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A new phylogenetic hypothesis is proposed for the relationships among the species within the genus Docosia Winnertz, 1863, based on a combined analysis of five DNA markers (28S, ITS2, COI, COII and CytB). Five new species are described, Docosia anatolica Ševčík sp. nov. from Turkey, D. japonica Kurina sp. nov. from Japan, D. peloponnensis Ševčík sp. nov. from Greece, D. svanetica Kurina sp. nov. from Caucasus and D. polyspina Kurina sp. nov. from the Russian Far East. New country records of the following species are presented: D. diutina Plassmann, 1996 (Turkey), D. flavicoxa Strobl, 1900 (Georgia), D. gilvipes (Haliday in Walker, 1856) (Georgia), D. kerkini Kurina & Ševčík, 2011 (Bulgaria), D. moravica Landrock, 1916 (Georgia), D. pannonica Lastovka & Ševčík 2006 (Georgia) and D. rameli Kurina & Ševčík, 2011 (Slovakia).
A new monotypic genus of flatid planthoppers (Hemiptera: Fulgoromorpha: Flatidae), Medleria gen. nov., is described for Medleria caudata gen. et sp. nov. (type species) from the island of Socotra (Yemen). Habitus, male and female external and internal genital structures of the new species are illustrated and compared with similar taxa. Medleria caudata gen. et sp. nov. is probably endemic to Socotra where it is known to date from a small area of the Dixam mountain plateau only.
Review of Synapsis Bates (Scarabaeidae: Scarabaeinae: Coprini), with description of a new species
(2010)
Presented are a checklist, a discussion of and keys to species groups and their constituent species, and a description of one new species: Synapsis horaki. The species Synapsis cambeforti Krikken and S. thoas Sharp are synonymized with S. ritsemae Lansberge, Balthasar’s synonymy of S. yunnana Arrow with S. tridens Sharp is revived, and the status of six recently described species is left unresolved because of insufficient data.
Presented is a checklist of the world Trogidae (Coleoptera: Scarabaeoidea) including synonyms, geographic distributions, type repositories where known, lists of valid species by genera and subgenera, citations of all papers containing original descriptions, and a supplemental literature section containing works on various other aspects of the family. The Literature Cited and Supplemental Literature sections combine to form a comprehensive bibliography.
The spider genus Zaitunia Lehtinen, 1967 (Araneae, Filistatidae) is revised. It was found to include 24 species distributed in the Eastern Mediterranean, Middle East and Central Asia: ♀ Z. afghana (Roewer, 1962) (Afghanistan), ♀ Z. alexandri Brignoli, 1982 (Iran), ♀ Z. akhanii Marusik & Zamani, 2015 (Iran), ♂♀ Z. annulipes (Kulczyński, 1908) (Cyprus), ♂♀ Z. beshkentica (Andreeva & Tyshchenko, 1969) (Tajikistan, Uzbekistan), ♀ Z. brignoliana sp. nov. (Iran), ♂♀ Z. ferghanensis sp. nov. (Kyrgyzstan, Uzbekistan), ♀ Z. feti sp. nov. (Turkmenistan), ♀ Z. halepensis sp. nov. (Syria), ♀ Z. huberi sp. nov. (Afghanistan), ♀ Z. inderensis Ponomarev, 2005 (Kazakhstan), ♂♀ Z. kunti sp. nov. (Cyprus, Turkey), ♂♀ Z. logunovi sp. nov. (Kazakhstan, Kyrgyzstan), ♂♀ Z. maracandica (Charitonov, 1946) (Uzbekistan, Kazakhstan), ♂♀ Z. martynovae (Andreeva & Tyshchenko, 1969) (Tajikistan, Turkmenistan), ♀ Z. medica Brignoli, 1982 (Iran), ♂♀ Z. minoica sp. nov. (Greece), ♀ Z. minuta sp. nov. (Uzbekistan), ♀ Z. persica Brignoli, 1982 (Iran), ♂ Z. psammodroma sp. nov. (Turkmenistan), ♂♀ Z. schmitzi (Kulczyński, 1911), the type species (Egypt, Israel), ♂♀ Z. spinimana sp. nov. (Kazakhstan, Turkmenistan), ♂♀ Z. wunderlichi sp. nov. (Kyrgyzstan) and ♀ Z. zonsteini Fomichev & Marusik, 1969 (Kazakhstan). Twelve above-listed species are newly described, and males of Z. annulipes, Z. beshkentica, Z. maracandica and Z. martynovae are described for the first time. Two new combinations are established: Z. annulipes (Kulczyński, 1908) comb. nov., ex Filistata, and Pholcoides monticola (Spassky, 1941) comb. nov., ex Zaitunia. New data on distribution of the considered taxa are provided.
The genus Raveniola Zonstein, 1987 is found to be represented in Western Asia by 16 species: ♂♀ R. adjarica sp. nov. (Georgia), ♂ R. anadolu sp. nov. (Turkey), ♂ R. arthuri Kunt & Yağmur, 2010 (Turkey), ♂ R. birecikensis sp. nov. (Turkey), ♂♀ R. dunini sp. nov. (Armenia, Azerbaijan, Iran), ♂♀ R. hyrcanica Dunin, 1988 (Azerbaijan), ♂ R. marusiki sp. nov. (Iran), ♂ R. mazandaranica Marusik, Zamani & Mirshamsi, 2014 (Iran), ♂♀ R. micropa (Ausserer, 1871) (Turkey), ♀ R. nana sp. nov. (Turkey), ♂♀ R. niedermeyeri (Brignoli, 1972) (Iran), ♂♀ R. pontica (Spassky, 1937) (Russia, Georgia), ♀ R. sinani sp. nov. (Turkey), ♂♀ R. turcica sp. nov. (Turkey), ♂♀ R. vonwicki Zonstein, 2000 (Iran) and ♂♀ R. zaitzevi (Charitonov, 1948) (Azerbaijan, Georgia) = ♀ Brachythele recki Mcheidze, 1983, syn. nov. Eight species are described as new; others are redescribed from types and/or conspecific material. Males of R. micropa and R. zaitzevi, hitherto unknown, are described for the first time. Data on the variability, relationships, distribution and ecology of all considered species are also provided.
The Central Asian spider genus Anemesia Pocock, 1895 is rediagnosed and revised. The genus was found to contain 14 species: ♂♀ A. andreevae sp. nov. (Uzbekistan, Tajikistan); ♂♀ A. birulai (Spassky, 1937) (Turkmenistan); ♂♀ A. castanea sp. nov.; ♂♀ A. incana Zonstein, 2001, ♂♀ A. infumata sp. nov.; ♂♀ A. infuscata sp. nov.; ♂♀ A. karatauvi (Andreeva, 1968) (all Tajikistan); ♂ A. koponeni Marusik, Zamani & Mirshamsi, 2014 (Iran); ♂♀ A. oxiana sp. nov.; ♂♀ A. pallida sp. nov.; ♂ A. parvula sp. nov. (all Tajikistan); ♂♀ A. pococki sp. nov. (Turkmenistan); ♂♀ A. sogdiana sp. nov. (Uzbekistan, Tajikistan) and ♂♀ A. tubifex (Pocock, 1889), the type species (Afghanistan, Turkmenistan). Nine species are newly described; others are redescribed from types and/or conspecific material. Males of A. tubifex and females of A. birulai, hitherto unknown, are described for the first time. Data on the variability, relationships, distribution, and ecology of all considered species are provided.
Allopatric alpine populations of Phtheochroa frigidana s. lat. (Lepidoptera, Tortricidae) are reviewed. In addition to traditional diagnostic characters of external morphology, the genitalia structures of everted vesicae in male genitalia and DNA barcodes are analysed. This new approach supports the existence of five rather than two species in Europe: Phtheochroa schawerdae (Rebel, 1908) comb. nov. (Dinaric Mts, Rila Mts, Pirin Mts) = P. drenowskyi (Rebel, 1916) syn. nov.; P. alpinana sp. nov. (SW Alps); P. apenninana sp. nov. (Apennines); P. frigidana (Guenée, 1845) stat. rev. (Pyrenees) = P. flavidana (Guenée, 1845) = P. sulphurana (Guenée, 1845) = P. andorrana (Millière, 1865); P. cantabriana sp. nov. (Cantabrian Mts). In order to stabilize the nomenclature, a neotype for Eupoecilia frigidana is designated.
