptera, Epichrysomallidae). In addition, the previously unknown female of O. lini is also described and illustrated. This publication constitutes the first report of species of Ormyrus associated with figs in Europe.
Six species of the spider genus Spinirta Jin & Zhang, 2020 from southern China (Araneae: Corinnidae)
(2024)
In the current study, six species of the genus Spinirta from southern China are recognized, four new species are described: S. hongyui Wang, Lu & Z.S. Zhang sp. nov. (♂♀), S. liuae Wang, Lu & Z.S. Zhang sp. nov. (♂♀), S. simianshan Wang, Lu & Z.S. Zhang sp. nov. (♂♀) and S. yintiaoling Wang, Lu & Z.S. Zhang sp. nov. (♂). Females of S. aviforma Jin & Zhang, 2020 (♂♀) and S. quadrata Jin & Zhang, 2020 (♂♀) are described for the first time. Photos of the body and copulatory organs, as well as the locality map are provided.
The ammonoids of the family Maenioceratidae from Givetian sedimentary rocks of the Anti-Atlas (Morocco) are investigated. The study is based on new collections stored in the Museum für Naturkunde, Berlin. The genera Maenioceras Schindewolf, 1933 and Afromaenioceras Göddertz, 1987 are revised; the genus Trimaenioceras is newly described. The species Maenioceras afroterebratum sp. nov., Maenioceras mzerrebense sp. nov., Maenioceras oufranense sp. nov., Maenioceras beckeri sp. nov., Afromaenioceras sulcatostriatum (Bensaïd, 1974), Afromaenioceras hiemale sp. nov., Afromaenioceras bensaidi sp. nov., Afromaenioceras brumale sp. nov., Afromaenioceras crassum (Bensaïd, 1974), Trimaenioceras klugi gen. et sp. nov., Trimaenioceras eculeus gen. et sp. nov., Trimaenioceras fuscina gen. et sp. nov. and Trimaenioceras paucum gen. et sp. nov. are described in detail.
A remarkable morphologically and genetically distinct species of the genus Ero C.L. Koch, 1836 is described based on both sexes from the cloud forest of the island of Saint Helena: Ero lizae sp. nov. Another new species, Ero natashae sp. nov., is also described on the basis of morphological differences in the male and female genitalia. Both species were initially reported a single species, Ero aphana (Walckenaer, 1802), from the island by Unzicker (1977).
High resolution images are provided for type specimens of Platygastroidea (Hymenoptera: Apocrita) in the Hope Entomological Collections, Oxford University. We formally resurrect Trissoscelio Kieffer revised status to accommodate T. bifasciata (Dodd) new combination, T. indica (Mani) new combination, T. nigriceps Kieffer revised combination, T. ruficeps Kieffer revised combination, and T. punctaticeps Kieffer revised combination. Paridris subplana (Dodd) new combination is transferred from Sceliacantha Dodd to Paridris Kieffer and treated as a senior synonym of P. coorgensis Sharma.
ZooBank registration. urn:lsid:zoobank.org:pub:EB97FE10-B01D-4D45-AF6E-D247ED0040BE
Two new species and one new genus of Lamiinae (Coleoptera: Cerambycidae) are described in Acanthocinini: Embera flava new species, new genus, from Panama; and Nealcidion lingafelteri new species from Costa Rica. The species-group name of Nealcidion napoensis Nascimento and McClarin, 2018 is corrected and morphological variations in Leptostylus cristulatus Bates, 1872 are reported. Trypanidius mimicavus Carelli, Monné, and Souza, 2013 is redescribed and transferred to Carphina Bates, 1872, forming the new combination Carphina mimicavus. Colombicallia curta Galileo and Martins, 1992 (Calliini) is recorded from Panama for the first time.
