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Coastal species are under considerable threat from recreational activities and climate change. The tiger beetle Cicindela hirticollis Say (Coleoptera: Cicindelidae) was recorded historically from 30 locations along the shores of New York City and Long Island, New York. We conducted surveys for extant populations of this species at 40 sites from 1989 to 2010. Adults of C. hirticollis were found at 13 beaches. Only four sites had 40 or more adults of C. hirticollis active at the time the beach was surveyed. No beetles were detected on the large coastal beaches that were formerly occupied by this species. Many coastal beaches of New York receive heavy human foot and vehicle traffic and are therefore unlikely to provide suitable habitat for C. hirticollis without a shift in beach management that recognizes the potential of some beaches as wilderness systems capable of supporting the full array of beach-dependent species.
The primary types of Onciderini Thomson, 1860 (Coleoptera: Cerambycidae: Lamiinae) deposited at the Carnegie Museum of Natural History (CMNH) are catalogued and illustrated. Data on the original combination, current name, and type locality are verified and presented. There are 36 primary types of Onciderini including 12 in Oncideres Lacordaire, 1830; four in Hypsioma Audinet-Serville, 1835; three each in Hesychotypa Thomson, 1868; Cacostola Fairmaire and Germain, 1859; and Sternycha Dillon and Dillon, 1945. A brief history of the CMNH is also presented.
A key is provided for the twelve taxonomic groups within the genus Therates Latreille (Coleoptera: Cicindelidae). The chennelli group is reviewed here and a key to the 53 species known for this group is provided. Each species is illustrated, and a detailed description of its morphology and distributional records are provided. Therates khaoyaii, Th. pearsoni, Th. safraneki, Th. schuelei and Th. sigridgeissleri are described as new to science. Therates differens Sawada and Wiesner, 1999 is placed into synonymy under Th. concinnus Gestro, 1888. Therates tonkinensis kubani Wiesner, 1988 is placed into synonymy under Th. tonkinensis Horn, 1902 and Th. belokobylskiyi Matalin and Wiesner, 2006 is placed into synonymy under Th. haucki Moravec and Wiesner, 2001. Lectotypes are designated for Th. waagenorum Horn, 1900, Th. clavicornis Horn, 1902, Th. tonkinensis Horn, 1902 and Th. annandalei Horn, 1908. Therates baolocensis Wiesner, 1996 is elevated to species rank.
The ostracod genus Bennelongia De Deckker & McKenzie, 1981 is endemic to Australia and New Zealand. Extensive sampling in Western Australia (WA) revealed a high specific and largely undescribed diversity. Here, we describe seven new species belonging to the B. barangaroo lineage: B. timmsi sp. nov., B. gnamma sp. nov., B. hirsuta sp. nov., B. ivanae sp. nov., B. mcraeae sp. nov., B. scanloni sp. nov. and B. calei sp. nov., and confirm the presence of an additional species, B. dedeckkeri, in WA. For five of these eight species, we could construct molecular phylogenies and parsimonious networks based on COI sequences. We also tested for cryptic diversity and specific status of clusters with a statistical method based on the evolutionary genetic species concept, namely Birky’s 4 theta rule. The analyses support the existence of these five species and a further three cryptic species in the WA B. barangaroo lineage. The molecular evidence was particularly relevant because most species described herein have very similar morphologies and can be distinguished from each other only by the shape, size and position of the antero-ventral lapel on the right valve, and, in sexual populations, by the small differences in shape of the hemipenes and the prehensile palps in males. Four species of the WA B. barangaroo lineage occur in small temporary rock pools (gnammas) on rocky outcrops. The other four species are mainly found in soft bottomed seasonal water bodies. One of the latter species, B. scanloni sp. nov., occurs in both claypans and deeper rock pools (pit gnammas). All species, except for B. dedeckkeri, originally described from Queensland, have quite clearly delimited distributions in WA. With the seven new species described here, the genus Bennelongia now comprises 25 nominal species but several more await formal description.