The Scarabaeus subgenus Scarabaeolus Balthasar, 1965 (Scarabaeidae: Scarabaeinae: Scarabaeini) is defined and nine new species are described, six from the Republic of South Africa [S. (S.) fragilis sp. n., S. (S.) krugeri sp. n., S. (S.) lizleri sp. n., S. (S.) orientalis sp. n., S. (S.) rugosipennis sp. n., S. (S.) similis sp. n.] and one each from Angola [S. (S.) cunene sp. n.], Namibia [S. (S.) namibensis sp. n.] and Kenya [S. (S.) werneri sp. n.], bringing the number of recorded species up to 41. Species accounts listing original descriptions, subsequent accounts, type localities, type repositories, and geographic distributions are provided for all the species. The status of S. (S.) reichei Waterhouse, 1890 and of S. (S.) bohemani Harold, 1868, both formerly synonymized and reinstated, are discussed. It is concluded that S. (S.) reichei is synonymous with S. (S.) canaliculatus Fairmaire, 1888, whereas S. (S.) bohemani differs from S. (S.) palemo Olivier, 1789 in the color of antennal club and shape of paramere tips, and is upheld pending availability of molecular data.
The taxonomic position of Onthophagus (Palaeonthophagus) lemuroides d’Orbigny, 1898 and Onthophagus
(Palaeonthophagus) fortigibber Reitter, 1909 is discussed (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini).
A key to the species is given. Photos of type specimens of the two taxa and significant chromatic varieties, and
drawings of aedeagi are presented.
A re-description and new records of Onthophagus viriditinctus Reitter, 1892 (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini), an uncommon species from Iran, are provided. The taxonomic position and some nomenclatural problems are discussed. The placement in the subgenus Exonthophagus Kabakov, 2006 is proposed. Images of the male, female, aedeagus and drawings of lamella copulatrix of Onthophagus viriditinctus and Onthophagus haroldi Ballion, 1871, the only other species included in the subgenus, are supplied. A key for distinguishing the two species is provided.
A new Iranian, probably pholeophilous species of the scarabaeine genus Onthophagus Latreille, 1802, O. roessneri n. sp., is described, illustrated, and its placement in the semicornis group within the subgenus Palaeonthophagus Zunino, 1979 is discussed. A key to the species of the group is presented.
This revision concerns a small group of Western Palaearctic Copris species (Coleoptera: Scarabaeoidea:
Scarabaeidae) distinguished by having three lateral teeth on the foretibae. According to the literature, this group consists of four taxa: Copris armeniacus Faldermann, 1835, C. felschei Reitter, 1892, C. pueli Mollandin de Boissy, 1905 and C. umbilicatus Abeille de Perrin, 1901. Copris armeniacus is herein deemed a species inquirenda, and C. felschei, for which a neotype is designated, is deemed valid. Copris umbilicatus is recorded for the fi rst time from Turkey. A key to all species of the genus Copris known from the Western Palaearctic is provided. Variability of the cephalic and pronotal armature, and morphology of the parameres are illustrated.
In 2014, 56 localities in four provinces of Lesser Himalaya in Pakistan were studied. A total of 28 species have been recorded. A female of the data deficient, threatened species Coeliccia vacca was recorded from Charhaan. The record of Drepanosticta carmichaeli is a new addition to the list of Odonata of Pakistan, and expand the range of this species further to the west. The taxonomical status of Ischnura aurora aurora – considered common in Pakistan, following baseline literature of Fraser (1933) – now turns out to be Ischnura aurora rubilio.
Damselflies recorded before the administrative partition of the Indian Subcontinent and now housed at National Insect Museum (NIM), Islamabad were reviewed and catalogued. This collection is the divided part of National PUSA Collection (NPC) transferred to the Pakistan during 1947. Data for this collection had never been available or published. A record of 104 taxa is reported herein. Few of the species were found double named, misidentified and not updated as per valid classification. Some of the specimens were found unidentified. All such issues were resolved by following regional literature.
A new species complex, the eparmata complex, is established within the subgenus Phortica s. str., based on eight known and five new species, all of which are endemic to the Oriental Region: P. bipartita (Toda & Peng, 1992), P. eparmata (Okada, 1977), P. lanuginosa Chen & Toda, 2007, P. latipenis Chen & Gao, 2005, P. pangi Chen & Wen, 2005, P. setitabula Chen & Gao, 2005, P. unipetala Chen & Wen, 2005 and P. zeta Chen & Toda, 2007; P. jadete sp. nov., P. kava sp. nov., P. mengda sp. nov., P. wongding sp. nov. and P. yena sp. nov. A key to all species of this complex is provided. Barcoding sequences (mitochondrial COI gene) were obtained for 22 specimens of five known and the five abovementioned new species. The intra- and inter-specific pairwise K-2P (Kimura’s two-parameter) distances of COI were determined. Phylogenetic analysis was performed using Bayesian inference based on COI sequences, confirming the monophyletic status of the eparmata complex, which is distinct from the species complexes of magna, omega, variegata and another two ungrouped species.
A new mesoserphid wasp from the Middle Jurassic of northeastern China (Hymenoptera, Proctotrupoidea)
(2017)
A new genus and species of Mesoserphidae (Hymenoptera), Juraserphus modicus gen. et sp. nov., is described based on a well-preserved fossil specimen from the Middle Jurassic Jiulongshan Formation of northeastern China. It is characterized by the following forewing features: the forking of Rs+M located approximately one-third of the distance between 1m-cu and 2r-rs, both 1cu-a and 2cu-a antefurcal; 1-M more than twice as long as 1m-cu and hind wing with cells r and rm closed. In addition, it has a short ovipositor, only extending slightly beyond the metasomal apex. Its new morphological characters broaden the diversity of Mesoserphidae in the Mesozoic and provide new insights into the evolution and relationships of Mesoserphidae.
The paper compiles records from four excursions to study the Odonata fauna of southern Guizhou, China. Between 2007 and 2010 in Xiaoqikong Park and Maolan National Nature Reserve, 104 taxa have been recorded. Some interesting species are discussed, compared with sibling taxa, and information on habitats and habits is given.
Between 2009 and 2016, a total of 174 Odonata species (Tab. 1) have been recorded in the Dehong Dai and Jingpo Autonomous Prefecture, China. 21 of these species are new records for China, and additional 26 taxa have to be described as new to science. Brief comments on selected species refer to morphological characters, distribution and seasonality.
Four new species of the genus Coecobrya, C. gejianbangi sp. nov., C. annulata sp. nov., C. ciliata sp. nov., and C. oculata sp. nov., are described from Guangxi caves as the representative of the genus in China. Coecobrya oculata sp. nov. of the boneti-group has 1+1 eyes and a serrate outer edge of the unguiculus. The other three species, devoid of eyes and with a tiny outer tooth on the unguiculus, are assigned here to the tenebricosa-group, assuming that the large tooth on the unguiculus is transformed into a tiny one in cave-obligate species. Clypeal chaetae in Entomobryoidea are systematically surveyed for the first time, and are found to be well diversified at species level. They have a potential taxonomical value in discriminating taxa of morphologically conserved groups.
A preliminary checklist of Calpini is provided, incorporating corrections and changes to publication dates and nomenclature as presented in the checklists of Poole (1989), Fibiger and Lafontaine (2005), and Holloway (2005). Culasta Moore is removed from synonymy with Calyptra Ochsenheimer. Eudocima talboti (Prout) and Graphigona antica Walker are placed in synonymy with E. cajeta (Cramer) and G. regina (Guenée), respectively. Africalpe Krüger, Ferenta Walker, Gonodonta Hübner, Graphigona Walker, Oraesia Guenée, and Tetrisia Walker, are added to the tribe based on shared characters. The genera Cecharismena Möschler, Goniapteryx Perty, Pharga Walker, Phyprosopus Grote, Psammathodoxa Dyar, and Radara Walker are removed and considered incertae sedis. Hemiceratoides and Phyllodes are not considered to be members of Calpini.