ZooBank registration. urn:lsid:zoobank.org:pub:7DD2EF0F-7DC9-4A85-83DA-10F4EE8CA8CB
A first-time analysis of taxonomically relevant characters, functional morphology, geographic distribution, ecoregion preference, and hypothetical host spiders of Pepsis basifusca Lucas (Hymenoptera: Pompilidae: Pepsinae) is presented. This analysis is compared with other Nearctic species in Vardy’s (2005) Pepsis menechma species-group, particularly P. cerberus Lucas and P. elegans Lepeletier which are suspected parasitoids of trapdoor spiders. Pepsis basifusca females differ from females of these species in possessing a rounded gena-postgena in dorsal view; straight mid and hind tibial spurs; short hind tibial inner spur; and short, very stout, and backward slanted hind tibial bristles. Pepsis basifusca Level III Ecoregions comprise mountains, plateaus, highlands, and tablelands, often at high elevation (~3,000–5,000 feet (914–1,524 meters), from Utah, Colorado, Kansas, and Missouri to Panama. Pepsis basifusca, the smallest Nearctic congener, should be expected to capture comparatively small mygalomorph spiders like some other species in Vardy’s (2005) Pepsis menechma species-group. Based on taxonomic, morphological, biogeographical, and potential host spider criteria, P. basifusca should probably be removed from this group and transferred to another species-group.
ZooBank registration. urn:lsid:zoobank.org:pub:4D3BF37C-3358-4C25-B56A-104C57301AE2
Identifications of recently collected Eucnemidae (Coleoptera: Elateroidea) borrowed from two collections have resulted in the discovery of two new species: Entomophthalmus abbreviatus Otto (Cuba) and Trigonopleurus cordobaalfaroi Otto (Guatemala and Louisiana, USA). Images of the two newly described species along with two New World Entomophthalmus Bonvouloir species for comparative purposes are included.
ZooBank registration. urn:lsid:zoobank.org:pub:E83B2AC6-33ED-4680-8F2E-4E455A26D4C8
Five new state records for Idaho in the genera Agrilus Curtis, Anthaxia Eschscholtz, and Buprestis Linnaeus (Coleoptera: Buprestidae) are included herein, with brief comments thereon. Two of the species are believed to be introduced. Agrilus liragus Barter and Brown is elevated to a full species, resurrected status.
ZooBank registration. urn:lsid:zoobank.org:pub:3AE72784-F368-45A2-AC90-B31E48D0D5CC
Sixteen species of Issidae (Hemiptera: Fulgoromorpha) were recorded in a single day of collecting in Dong Son-Ky Thuong Nature Reserve, Quang Ninh Province, Vietnam, of which a checklist is provided. Among them, eight species are new to science and described in different tribes and subtribes: in the Hemisphaeriini Hemisphaeriina: Gergithoides olivaceus sp. nov., Hemisphaerius bresseeli sp. nov. and Neogergithoides scapularis sp. nov.; Hemisphaeriini Mongolianina: Mongoliana vietnamica sp. nov.; Kodaianellini: Kodaianella mua sp. nov.; Parahiraciini Parahiraciina: Pusulissus quangninhensis sp. nov. and Rostrolatum curviceps sp. nov.; Sarimini: Pseudocoruncanius obliquus sp. nov. The new genus Melichergithus gen. nov. is described to accommodate Gergithus gravidus Melichar, 1906 and the new combination Melichergithus gravidus (Melichar, 1906) gen. et comb. nov. is proposed; a black form of the species is recorded for the first time and described. Two species, Longieusarima lunulia Wang, Bourgoin & Zhang, 2017 and Parallelissus fuscus Meng, Qin & Wang, 2020, are recorded from Vietnam for the first time, while another species Maculergithus luteomaculatus (Constant & Pham, 2016) is recorded for the first time from Quang Ninh Province. Four species were represented by single females that could not be identified to species level, belonging to the genera Clypeosmilus Gnezdilov & Soulier-Perkins, 2017, Fortunia Distant, 1909 (two species) and Kodaianellissus Wang, Bourgoin & Zhang, 2017. The five following genera are recorded from Vietnam for the first time: Kodaianella Fennah, 1956, Rostrolatum Che, Zhang & Wang, 2020, Longieusarima Wang, Bourgoin & Zhang, 2017, Parallelissus Meng, Qin & Wang, 2020 and Kodaianellissus. The type material of Hemisphaerius rufovarius Walker, 1858 and its junior synonyms H. scymnoides Walker, 1862, H. testaceus Distant, 1906 and H. virescens Distant, 1906 was studied and the male terminalia described; as a result, H. rufovarius is removed from the fauna of Vietnam and China. Hemisphaerius bipunctatus Melichar, 1906 is also removed from the fauna of Vietnam after examination of the corresponding specimens. This study provides a 25% increase in the number of Issidae of Vietnam; a complete list of the 51 species recorded from the country, as well as a map of the number of recorded species per province, are given and discussed.