Two new genera and five new species of Selachinematidae are described from the New Zealand upper continental slope (350-1240 m depth). Synonchiella rotundicauda sp. nov. is characterised by cephalic setae 0.25 cbd long, mandibles each with two pairs of hooks and two wing-like projections laterally, eight cup-shaped pre-cloacal supplements and short rounded tail. Pseudocheironchus gen. nov. is similar to Cheironchus, but differs from the latter in having a cuticle without lateral differentiation, cephalic setae only slightly longer than the outer labial sensillae, and a posterior buccal cavity with three equal mandibles. Pseudocheironchus ingluviosus gen. et sp. nov. is characterised by mandibles with eight blunt teeth, multispiral amphideal fovea with five turns, and a short rounded tail. Males of this new species with 17-19 cup-shaped pre-cloacal supplements. Males of the genus Cobbionema are described for the first time; C. trigamma sp. nov. is characterised by four long cephalic setae and six smaller outer labial setae in one circle, six rhabdions surrounding the anterior buccal cavity, each with two pairs of pointed projections at their posterior extremities, posterior buccal cavity widening posteriorly, with three pairs of rhabdions fused posteriorly and widening anteriorly, males with two testes pointing anteriorly and with reflexed posterior testis, and no pre-cloacal supplements. Gammanema agglutinans sp. nov. is characterised by a short, stout body often covered in adhering mucus and detritus, cuticle with minute spines, leaf-shaped somatic setae with ducts, sexual dimorphism in the shape of the amphideal fovea (loop-shaped in males and spiral in females), posterior buccal cavity with three pairs of broad, column-shaped rhabdions fused anteriorly, intestine cells with orange-brown granules, and small tubular pre-cloacal supplements. Bendiella gen. nov. is most similar to Halichoanolaimus, but differs from the latter, and all other genera of the family Selachinematidae, in having a cuticle with lateral differentiation consisting of longitudinal rows of larger dots, and from all other genera of the Choniolaiminae in lacking pre-cloacal supplements. Bendiella thalassa gen. et sp. nov. is characterised by amphideal fovea with 5.25 turns, anterior buccal cavity with twelve rhabdions, each with a pair of pointed projections at posterior extremity, posterior buccal cavity with three Y-shaped pairs of slender rhabdions fused from two thirds of distance from anterior ends, and conico-cylindrical tail.
New Zealand species of Iphimediidae, Amphipoda, are revised. Based on new material from the Chatham Rise, east of New Zealand, two new species are described in detail: Labriphimedia meikae sp. nov. and Labriphimedia martinae sp. nov. A key to the six species belonging to three genera of New Zealand Iphimediidae is provided.
A new status (as subgenera of Diostracus Loew, 1861) for Sphyrotarsus Mik, 1874, Lagodechia Negrobov & Tsurikov, 1996 and Ozmena Özdikmen, 2010 stat. nov. is proposed. A new species, Diostracus (Sphyrotarsus) kustovi sp. nov., is described from the Russian Caucasus. The following recombinations (comb. nov.) are also proposed: Diostracus (Sphyrotarsus) argyrostomus (Mik, 1874); D. (S.) caucasicus (Negrobov, 1965); D. (S.) hervebazini (Parent, 1914); D. (S.) hessei (Parent, 1914); D. (S.) hygrophilus (Becker, 1891); D. (S.) leucostomus (Loew, 1861); D. (S.) parenti (Hesse, 1933); D. (Lagodechia) spinulifer Negrobov & Tsurikov, 1988; and D. (Ozmena) stackelbergi (Negrobov, 1965). A key to ten Diostracus species inhabiting the West Palaearctic Region is provided.
Two new colourful species of direct-developing frogs of the genus Pristimantis are described from the summit of two isolated tepuis (sandstone table mountains) in the Eastern Pantepui District of the Guiana Shield highlands. Pristimantis jamescameroni sp. nov. is described from the summit of Aprada-tepui from 2557-2571 m elevation, and P. imthurni sp. nov. is described from the summit of Ptaritepui at 2471 m elevation. Both species share the absence of a differentiated tympanic membrane and external tympanic annulus (but presence of tiny pharyngeal ostia), the presence of nuptial pads in males, and the presence of lateral fringes on fingers and toes, a combination of characters that immediately distinguishes them from all other known Pantepui congeners. The two new species are morphologically similar to each other and are phylogenetically closely related, but they can be distinguished based on colour pattern and morphological characters such as head proportions, dorsal skin texture, and condition of the supratympanic fold. The IUCN conservation status of the new species is considered as Endangered (EN) owing to their apparent very restricted ranges. The number of described Pristimantis species occurring exclusively on tepui (and faunistically related granitic mountains) summits and upper slopes now reaches eleven.