The present study is based on the genus Cotesia Cameron,1891 collected from Khuzestan Province in the Southwestern part of Iran during 2016–2017. Nine species (+200 specimens) of the genus Cotesia were collected and identified. We recognised three new species, which we describe and illustrate here: Cotesia elongata Zargar & Gupta sp. nov., C. khuzestanesis Zargar & Gupta sp. nov. and C. zagrosensis Zargar & Gupta sp. nov. Two species are recorded for the first time from Iran: Cotesia cynthiae (Nixon, 1974) and C. glabrata (Telenga, 1955). A faunistic list with species distribution in Iran, a modified key to include the new species and brief diagnoses for the new records from Iran are provided.
The Iranian species of the genus Anomalon Panzer, 1804 (Hymenoptera: Ichneumonidae: Anomaloninae) are reviewed. Four species, Anomalon amseli (Hedwig, 1961), A. chinense (Kokujev, 1915), A. cruentatum (Geoffroy, 1785) and A. narinae Zardouei & Rakhshani sp. nov., are found to occur in Iran. The female of A. amseli is described for the first time. Anomalon chinense is a new record for Iran. A key to the known Anomalon species of Iran is provided.
A new fossil diatom species, Tertiarius minutulus sp. nov., is described from a sediment sequence DEEP-5045-1 of Lake Ohrid. The species is characterized by small valves (3.0–8.0 μm) with a round shape, a marginal area with radially arranged costae and a central uneven area with scattered areolae. Externally, the alveoli are occluded by cribra perforated by irregularly arranged pores. Internally, the alveoli are simple and areolae are occluded with domed cribra. The marginal fultoportulae are situated on costae close to the valve margin, located on every 5th or 7th thick internal costa. One to three fultoportulae are present on the valve face, each surrounded by two to three satellite pores. One rimoportula is present, positioned on a costa at the valve face / mantle junction. The species is compared with morphologically similar taxa and a detailed differential diagnosis is provided. Tertiarius minutulus sp. nov. is known only as a fossil taxon from Lake Ohrid, with a first occurrence during the early stages of lake basin development.
Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae)
(2014)
The North American species of the genus Eusphalerum Kraatz (Coleoptera, Staphylinidae, Omaliinae) are revised. The taxonomic history, natural history, geographical distribution of the genus, characters, species groups, diversity, and biogeography of North American species are presented. Two main phylogenetic lineages and 13 species groups are provisionally recognized. The following new synonymies are proposed: Eusphalerum farrarae (Hatch, 1944) = E. lawrencei Hatch, 1957; Eusphalerum californicum (Fauvel, 1878) = E. atriventre (Casey, 1894), = E. nigerrimum (Casey, 1894), = E. gilvipenne (Casey, 1894), = E. dichroum (Fall, 1922), = E. bonnelli (Hatch, 1944), = E. lunae Hatch, 1957; Eusphalerum fraternum (Casey, 1894) = E. minskae (Hatch, 1944); Eusphalerum rugulosum (Mäklin, 1853) = E. grayae (Hatch, 1944); Eusphalerum orientale (Bernhauer, 1912) = E. frosti (Bernhauer, 1928). The following lectotypes are designated: E. subangulatum (Casey), E. californicum (Fauvel), E. gilvipenne (Casey), E. diversicolle (Casey), E. convexum (Fauvel), E. fraternum (Casey), E. horni (Fauvel), E. orientale (Bernhauer), E. pothos (Mannerheim), and E. punctatum (Casey). The following new species are described: Eusphalerum pilosum (California); E. klimaszewskii (British Columbia); E. chatzimanolisi (California); E. carolinensis (Kentucky, Missouri, North Carolina, Ohio, Pennsylvania, Tennessee, Virginia); E. caterinoi (California); E. luteipes (California); E. thayeranum (Alberta, British Columbia, Idaho, Indiana (doubtful record), Oregon, Washington); E. margaretae (Tennessee); E. newtoni (British Columbia, Oregon, California); E. parvispiculum (California, Oregon); E. uncinatum (British Columbia, California, Oregon, Washington). Eusphalerum lapponicum (Mannerheim, 1830) is excluded from the North American fauna. The following new combination is proposed: Xylodromus segmentarius (Mäklin, 1852: 322) (ex Omalium), wrongly attributed to Eusphalerum in the literature. Omalium marginatum Say, 1832 is considered a doubtful species, probably not Eusphalerum. A key to the 27 recognized North American Eusphalerum species and a catalog of the species are provided.
Cochlostoma revised: the subgenus Lovcenia Zallot et al., 2015
(Caenogastropoda, Cochlostomatidae)
(2018)
Five species of the subgenus Lovcenia of Cochlostoma (Cochlostomatidae) are recognized, three of which are described as new to science: C. (L.) tropojanum sp. nov., C. (L.) jakschae sp. nov. and C. (L.) lanatum sp. nov. A lectotype is designated for C. (L.) erika (A.J. Wagner, 1906). The shell and the genital tracts are described for all species and the distributional data are summarized.
The classification of the largest subfamily of leafhoppers, Deltocephalinae, including 38 tribes, 923 genera, and 6683 valid species, is reviewed and revised. An updated phylogeny of the subfamily based on molecular (28S, Histone H3) and morphological data and an expanded taxon sample (37 taxa not included in previous analyses) is presented. Based on the results of these analyses and on the morphological examination of many representatives of the subfamily, the classification of the tribes and subtribes of Deltocephalinae is revised. Complete morphological descriptions, illustrations, lists of the included genera, and notes on their distribution, ecology, and important vector species are provided for the 38 recognized tribes and 18 subtribes. A dichotomous key to the tribes is provided. All names in the taxonomic treatments are hyperlinked to online resources for individual taxa which are supported by a comprehensive database for Deltocephalinae compiled using the taxonomic database software package 3I. The online functionality includes an interactive key to tribes and subtribes and advanced database searching options. Each taxon (subspecies through subfamily) has a unique taxon webpage providing nomenclatural information, lists of included taxa, an automated description (if available), images (if available), distributional information, bibliographic references and links to outside resources. Some observations and trends regarding the history of taxonomic descriptions in Deltocephalinae are reported. Four new tribes are described: Bahitini tribe nov. (25 genera), Bonsapeiini tribe nov. (21 genera), Phlepsiini tribe nov. (4 genera), and Vartini tribe nov. (7 genera). The circumscription and morphological characterization of Scaphoideini Oman, 1943 (61 genera) is substantially revised. Eleven new species are described: Acostemma stilleri sp. nov., Arrugada linnavuorii sp. nov., Drabescus zhangi sp. nov., Parabolopona webbi sp. nov., Goniagnathus emeljanovi sp. nov., Hecalus hamiltoni sp. nov., Scaphoideus omani sp. nov., Dwightla delongi sp. nov., Abimwa knighti sp. nov., Gannia viraktamathi sp. nov., and Doratulina dmitrievi sp. nov. Some family-group level taxonomic changes are made: Platymetopiini Haupt, 1929, Anoterostemmini Haupt, 1929, and Allygidiina Dmitriev, 2006 are synonymized with Athysanini Van Duzee, 1892, syn. nov.; Procepitini Dmitriev, 2002 is synonymized with Cicadulini Van Duzee, 1892, syn. nov.; Listrophorini Boulard, 1971 is synonymized with Chiasmini Distant, 1908, syn. nov.; Adamini Linnavuori & Al-Ne’amy, 1983, Dwightlini McKamey, 2003, and Ianeirini Linnavuori, 1978 are synonymized with Selenocephalini Fieber, 1872 syn.nov., and all three are now recognized as valid subtribes in their parent tribe. New placements of many genera to tribe and subtribe are made, and these are described in individual taxon treatments.