A new family-level taxon of deep-sea isopods, Basoniscus hikurangi gen. et sp. nov., was recovered from the Hikurangi Plateau in the deep sea off eastern New Zealand. The broad-bodied, eyeless, seemingly unremarkable isopod was unusual in its possession of features that characterize two different families: the shallow water Joeropsididae Nordenstam, 1933 and the deep-sea Haploniscidae Hansen, 1916. An analysis of superfamily Janiroidea G.O. Sars, 1897 was conducted to establish the affinities of the species. Multiple analyses were done using unweighted and implied weighted characters. Existing families were well supported, with B. hikurangi intermediate between Joeropsididae and Haploniscidae. The new species, however, cannot be placed in either family owing to its lack of important defining synapomorphies of each family. As a result, Basoniscidae fam. nov. was created to contain this new species. That rocky hard substrates are undersampled is another implication. Our understanding of deep-sea species richness will not be accurate until more efforts are made to survey these habitats, especially more sites in the southern hemisphere. These gaps in our knowledge of the deep sea impairs any general claims about the distribution of biodiversity on a global scale. This find demonstrates that museums hold underused but valuable resources for understanding and describing the biodiversity of the deep sea.
Type specimens of Oedionychina Chapuis, 1875 described by Fabricius from the Kiel collection are examined and illustrated. Lectotypes are designated for the following species: Chrysomela albicollis Fabricius, 1787; Chrysomela nobilitata Fabricius, 1787; Chrysomela quadrifasciata Fabricius, 1787; Chrysomela quadriguttata Fabricius, 1781; Galleruca atomaria Fabricius, 1801; Galleruca decemguttata Fabricius, 1801; Galleruca fasciata Fabricius, 1798; Galleruca humeralis Fabricius, 1801; Galleruca lunata Fabricius, 1801; Galleruca nitida Fabricius, 1801; Galleruca obsoleta Fabricius, 1801; Galleruca petaurista Fabricius, 1801; Galleruca quadrinotata Fabricius, 1798; Galleruca sellata Fabricius, 1801. The species status is restored for Chrysomela quadriguttata Fabricius, 1781 and Alagoasa areata (Germar, 1824) comb. nov. The following new combinations are proposed: Phenrica quadriguttata (Fabricius, 1781), Asphaera nitida (Fabricius, 1801), Phenrica obsoleta (Fabricius, 1801), Alagoasa areata areata (Germar, 1824), Alagoasa areata decempunctata (Latreille, 1833), Alagoasa areata escuintla Bechyné, 1955, Alagoasa areata macromela Bechyné, 1958, Alagoasa areata praecessa Bechyné, 1959, Alagoasa areata recuperata Bechyné, 1959; all comb. nov. New placement: Galleruca avicenniae Fabricius, 1792 is removed from Alticini and placed in Galerucini incertae sedis; Galleruca trifasciata Fabricius, 1801 is removed from Chrysomelidae and placed in genus Ora Clark, 1865 (Scirtidae Fleming, 1821).