This paper summarizes current knowledge about West African pholcids. West Africa is here defined as the area south of 17°N and west of 5°E, including mainly the Upper Guinean subregion of the Guineo-Congolian center of endemism. This includes all of Senegal, The Gambia, Guinea Bissau, Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana, Togo and Benin. An annotated list of the 14 genera and 38 species recorded from this area is given, together with distribution maps and an identification key to genera. Five species are newly described: Anansus atewa sp. nov., Artema bunkpurugu sp. nov., Leptopholcus kintampo sp. nov., Spermophora akwamu sp. nov., and S. ziama sp. nov. The female of Quamtana kitahurira is newly described. Additional new records are given for 16 previously described species, including 33 new country records. Distribution patterns of West African pholcids are discussed, as well as possible explanations for relatively low West African pholcid species diversity as compared to Central and East Africa.
After our taxonomic revision of Ootheca Chevrolat, 1837, and the description of Oothecoides Kortenhaus & Wagner, 2011 and Ootibia Kortenhaus & Wagner, 2012, it became clear that a further four galerucine species, closely related to the above named taxa, form a distinct monophyletic group, that constitutes a new genus, Oosagitta gen. nov. with O. anningae sp. nov., O. geescheae sp. nov., O. melanopicta sp. nov. and O. thomasi sp. nov.. Exosoma angolensis Laboissière, 1939, the type species of the new genus, and Ergana minuta Laboissière, 1937 are newly transferred to Oosagitta gen. nov. All species of Oosagitta gen. nov. are characterized by a broad body and pronotum, a more or less convex dorsum and short legs, and as such are most similar to the other above named genera. The antennae of Oosagitta gen. nov. are distinctly longer than those of Ootheca, Oothecoides and Ootibia. Genital structures of the males allow a reliable identifi cation of the genus. (Re-) descriptions are given for all species, including semi-schematic illustrations depicting the habitus outline, shape of the basal antennomeres and the median lobe. Photographs of the name-bearing types and distribution maps are provided.
The osteology of Gladiopycnodus karami gen. et sp. nov., of Monocerichthys scheuchzeri gen. et sp. nov. and of Rostropycnodus gayeti gen. et sp. nov., three new fossil fishes from the marine Cenomanian (Late Cretaceous) of Lebanon, is studied in detail. Some of their cranial characters and the presence of a postcoelomic bone clearly refer these fishes to the order Pycnodontiformes. However, they differ from all other described Pycnodontiformes by two important characters. Their snout is elongated as a rostrum, formed by the enlarged prefrontal and the toothless premaxilla, with this premaxilla sutured by its upper margin to the lower margin of the prefrontal. Their pectoral fin is replaced by a strong spine articulated with the cleithrum. These two apomorphies justify the erection of a new family, the Gladiopycnodontidae. The skull of Monocerichthys scheuchzeri sp. nov. does not differ greatly from a classical pycnodontiform skull and this species seems to be the more primitive member of this new family. Gladiopycnodus karami gen. et sp. nov. and Rostropycnodus gayeti gen. et sp. nov. are much more specialized. They share some apomorphies not present in Monocerichthys scheuchzeri gen. et sp. nov., i. e., an extremely long rostrum and an elongated first anal pterygiophore that sustains with the postcoelomic bone a strong and long anal spine. Gladiopycnodontidae fam. nov. and Coccodontidae share a series of apomorphies that justify the erection of a new superfamily, Coccodontoidea, grouping these two families.
A new species, Memecylon pseudomegacarpum (Melastomataceae), is described from southern Peninsular Thailand, Peninsular Malaysia and Singapore. This taxon was previously known under the misapplied name M. megacarpum, which is now considered endemic to Borneo. Memecylon pseudomegacarpum sp. nov. differs from M. megacarpum in having smaller leaves (8–)10.5–17(–22.5) cm rather than (10–)17–28(–35) cm long, with an elliptic lamina (not lanceolate) with a raised mid-rib (not sunken) and a marginal vein which is 2–4 mm from the margin (not 5–12 mm). Both species have similar flowers and share large (c. 15 mm diameter) globose fruits.
Two new species of the family Trefusiidae, viz., Trefusia piperata sp. nov. and Trefusialaimus idrisi sp. nov., are described from the crest of the Chatham Rise, Southwest Pacific Ocean (350 m water depth). The present study provides the first species records for this family in the region. Trefusia and Trefusialaimus comprise twenty and three valid species, respectively. A key to males of Trefusia is provided.