Two species of Tomoceridae were found near Dalat, southern Vietnam. Tomocerus ocreatus Denis, 1948 is redescribed based on a neotype specimen. Previous records of Tomocerus ocreatus in non-type localities are reevaluated. A new species Tomocerina annamitica sp. nov. is described. The new species is mainly characterized by its small body size, pointed tenent hair, compound dental spines and the absence of intermediate teeth on mucro.
A recent paper on the phylogenetic relationships of species within the cephalopod family Mastigoteuthidae meant great progress in stabilizing the classification of the family. The authors, however, left the generic placement of Mastigoteuthis pyrodes unresolved. This problem is corrected here by placing this species in a new monotypic genus, Mastigotragus, based on unique structures of the photophores and the funnel/mantle locking apparatus.
Examination of the type series of Schizotus gibbifrons Lewis 1887 and Pyrochroa higoniae Lewis 1895, together with examination of the historical Lewis collection of Pseudopyrochroa japonica (Heyden 1879), provide hints as to the conspecifi city of these binomials. This evidence, together with concurrent collection events spanning more than 100 years and general anatomy suggests Pseudopyrochroa japonica (Heyden 1879) and Pseudopyrochroa gibbifrons (Lewis 1887) are conspecifi c. Schizotus gibbifrons Lewis is proposed as a new junior synonym of Pyrochroa japonica Heyden, where it joins the established synonym, Pyrochroa higoniae Lewis (1895). Schizotus theresae Pic 1911 is also proposed as a new junior synonym of Pyrochroa basalis Pic 1906, where it joins a long list of synonyms. Complete synonymies are presented for both species.
Sundapyrochroa, a new genus of pyrochroine Pyrochroidae, is described from three Sunda Shelf species most recently assigned to Pseudopyrochroa Pic: Sundapyrochroa atricolor (Pic), comb. nov., recorded from Peninsular Malaysia, Sundapyrochroa nigripennis (Pic), comb. nov., recorded from Peninsular Malaysia, and Sundapyrochroa sumatrensis (Pic), comb. nov., recorded from Borneo (Malaysia: Sabah) and Sumatra (Indonesia). Schizotus rotundicollis Pic, syn. nov., is proposed as a junior synonym of S. sumatrensis. No evidence could be found to support subspecifi c categories for S. nigripennis, thus, Pseudopyrochroa nigripennis notaticeps Pic, syn. nov., is proposed as a junior synonym of S. nigripennis. Gross anatomy of the cranium (female, male), antennae (female, male), and genitalia (male) are both diagnostic and enigmatic, suggesting no clear relationships with other pyrochroine genera.
Pyroghatsiana, a new genus of pyrochroine Pyrochroidae is described from the Southern Ghats of the Indian continental southern tip. The only known specimen is a female, Pyroghatsiana madurensis (Pic), new combination, originally placed in Dendroides Latreille, and subsequently transferred to Pseudodendroides Blair. Several striking differences including the dorsal interocular width between the compound eyes, shape and length of the third antennal segment, and shape of the pronotum preclude placement of Pyroghatsiana in either Dendroides, Pseudodendroides, or any other existing pyrochroine genus.
The endophallic structure of the genus Laius is studied and discussed based on the examination of 19 species from Asia to the Indian Ocean. The structure contains two primary sclerites (named gonoporal piece and ligula), a secondary sclerite on the basal part of the gonoporal piece (named additional sclerite) in some species, and a membranous basal area closely covered with many spines (named spinous area). Five species groups are recognized based on the morphology of the endophallic sclerites. The sympatric species have different body sizes and quite distinguishable endophallic sclerites (= different species group), while the allopatric species have overlapping body sizes and similar endophallic sclerites (= same species group). Three new species are described and six previously known species are redescribed with endophallic sclerites, and the descriptions of endophallic sclerites of the remaining ten species are added. The larva of Laius rodriguesensis sp. nov. is also described. The genus
Nossibeus Evers, 1994 is synonymised with Laius Guérin-Méneville, 1830.
The Asian species of the subfamily Thaumastodinae Champion, 1924 are reviewed. Seven new species are described: Acontosceles borneensis sp. nov., Pseudeucinetus papuanus sp. nov., Mexico ogasawaraensis sp. nov., M. baliensis sp. nov., M. papuanus sp. nov., M. palauensis sp. nov. and M. borneensis sp. nov. The genus Babalimnichus Satô, 1994 is treated as a junior synonym of the genus Mexico Spilman, 1972, and three known species of the genus Babalimnichus are transferred to Mexico, viz. M. taiwanus (Satô, 1994) comb. nov., M. masamii (Satô, 1994) comb. nov. and M. splendens (Hernando & Ribera, 2003) comb. nov. Additional specimen data are shown, and new distributional records are as follows: Acontosceles chujoi Yoshitomi & Satô, 2005 from Vietnam; A. zetteli Pütz, 2008 from Laos; Pseudeucinetus javanicus Yoshitomi & Putra, 2010 from Lombok Island; Mexico taiwanus (Satô, 1994) comb. nov. from Lutao, Lanhsu and the Yonaguni-jima Islands; and M. masamii (Satô, 1994) comb. nov. from Kume-jima. A species list of the subfamily Thaumastodinae is given, with ZooBank LSIDs. The phylogenetic relationships of the thaumastodine genera are discussed.
Four new species of the genus Caccothryptus (Coleoptera: Limnichidae) are described: C. taiwanus from Taiwan; C. orion from Okinawa; C. tibetanus and C. chayuensis from Tibet. All the species belong to the testudo species group (sensu Hernando & Ribera 2014). Additional specimen data and an updated species list are also given, and C. testudo Champion, 1923 is newly recorded from Thailand.
The 'D. bakoue species complex' Rafael, 1984 ('D. montium species group' Da Lage et al., 2007) comprises seven Afrotropical species. Using complete mitochondrial genome sequences and detailed morphological analysis, we revised thephylogenetic relationships between these species including two new ones. We found the 'D. bakoue species complex' to be a junior synonym of the 'D. seguyi species complex' Lachaise, 1971 and its seven species polyphyletic. We thus classified the species into three complexes, the 'D. seguyi species complex' comprises D. seguyi Smart, 1945, D. malagassya Tsacas & Rafael, 1982, D. curta Chassagnard & Tsacas, 1997 and D. chocolata sp. nov., the new 'D. tsacasi species complex' comprises D. tsacasi Bock & Wheeler, 1972 and D. seguyiana Chassagnard & Tsacas, 1997, and the new 'D. vulcana species complex' comprises D. vulcana Graber, 1957 and D. mylenae sp. nov. Drosophila bakoue Tsacas & Lachaise, 1974 could not be assigned to any of the defined complexes. The two new species are endemic to Madagascar and we report the presence of D. seguyi and D. curta in Mayotte and Madagascar, respectively. The results hence represent a significant step towards understanding the diversity and evolution of this species group in Africa and the islands of the Western Indian Ocean.
Cloeon perkinsi was described from South Africa in 1932 by Barnard. Despite being relatively common in Africa, it was mentioned in the literature quite rarely, and its known distribution to date includes most of sub-Saharan Africa. Material collected recently in Ethiopia, Israel, Saudi Arabia, and Yemen extends its distribution in East Africa, Arabian Peninsula and the Levant. We examined this material, and provide a re-description of adults (females and males) and nymphs of the species. It represents a much-needed urge mainly due to inconsistencies in literature reports regarding colouration, and sometimes incomplete morphological description of all stages. We demonstrate the intraspecific morphological variability that we have witnessed, and provide information regarding the range of habitats colonised by C. perkinsi. Based on geological and climatic history of the studied region, taken together with among countries genetic distances of the mitochondrial barcoding gene COI, we propose colonisation mechanisms for the north-easternmost limit of distribution. The fragmented distribution pattern of the species highlights the conservation importance of isolated aquatic habitats in the region, as well as current knowledge gaps.