The genus Spaeleoleptes was proposed by H. Soares in 1966 to accommodate the first Brazilian troglobitic species of harvestmen, Spaeleoleptes spaeleus H. Soares, 1966. In this work, we redescribe this species, including digital images of the type material and drawings of the male genitalia. Since its description, Spaeleoleptes has remained monotypic, and after 56 years, herein is described the second species of the genus, the troglobitic Spaeleoleptes gimli sp. nov. Both species share sexually dimorphic legs I and II with modified regions and swelling on the tibiae and patellae I and II; a penis with robust conductors covering all or part of the capsula interna and a capsula interna with two lateral projections. They are clearly separated by the shape of the modified region of the tibia; by the presence of an apical projection on the apical lamina of the pars distalis in S. spaeleus; and the lateral projections of the capsula interna, which is flattened in S. gimli. Spaeleoleptes gimli greatly increases the distributional range of the genus, as it is now recorded from caves located in two Brazilian phytophysiognomies from the Cerrado of Minas Gerais to the Caatinga of Bahia.
Several adult female monstrilloid copepods, collected in March 2022 from the protected reef area of Xcalak, on the southern part of the Mexican Caribbean coast, proved to belong to two undescribed species of Monstrilla Dana, 1849. They are described here as M. xcalakensis sp. nov. and M. annulata sp. nov., partly by use of scanning electron microscopy. Females of the two species are generally similar but differ in: (1) antennular segmental structure, (2) antennular armature and ornamentation, (3) structure and setation of the fifth leg, (4) number and modifications of the caudal setae, and (5) integumental ornamentation. Comparison with congeneric species revealed distinctive features for both species that support their status as new. These two species are interesting additions to the reef-dwelling monstrilloid copepod fauna of the Mexican Caribbean and confirm Monstrilla as the most diverse genus of monstrilloids in this area, now represented by nine species.
SAFE Update February 2024
(2024)
This thesis contains three theoretical works about certain aspects of the interplay of electronic correlations and topology in the Hubbard model.
In the first part of this thesis, the applicability of elementary band representations (EBRs) to diagnose interacting topological phases, that are protected by spatial symmetries and time-reversal-symmetry, in terms of their single-particle Matsubara Green’s functions is investigated. EBRs for the Matsubara Green’s function in the zero-temperature limit can be defined via the topological Hamiltonian. It is found that the Green’s function EBR classification can only change by (i) a gap closing in the spectral function at zero frequency, (ii) the Green’s function becoming singular i.e. having a zero eigenvalue at zero frequency or (iii) the Green’s function breaking a protecting symmetry. As an example, the use of the EBRs for Matsubara Green’s functions is demonstrated on the Su-Schriefer-Heeger model with exact diagonalization.
In the second part the Two-Particle Self-Consistent approach (TPSC) is extended to include spin-orbit coupling (SOC). Time-reversal symmetry, that is preserved in the presence of SOC, is used to derive new TPSC self-consistency equations including SOC. SOC breaks spin rotation symmetry which leads to a coupling of spin and charge channel. The local and constant TPSC vertex then consists of three spin vertices and one charge vertex. As a test case to study the interplay of Hubbard interaction and SOC, the Kane-Mele-Hubbard model is studied. The antiferromagnetic spin fluctuations are the leading instability which confirms that the Kane-Mele-Hubbard model is an XY antiferromagnet at zero temperature. Mixed spin-charge fluctuations are found to be small. Moreover, it is found that the transversal spin vertices are more strongly renormalized than the longitudinal spin vertex, SOC leads to a decrease of antiferromagnetic spin fluctuations and the self-energy shows dispersion and sharp features in momentum space close to the phase transition.
