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Numerous footprints areas has been found, since 1962, in Permian, Triassic and Hettango-Sinemurian. All show precised oriented trackways. So that they were undestood like walking ways but their precise meaning is still enigmatic.
Footprints are numerous in France where they found from Carboniferous to Quaternary levels. Those of Middle Triassic (Anisian-Ladinian) are especialy interesting because they are tridactyl footprints made probably by first dinosaurs. First skelettons are known from Upper Ladinian / lower Carnian.
In the inter/supratidal algal mats, the early diagenesis products a progressive induration. It is enough to preserve all the deformations affecting the mats, including footprints of vertebrates, tracks and burrows of small invertebrates, and physical actions (desiccation cracks).