Nine individuals of Apatidelia from Zhejiang Province, China were examined and their barcode sequences were generated and analyzed. A new species, A. morsei Xu & Sun sp. nov., is described and illustrated. The larva, male and female of A. acuminata Leng & Yang, 1998 and the male and female of A. morsei Xu & Sun sp. nov. are associated by mtCOI gene sequences. The male of A. acuminata Leng & Yang, 1998 is re-described and re-illustrated, and the female and the larva of the same species are also described and illustrated. Females and larvae of the genus are here reported for the first time.
Three species of the genus Cryptochetum Rondani, 1875 from China are described and figured as new to science: C. euthyiproboscise sp. nov., C. glochidiatusum sp. nov., and C. longilingum sp. nov. An identification key to the known species of Cryptochetum from China is presented. The type specimens of the new species are deposited in the Henan Agricultural University.
Important missing specimen data are provided for the original description of Ozophora woodruffi Slater (2005: 245) (Hemiptera: Lygaeidae), along with additional comparative relationships. Because of the missing type information, according to ICZN rules (1999), the species became a nomen nudum. This paper now serves to validate the name, and authorship becomes Slater (2012).
In the eastern United States, the genus Cotinis Burmeister previously contained only C. nitida (L.), the common economic pest known as the "Green June Beetle". A new species from the Florida Keys, Cotinis aliena, is here described and illustrated. A checklist is provided for the genus, which includes 27 valid New World species, and 44 synonyms.
The first New World amber member of the family Lucanidae is described from the Dominican Republic. Its age is presumed to be Miocene (20-30 million YBP). It is also the fourth known amber species, the second Miocene fossil species, the second fossil species in the subfamily Syndesinae, and the first species (fossil or extant) of Lucanidae from the entire Caribbean. It is especially interesting because it is a member of the Australasian genus Syndesus MacLeay. Other such disjunct Dominican amber insect fossils are also discussed.
Evidence is presented that the subspecies Chrysobothris thoracica guadeloupensis Descarpentries, 1981
(Coleoptera: Buprestidae) should be recognized at the species level. Character evidence is provided to separate C.
guadeloupensis, new status, from C. thoracica Fabricius, 1798. Both species are illustrated with habitus photographs
and images of the male genitalia.
Este trabajo actualiza el conocimiento sobre la presencia de cochinillas protegidas en el olivo, Olea europaea Linnaeus (Oleaceae), en Brasil. Aonidiella aurantii (Maskell), Hemiberlesia cyanophylli (Signoret), Hemiberlesia lataniae (Signoret) y Pinnaspis strachani (Cooley) son nuevos registros en olivos en Brasil. Melanaspis obscura (Comstock) es la primera cita en O. europaea en el mundo y una nueva sinonimia se propone: Melanaspis jaboticabae (Hempel) = Melanaspis obscura (Comstock).
Revised definitions are given for the genus Mysidium Dana, 1852, and its eight previously known species, based on material from Curaçao, Bonaire and SE-Brazil, along with the evaluation of published data. Type material of Diamysis columbiae Zimmer, 1915, M. cubanense Băcescu & Ortiz, 1984 and M. rubroculatum Băcescu & Ortiz, 1984 is examined. A lectotype is designated for D. columbiae Zimmer, 1915, a senior synonym of Mysidium columbiae (Zimmer, 1915). Two new species are described, M. triangulare Wittmann sp. nov. from Curaçao and M. antillarum Wittmann sp. nov. from Curaçao and Bonaire. Known ranges are extended by first records of M. cubanense from Curaçao and Bonaire and of M. integrum W.M. Tattersall, 1951 from SE Brazil. Three morphologically different groups are established at the subgeneric level: (1) the nominotypical subgenus Mysidium Dana, 1852 with M. gracile (Dana, 1852), M. integrum, M. cubanense, M. rubroculatum and M. triangulare sp. nov. from the West Atlantic plus M. rickettsi Harrison & Bowman, 1987 from the East Pacific; (2) Occimysidium Wittmann subgen. nov. represented only by M. pumae Ortiz, Hendrickx & Winfield, 2017 from the Pacific coast of Mexico; and finally (3) Orientomysidium Wittmann subgen. nov. comprising M. columbiae and M. antillarum sp. nov. from the West Atlantic. The poorly known M. iliffei Băcescu, 1991 is not assigned to any subgenus. A key to the resulting three subgenera and ten nominal species of the genus Mysidium is given.
Three species of Lophogastrida and eight Mysida are documented for samples from 5161–5497 m bottom depth in the Angola Basin. Previously known latitudinal ranges are extended southward for five species, and bathymetric ranges extended beyond 5000 m for six species. Upon revision of the subfamily Petalophthalminae (Mysidae), four species previously attributed to the genus Petalophthalmus are integrated into Ipirophthalmus gen. nov. as I. liui gen. et comb. nov., I. caribbeanus gen. et comb. nov., I. oculatus gen. et comb. nov., and I. macrops gen. et comb. nov., mainly based on the structure of eyes and presence of setae on the telson. Petalophthalmus cristatus sp. nov. is described based on its reduced cornea and the structure of eyestalks, rostrum, mandibles, and telson. The structure of mouthparts, foregut and maxillipeds suggests an omnivorous mode of life. The diagnosis of the tribe Calyptommini (Mysidae: Erythropinae) is widened to cover the 3-segmented, uniramous fourth male pleopod and the non-incised eyeplate with horn-like rudiments of eyestalks in Abyssomysis cornuta gen. et sp. nov. The structure of mandibles, foregut, and second maxilliped suggest detritus feeding in this species. Keys to the Calyptommini and Petalophthalminae are given.
Dendritic and/or rosetted microborings in calcareous and osteic skeletal substrates have a diverse trace fossil record, spanning most of the Phanerozoic, whereas the ichnodiversity of comparable bioerosion traces produced in modern seas is rather limited. The most prominent occurrences are known from Devonian brachiopods and from Upper Cretaceous belemnite rostra. Ichnotaxonomically, they are comprised within one of the few ichnofamilies established to date, the Dendrinidae Bromley et al., 2007. As an outcome of the present revision of this ichnofamily, the plethora of 84 ichnospecies established within 25 ichnogenera since the erection of the type ichnogenus Dendrina Quenstedt, 1849 was considerably condensed to 22 ichnospecies included in 7 ichnogenera, based on a coherent morphological categorisation and ichnotaxobasis assessment. The suite of ichnogenera now subsumed within the Dendrinidae includes Dendrina Quenstedt, 1849; Clionolithes Clarke, 1908; Calcideletrix Mägdefrau, 1937; Dictyoporus Mägdefrau, 1937; Abeliella Mägdefrau, 1937; Nododendrina Vogel et al., 1987; and Pyrodendrina Tapanila, 2008. New combinations thereby concern Dendrina dendrina (Morris, 1851) comb. nov., Clionolithes pannosus (Solle, 1938) comb. nov., C. alcicornis (Vogel et al., 1987) comb. nov., C. convexus (Hofmann, 1996) comb. nov., Calcideletrix anomala (Mägdefrau, 1937) comb. nov., C. fastigata (Radtke, 1991) comb. nov., Dictyoporus balani (Tavernier et al., 1992) comb. nov., Nododendrina europaea (Fischer, 1875) comb. nov., N. incomposita (Mägdefrau, 1937) comb. nov. and N. paleodendrica (Elias, 1957) comb. nov. Investigation of new material and a reassessment of 63 dendrinid microborings previously addressed in informal nomenclature allowed the establishment of two complementing ichnogenera, Rhopalondendrina igen. nov. and Antodendrina igen. nov., and eight new ichnospecies, comprising Pyrodendrina arctica isp. nov., P. belua isp. nov., P. villosa isp. nov., Rhopalondendrina avis igen. et isp. nov., R. acanthina igen. et isp. nov., R. contra igen. et isp. nov., R. tigris igen. et isp. nov. and Antodendrina ligula igen. et isp. nov. In densely bioeroded calcareous substrates, different dendrinids and other bioerosion traces may be found in direct contact with each other, forming composite trace fossils, but some of these associations appear rather systematic in nature and could be the work of the same tracemaker under different behavioural modes, thus forming compound trace fossils. In these cases, however, the distinction between the two concepts remains largely equivocal. Dendrinid microborings are primarily found in living and dead calcareous skeletal substrates of bivalves, brachiopods, belemnites and corals, with complementing records from six other substrate types. Facing considerable sampling artefacts, evidence for true substrate specificity or symbiotic relationships is inconclusive as yet, whereas there is direct evidence for post-mortem infestation in several cases, such as the diverse dendrinid associations in Upper Cretaceous belemnite guards. Despite a wealth of available interpretations, the actual biological identity of the dendrinids’ tracemakers remains largely speculative. The most convincing evidence has been put forward in support of foraminiferans as the producers of Nododendrina, and excavating micro-sponges producing Clionolithes and some Calcideletrix. Since most of the dendrinids are found in aphotic (palaeo-)environments, these two principal types of organotrophic tracemakers are also potential candidates for the other ichnogenera. With regards to evolutionary patterns through geologic time, strong adaptive radiations are evident from the ichnodiversity of dendrinid ichnospecies in the Early to Mid-Palaeozoic, reflecting the “Ordovician Bioerosion Revolution” (sensu Wilson & Palmer 2006) and the “Mid-Palaeozoic Precursor of the Mesozoic Marine Revolution” (sensu Signor & Brett 1984), respectively, and in the Mesozoic, coinciding with the prominent “Marine Mesozoic Revolution” (sensu Vermeij 1977). This pattern mimics that of
other micro- and macro-bioerosion trace fossils and is interpreted as a reflection of increased predation pressure and consequent infaunalisation. For extinction events, in turn, a differential effect is recorded in that the first four of the “Big Five” mass extinctions appear not to have had any noticeable effect on dendrinid ichnodiversity, whereas the end-Cretaceous mass-extinction resulted in a 77% drop following the Cretaceous peak ichnodiversity of 13 dendrinid ichnospecies.