The molecular phylogeny of Miliusa (Annonaceae) is reconstructed, with 27 (of ca. 50) species included, using a combination of seven plastid markers (rbcL exon, trnL intron, trnL-F spacer, matK exon, ndhF exon, psbA-trnH spacer, and ycf1 exon) constituting ca. 7 kb. In addition, two new species of Miliusa are described from the Malesian area: M. butonensis sp. nov. from Buton Island, Indonesia and M. viridifl ora sp. nov. from Papua New Guinea. The former is included in the molecular phylogenetic analysis. The reconstructed phylogeny corresponds well to the informal morphological grouping proposed earlier. A revised key to 13 Austro-Malesian species of Miliusa is provided.
Type material of the species of Alloxysta described by Cameron and Fergusson and deposited in the Natural History Museum of London has been revised. Seven species are considered valid: Alloxysta abdera Fergusson, 1986, A. basimacula (Cameron, 1886), A. crassa (Cameron, 1889), A. mullensis (Cameron, 1883), A. piceomaculata (Cameron, 1883), A. pleuralis (Cameron, 1879) and A. semiaperta Fergusson, 1986. A. basimacula, A. crassa, A. maculicollis (Cameron, 1886), A. perplexa (Cameron, 1889) and A. piceomaculata are here removed from synonymy with A. macrophadna (Hartig, 1841). A. rufi ceps (Cameron, 1883) is removed from synonymy with A. victrix (Westwood, 1833). A. caledonica (Cameron, 1886) and A. perplexa are here synonymized with A. basimacula. A. maculicollis, A. ruficeps and A. ruficollis (Cameron, 1883) are here synonymized with A. castanea (Hartig, 1841). A. ancylocera (Cameron, 1886) was correctly synonymized with A. fuscicornis (Hartig, 1841), A. curvicornis (Cameron, 1883) was correctly synonymized with A. victrix and A. filicornis (Cameron, 1889) was correctly synonymized with A. macrophadna. Complete redescriptions and illustrations are given for valid species. A key for all the Alloxysta species found so far in Great Britain is given.
Several populations of four known species of the genus Pungentus (P. clavatus, P. engadinensis, P. marietani and P. silvestris), collected in the wild and in cultivated soils from the Iberian Peninsula, are studied. Detailed redescriptions and morphometrics are presented for each species. Illustrations are provided, including line drawings, light microscopy pictures of the four species as well as scanning electron microscopy observations of P. engadinensis. The Iberian populations are compared to type and other known populations, and new data are given that provide a better characterization of these taxa. Pungentus engadinensis is the most widely distributed species in the Iberian Peninsula.
We revise the species-level taxonomy of the Crematogaster (Crematogaster) degeerispecies-assemblage, a group of related ants occuring in Madagascar and the wider Malagasy region, and further provide an identification key to all species-groups of the genus Crematogaster in this region. Within the C. degeeri-assemblage, we recognize twelve species based upon morphological data from worker, queen and male ants, as well as genetic data from the barcode region of cytochrome oxidase I. Seven new species are described: Crematogaster alafara Blaimer sp. nov., C. bara Blaimer sp. nov., C. mafybe Blaimer sp. nov., C.maina Blaimer sp. nov., C. malahelo Blaimer sp. nov., C. masokely Blaimer sp. nov., C. ramamy Blaimer sp. nov. Crematogaster tricolor Gerstäcker, 1859 (stat. rev.) and C. dentata Dalla Torre, 1893 (stat. nov.) are raised to species level, and the following new synonymies are proposed: Crematogaster degeeri lunaris Santschi, 1928 as a synonym of C. degeeri Forel, 1886; Crematogaster sewelli improba Forel, 1907 and C. sewelli mauritiana Forel, 1907 as synonyms of C. dentata Dalla Torre, 1893, and C. pacifi ca Santschi, 1919 as a synonym of C. lobata Emery, 1895. Species descriptions, images, and distribution maps and identification keys based on worker ants, as well as on queen ants where available, are presented for all twelve species. In addition, we present a molecular gene tree for cytochrome oxidase I and summarize levels of sequence divergence within and between species of the C. degeeri-species-assemblage. Our findings are discussed in the light of previous work on Malagasy Crematogaster ants.
The species richness of the frugivorous fruit fly fauna of western African (in particular of Ivory Coast, Ghana, Togo, Benin and Nigeria) is discussed. The diversity is compared at a national level and between the ecoregions within the national boundaries of the study area. A new species, Dacus goergeni sp. nov. is described and additional taxonomic notes are presented.