In the third part TPSC with SOC is used to calculate the spin Hall conductivity in the Kane-Mele-Hubbard model at finite temperature. The spin Hall conductivity is calculated once using the conductivity bubble and once including vertex corrections. Vertex corrections for the spin Hall conductivity within TPSC corresponds to the analogues of the Maki-Thompson contributions which physically correspond to the excitation and reabsorption of a spin, a charge or a mixed spin-charge excitation by an electron. At all temperatures, the vertex corrections show a large contribution in the vicinity of the phase transition to the XY antiferromagnet where antiferromagnetic spin fluctuations are large. It is found that vertex corrections are crucial to recover the quantized value of −2e^2/h in the zero-temperature limit. Further, at non-zero temperature, increasing the Hubbard interaction leads to a decrease of the spin Hall conductivity. The results indicate that scattering of electrons off antiferromagnetic spin fluctuations renormalize the band gap. Decreasing the gap can be interpreted as an effective increase of temperature leading to a decrease of the spin Hall conductivity.
Highlights
• Three ecological groups were identified based on distributional patterns.
• Old assessments were confirmed with the latest occurrence data.
• For each group, we derived different population trends in times of global change.
• Global change elevates importance of vector-borne diseases.
• Our results serve as base for effective Simuliidae monitoring.
Abstract
The black fly genus Simulium includes medically and ecologically important species, characterized by a wide variation of ecological niches largely determining their distributional patterns. In a rapidly changing environment, species-specific niche characteristics determine whether a species benefits or not. With aquatic egg, larval and pupal stages followed by a terrestrial adult phase, their spatial arrangements depend upon the interplay of aquatic conditions and climatic-landscape parameters in the terrestrial realm. The aim of this study was to enhance the understanding of the distributional patterns among Simulium species and their ecological drivers. In an ecological niche modelling approach, we focused on 12 common black fly species with different ecological requirements. Our modelling was based on available distribution data along with five stream variables describing the climatic, land-cover, and topographic conditions of river catchments. The modelled freshwater habitat suitability was spatially interpolated to derive an estimate of the adult black flies' probability of occurrence. Based on similarities in the spatial patterns of modelled habitat suitability we were able to identify three biogeographical groups, which allows us to confirm old assessments with current occurrence data: (A) montane species, (B) broad range species and (C) lowland species. The five veterinary and human medical relevant species Simulium equinum, S. erythrocephalum, S. lineatum, S. ornatum and S. reptans are mainly classified in the lowland species group. In the course of climatic changes, it is expected that biocoenosis will slightly shift towards upstream regions, so that the lowland group will presumably emerge as the winner. This is mainly explained by wider ecological niches, including a higher temperature tolerance and tolerance to various pollutants. In conclusion, these findings have significant implications for human and animal health. As exposure to relevant Simulium species increases, it becomes imperative to remain vigilant, particularly in investigating the potential transmission of pathogens.
Highlights
• We study dormancy in the ‘rare mutation’ regime of stochastic adaptive dynamics.
• We first derive the polymorphic evolution sequence, based on prior work.
• Our evolutionary branching criterion extends a result by Champagnat and Méléard.
• In a classical model dormancy can favour evolutionary branching.
• Dormancy also affects several more population characteristics.
Abstract
In this paper, we investigate the consequences of dormancy in the ‘rare mutation’ and ‘large population’ regime of stochastic adaptive dynamics. Starting from an individual-based micro-model, we first derive the Polymorphic Evolution Sequence of the population, based on a previous work by Baar and Bovier (2018). After passing to a second ‘small mutations’ limit, we arrive at the Canonical Equation of Adaptive Dynamics, and state a corresponding criterion for evolutionary branching, extending a previous result of Champagnat and Méléard (2011).
The criterion allows a quantitative and qualitative analysis of the effects of dormancy in the well-known model of Dieckmann and Doebeli (1999) for sympatric speciation. In fact, quite an intuitive picture emerges: Dormancy enlarges the parameter range for evolutionary branching, increases the carrying capacity and niche width of the post-branching sub-populations, and, depending on the model parameters, can either increase or decrease the ‘speed of adaptation’ of populations. Finally, dormancy increases diversity by increasing the genetic distance between subpopulations.