Arising from a number of 2019 IUCN Red List assessments for a subset of Chinese Odonata, a series of corrections and taxonomic revisions were made to the World Odonata List. The rationale for these amendments is provided here. Paragomphus wuzhishanensis Liu, 1988 is shown to be a junior synonym of Paragomphus pardalinus Needham (1942). Epophthalmia kuani Jiang 1998 is synonymised as a junior synonym of Epophthalmia. elegans (Brauer, 1865) and Epophthalmia bannaensis Zha & Jiang, 2010 is treated as a junior synonym of Epophthalmia vittata Burmeister, 1839. Idionyx pseudovictor Xu, 2013 is shown to be junior synonym of Idionyx claudia Ris, 1912 and Sympetrum anomalum Needham, 1930 is treated as a junior synonym of Sympetrum maculatum Oguma, 1922.
We describe a new butterfl y species, Anaeomorpha mirifi ca Simon and Willmott, n. sp. (Lepidoptera, Nymphalidae, Charaxinae), from premontane rain forest of the Chocó region of northwestern Ecuador. This represents the second known species and the fi rst record outside of the Amazon basin for this taxonomically and biologically enigmatic genus. Although the two species are not known to be sympatric, we identifi ed 21 characters in the external color pattern, wing shape and male genitalia that together support distinct species status. Most notably, the new species possesses an ocellus in the ventral hind wing tornus, a character which occurs in the Neotropical Charaxinae only in the genus Prepona Boisduval, 1836. A mean divergence of 6.8% in the COI ‘barcodes’ between the two species underlines their taxonomic distinctness.
Species are often presumed to be apparent in nature, but in practice they may be difficult to recognise, especially when viewed across continents rather than within a single site. Coalescent-based Poisson-tree-process (PTP) models applied to fast-evolving genes promise one quantitative criterion for recognising species, complete with the estimates of uncertainty that are required of a scientific method. Such methods face challenges especially in discerning between widespread polytypic species and complexes of closely related, restricted-range species. In particular, ‘over-sampling’ of many closely related individuals within one species could risk causing groups of less closely-related individuals within other species appearing relatively more distinct and consequently could risk them being interpreted falsely as separate species. Some of the most persistent taxonomic problems among bumblebees (genus Bombus Latreille, 1802) are within the subgenus Melanobombus von Dalla Torre, 1880. For a global revision of Melanobombus species, we use COI barcodes and seek to reduce the risk from localised over-sampling by filtering the data to include only unique haplotypes. Unique haplotypes give more conservative results than unfiltered data, but still increase the number of species in comparison with recent morphological treatments. After integrative assessment of COI coalescents in comparison with morphological groups, the number of accepted species shows a non-linear increase with sample size that plateaus to an increase of 47% (to 25 species) compared with a previous estimate (of 17) based on morphology alone. For the most widespread and variable species-complexes, our revised species improve the match to the patterns expected of species, both for genetic divergence-with-distance and for sympatry, leading to three main inferences. (1) The particularly widespread polytypic Bombus sichelii Radoszkowski, 1859, is a single species. (2) We detect two candidates for species within previous broad concepts of each of the former B. lapidarius (Linnaeus, 1758), B. miniatus Bingham, 1897, and B. rufofasciatus Smith, 1852. Within B. lapidarius s. lat. we find insufficient evidence to corroborate the candidate species, with no coalescent or morphological support for a recent claim for a separate species, B. bisiculus Lecocq, Biella, Martinet & Rasmont, 2019 described from southern Italy, but rather we find a weak and uncorroborated coalescent for a different and much broader group of samples from across southeastern Europe but excluding Turkey. Within the former broad concepts of B. miniatus s. lat. and B. rufofasciatus s. lat. the coalescent evidence is stronger and subtle evidence from morphology corroborates recognising B. miniatus s. str. and B. eurythorax Wang, 1892 stat. rev. as separate species as well as B. rufofasciatus s. str. and B. prshewalskyi Morawitz, 1880 stat. rev. as separate species. (3) Our coalescent and morphological results ‘split’ more clearly what has long been interpreted as a single polytypic B. keriensis Morawitz, 1887, s. lat., by supporting novel concepts of the restricted-range species: B. alagesianus Reinig, 1930 stat. rev., B. incertoides Vogt, 1911 stat. rev., B. keriensis s. str., B. qilianensis sp. nov., B. separandus Vogt, 1909 stat. rev., and B. tibeticus sp. nov. A lectotype is designated for the name B. keriensis and a neotype is designated for the name B. alagesianus. We estimate the phylogeny of Melanobombus species by including three slower-evolving genes to provide more evidence for deeper relationships, to estimate the time calibration of this phylogeny, and to estimate ancestral distributions, all within a Bayesian framework. We provide the first keys for identifying all of the species of Melanobombus.
Females of Traumatomutilla André (Hymenoptera: Mutillidae) are organized into 14 species groups; 136 of the 138 Traumatomutilla taxa known from females were studied. Of these species groups, only the inermis species group was adequately established and diagnosed in the literature (Casal 1969). The remaining newly established species groups are as follows: americana, auriculata, bellica, bifurca, diabolica, gemella, indica, integella, juvenilis, quadrinotata, tabapua, trochanterata, and vitelligera. Diagnoses and lists of included species are provided for each species group. Eighty-three new country records are provided for 49 species.
Two new genera and species of tiger beetles from Baltic amber (Coleoptera: Carabidae: Cicindelinae)
(2017)
Two fossil tiger beetle species (Coleoptera, Carabidae, Cicindelinae) are described from Eocene Baltic amber using light microscopic and X-ray microscopic techniques. Both species are considered representatives of the subtribe Iresina Rivalier, 1971 due to the shared combination of character states: glabrous head, six labral and four suborbital setae, and glabrous pronotum. Palaeopronyssiformia groehni Wiesner, Will, and Schmidt, new genus, new species, is characterized by a glabrous and furrowed head with six labral setae, large eyes, presence of two supraorbital setae on each side, mandibles with two teeth of the incisor region, and a glabrous and furrowed pronotum. Palaeoiresina cassolai Wiesner, Will, and Schmidt, new genus, new species, is characterized by a unicolored, undentated labrum, mandibles with two teeth of the incisor region, glabrous head with six labral setae, two clypeal setae, two supraorbital setae on each side, and a glabrous pronotum, mesepisternum, mesepimeron, and metepisternum. The species described here represent the only known tiger beetle fossils preserved in Baltic amber.
Distributional records of the 177 tiger beetle species and subspecies known for Vietnam are given together with a key and habitus photos (Coleoptera: Carabidae: Cicindelinae). Tricondyla (Tricondyla) macrodera abruptesculpta Horn, 1925, Protocollyris festiva Naviaux, 2008, Neocollyris (Brachycollyris) purpureomaculata borea Naviaux, 1994, Neocollyris (Brachycollyris) torosa Naviaux, 2010, Neocollyris (Leptocollyris) rogeri Shook and Wu, 2006, Neocollyris (Leptocollyris) laosensis Naviaux, 1999, Neocollyris (Stenocollyris) rubens (Bates, 1878), Neocollyris (Pachycollyris) tricolor Naviaux, 1991, Neocollyris (Pachycollyris) panfi lovi Naviaux and Matalin, 2002, Therates rugulosus Horn, 1900, Therates bannapecolus Sawada and Wiesner, 1999, Calochroa interruptofasciata fl avolineata (Chaudoir, 1865), Cylindera (Ifasina) cyclobregma Acciavatti and Pearson, 1989, Cylindera (Ifasina) decolorata (Horn, 1907), Cylindera (Ifasina) humillima (Gestro, 1893) and Cylindera (Eugrapha) elisae elisae (Motschulsky, 1859) are reported for the fi rst time in the fauna of Vietnam. Neocollyris (Isocollyris) septentrionalis Naviaux, 1999, Neocollyris (Leptocollyris) linearis tenuicornis (Chaudoir, 1864), Neocollyris (Pachycollyris) aptera apicalis (Chaudoir, 1864), Neocollyris (Pachycollyris) contracta (Horn, 1905), Therates dimidiatus wallacei Thomson, 1857, Cylindera (Cylindera) pseudocylindriformis Horn, 1913, Cylindera (Ifasina) viridilabris (Chaudoir, 1852), Cylindera (Ifasina) psilica psilica (Bates, 1866), Cylindera (Eugrapha) venosa (Kollar, 1836) and Myriochila (Myriochila) undulata (Dejean, 1825) are excluded from the fauna of Vietnam
Distributional records of the 71 tiger beetle species and subspecies known for Argentina are given together with a key and habitus photos (Coleoptera: Cicindelidae). Brasiella (Brasiella) cuyabaensis Mandl, 1970 and Brasiella (Gaymara) rotundatodilatata (Horn, 1925) are reported as new for the fauna of Argentina. Brasiella (Brasiella) stamatovi (Sumlin, 1979) is transferred to Cylindera (Plectographa), becoming C. (P.) stamatovi (Sumlin), new combination. Cylindera (Plectographa) siccalacicola (Sumlin, 1979) is placed into synonymy under Cylindera (Plectographa) hassenteufeli (Mandl, 1960), new synonymy. A lectotype is designated for Cylindera (Plectographa) patagonica bergiana (Horn, 1895).
A key is provided for the twelve taxonomic groups within the genus Therates Latreille (Coleoptera: Cicindelidae). The chennelli group is reviewed here and a key to the 53 species known for this group is provided. Each species is illustrated, and a detailed description of its morphology and distributional records are provided. Therates khaoyaii, Th. pearsoni, Th. safraneki, Th. schuelei and Th. sigridgeissleri are described as new to science. Therates differens Sawada and Wiesner, 1999 is placed into synonymy under Th. concinnus Gestro, 1888. Therates tonkinensis kubani Wiesner, 1988 is placed into synonymy under Th. tonkinensis Horn, 1902 and Th. belokobylskiyi Matalin and Wiesner, 2006 is placed into synonymy under Th. haucki Moravec and Wiesner, 2001. Lectotypes are designated for Th. waagenorum Horn, 1900, Th. clavicornis Horn, 1902, Th. tonkinensis Horn, 1902 and Th. annandalei Horn, 1908. Therates baolocensis Wiesner, 1996 is elevated to species rank.
Three new fossil species from Burmese amber are described, one clearly in family Calamoceratidae, the other two in the highly variable family Odontoceridae. The family Odontoceridae contains 18 disparate genera, but there are no good diagnostic characters, which makes it difficult to place fossil taxa in this family. We here offer a revised diagnosis for the family, highlighting the lack of good diagnostic characters, and the need to use sets of characters to place (extinct) taxa in this family. On this taxonomic basis we describe two new species in the hitherto monotypic fossil genus Palaeopsilotreta Wichard & Wang, 2017 (Odontoceridae), Palaeopsilotreta burmanica sp. nov. and P. cretacea sp. nov., redescribe the type species based on additional information, and describe features of the female, based on two specimens, one of which is embedded adjacent to a male identified as P. xiai. Males of Palaeopsilotreta bear bipectinate antennae; the antennae of the females are simple and filiform. Similarly, bipectinate antennae are present in the third species we describe, Bipectinata orientalis gen. et sp. nov., which otherwise lacks the character combinations associated with the Odontoceridae, but clearly can be assigned to the family Calamoceratidae.
Examination of leucothoid amphipods of the Red Sea has revealed seven species not previously reported from this location. Leucothoe minoculis sp. nov., Leucothoe pansa sp. nov., Leucothoe reimeri sp. nov., and Paranamixis sommelieri sp. nov. are described and the range of Leucothoe predenticulata Ledoyer, 1978, L. acutilobata Ledoyer, 1978 and L. squalidens Ledoyer, 1978 is expanded to include the Red Sea. Clarification of reports of L. acanthopus Schellenberg, 1928 and L. bannwarthi (Schellenberg, 1928) is provided and Leucothoe alani sp. nov. is described from outside the Red Sea.
A new species, Chrysobothris cerceripraeda Westcott and Thomas (Coleoptera: Buprestidae), is described from prey specimens found in nests of the ground-nesting wasp, Cerceris fumipennis Say (Hymenoptera: Crabronidae), near Jacksonville, Florida, USA. A listing for all species of Buprestidae taken as prey of the wasp in that state is provided, four of which, including the species described herein, are new. Notes are also given for
four new state records for Buprestidae.
The concept of the jumping spider genus Pochytoides Berland & Millot, 1941 is reviewed, based on the examination of described and undescribed species. Pochytoides is elevated from the subgeneric to the generic rank and a short diagnosis and description of the genus are presented. Redescriptions or descriptions of all species are provided together with a key to the species. Two new combinations are proposed: Pochytoides perezi (Berland & Millot, 1941) comb. nov. and P. poissoni (Berland & Millot, 1941) comb. nov. (both from Pochyta). Pochyta remyi Berland & Millot, 1941 originally placed in the subgenus Pochytoides is excluded; new combination Thiratoscirtus remyi (Berland & Millot, 1941) comb. nov. is proposed for it (but its generic status is uncertain). Six new species are described: Pochytoides monticola sp. nov., P. obstipa sp. nov., P. lamottei sp. nov., P. patellaris sp. nov., P. securis sp. nov. and P. spiniger sp. nov. The genus has a West African distribution.
Dichromatobolus, a new genus of spirobolidan millipedes from Madagascar (Spirobolida, Pachybolidae)
(2020)
A new genus, Dichromatobolus gen. nov., belonging to the genus-rich mainly southern hemisphere family Pachybolidae of the order Spirobolida, is described based on D. elephantulus gen. et sp. nov., illustrated with color pictures, line drawings, and scanning electron micrographs. The species is recorded from the spiny bush of southwestern Madagascar. Dichromatobolus elephantulus gen. et sp. nov. shows an unusual color pattern, sexual dichromatism with males being red with black legs and females being grey. Males seem to be more surface active, as mainly males were collected with pitfall traps. Females mainly come from the pet trade. The body of this species is short and very wide, being only 8 times longer than wide in the males. Live observations show the species is a very slow mover, digging in loose soil almost as fast as walking on the surface. The posterior gonopods of Dichromatobolus gen. nov. are unusually simple and well-rounded, displaying some similarities to the genera Corallobolus Wesener, 2009 and Granitobolus Wesener, 2009, from which the new genus differs in numerous other characters, e.g., size, anterior gonopods and habitus. Despite several attempts with fresh tissue samples and different primers, molecular barcoding did not work for Dichromatobolus gen. nov. Any relationships to the other 15 genera of Pachybolidae indigenous to Madagascar remain unknown.
A taxonomic revision of Desplatsia Bocq. (Malvaceae s. lat. Juss., subfamily Grewioideae Hochr., tribe Grewieae Endl.) based on about 800 herbarium specimens is presented. Desplatsia is a genus of trees and shrubs found in tropical West and Central Africa and is characterized by subulately divided stipules, the absence of an androgynophore, stamens that are fused to a tube at the base, and large and distinctive fruits that are dispersed by elephants. Four species are recognized (D. subericarpa Bocq., D. chrysochlamys (Mildbr. & Burret) Mildbr. & Burret, D. dewevrei (De Wild. & T.Durand) Burret and D. mildbraedii Burret) and 12 species names are placed into synonymy, two of which have been put into synonymy for the first time: D. floribunda Burret syn. nov. and D. trillesiana (Pierre ex De Wild.) Pierre ex A.Chev. syn. nov. All four species are widely distributed and their conservation status is assessed as Least Concern (LC). A key to the species, full species descriptions, illustrations, a specimen citation list and distribution maps are provided.
Acoma howdenorum, Acoma westcotti, Acoma quadrilaminata, and Acoma cimarron (Coleoptera: Scarabaeidae: Melolonthinae), all new species, are described from Yuma County, Arizona, USA, and Baja California Sur, Baja California (Norte), and Sonora, Mexico, respectively. Habitus of the four new species is illustrated, and an updated key to the described species in the genus is provided. Distribution and variation of Acoma glabrata Cazier are also discussed.
The Orizabus Fairmaire (Coleoptera: Scarabaeidae: Dynastinae: Pentodini) of the USA are reviewed. Orizabus pinalicus new species and O. mcclevei new species are described. Lectotypes are here designated for eight species names: Bothynus pyriformis LeConte, Pseudaphonus lucidus Casey, Orizabus snowii Horn, Orizabus cultripes Fairmaire, Orizabus isodonoides Fairmaire, Orizabus sallei Fairmaire, Orizabus fontinalis Casey, and Orizabus ponderosus Casey. Illustrations of diagnostic characters and a key to the five included species are presented. The Mexican species O. isodonoides and O. rubricollis Prell are also illustrated for comparison to the new species.
Plesioclytus morrisi Wappes and Skelley new species (Coleoptera: Cerambycidae) from the Ohoopee dune system in central Georgia is described with comments on the biology of the new species. The taxonomic placement of Plesioclytus Giesbert in the tribe Clytini is questioned as key characters are found to differ from the current characters used to define the tribe in the New World, resulting in its transfer to the newly erected Plesioclytini Wappes and Skelley new tribe, defined herein. Habitat photos for the new species and habitus photos for it and P. relictus Giesbert are provided.
A new Mexican species of Ochraethes Chevrolat, 1860 (Coleoptera, Cerambycidae, Cerambycinae, Clytini) is described: Ochraethes skillmani Wappes, Santos-Silva and Botero. Plocaederus mirim Martins and Monné, 2002 (Cerambycini) is redescribed and its female is figured for the first time. New geographical records in Plocaederus Dejean, 1835 are also provided.
The Bolivian species of Polyrhaphis Audinet-Serville, 1835, (Coleoptera, Cerambycidae, Lamiinae) are reviewed and illustrated, with P. skillmani new species described. A key is presented to the six species recorded from Bolivia (P. angustata Buquet, 1853; P. argentina Lane, 1978; P. gracilis Bates, 1862; P. pilosa Lane, 1965; P. spinosa (Drury, 1773); and P. skillmani Wappes and Santos-Silva, new species). Their collection localities, based on recently identified specimens examined by the authors, are plotted to show the distribution of species, and displayed next to an ecoregion map of Bolivia to illustrate biogeographical information for Polyrhaphis.
Nine new species of Rhinotragini (Coleoptera: Cerambycidae: Cerambycinae) are described from the Neotropical region: Odontocera galileoae (Mexico); Odontocera mthomasi (Guatemala); Ecliptoides vandenberghei (Nicaragua); Eclipta ricei and Odontocera skelleyi (Costa Rica); Eclipta nearnsi (Panama); Odontocera stangei (Venezuela); Chariergodes lingafelteri and Ischasia martinsi (Costa Rica and Panama). Ischasia rufi na Thomson, 1864 is formally excluded from the fauna of Costa Rica and Panama, and Nicaragua is added as a new country record for Eclipta fi cta Bezark, Martins and Santos-Silva, 2013. Notes on Ecliptoides monostigma (Bates, 1869)
are provided in the “Diagnosis” of E. vandenberghei.
A new genus and two new species of Argentine Rhinotragini
(Coleoptera, Cerambycidae, Cerambycinae)
(2017)
Rhopalessa irwini sp.nov. and Rhinion parkeri gen. nov., sp. nov. (Coleoptera, Cerambycidae, Cerambycinae, Rhinotragini) are described from Argentina. A key to species of Rhopalessa is provided.
Two Neoibidionini from Bolivia are described as new: Neocompsa schneppi sp. nov. and Tropidion nancyae sp. nov. The new species are compared to closely related species and previous keys by Martins and Galileo (2007, 2009) are modifi ed to include them. Illustrations of dorsal, ventral and lateral habitus, as well as head structures are included for both species. Comments are also included on color and pattern variation of each species.
The female of Nothopleurus subsulcatus (Dalman, 1823) (Coleoptera: Cerambycidae: Prioninae: Macrotomini) is described for the first time, and the female of Strongylaspis bullata Bates, 1872 is redescribed. Color photographs of the habitus of both, and key characters for the former are included. New distributional records within Mexico for N. subsulcatus and Strongylaspis championi Bates, 1884 are given.
Adesmus martinsi (Coleoptera, Cerambycidae, Lamiinae, Hemilophini), a new species from Bolivia, is described, illustrated, and included in a previous key. The new species displays gender dimorphism in the pubescent pattern and in anatomical structure. Thus, Adesmus becomes the second genus recorded in the Hemilophini to have visual chromatic dimorphism.
An additional 137 species and two tribes are added to the cerambycid fauna of Bolivia while 12 species are deleted. This brings the total number of species known from Bolivia to 1,561. Comments and statistics regarding the growth of knowledge on the Bolivian Cerambycid fauna and species endemicity are included.
Seventy-nine Cerambycidae and two Vesperidae species not previously recorded from Bolivia are listed along with the department where they were collected, and are thus added to the known fauna. An additional 22 species from existing publications, but whose Bolivian distribution is not recorded in the 2013 version of Bezark and Monné (2013), are listed separately to assist inclusion in this important reference. These records, along with the 60 new species described (through February, 2013) since Wappes et al. (2011), brings the total number of Cerambycidae and, closely related families Disteniidae, Oxypeltidae and Vesperidae, to 1,717 species known from Bolivia. New departmental records for another 254 species are listed. Color illustrations for 80 of the 81 species newly recorded from Bolivia are provided. Among the new records for Bolivia is Lathroeus oreoderoides Thomson, 1864 previously known only from South America without exact locality, hence this is its first recorded distribution. A male of Myzomorphus Dejean, 1835 collected at the same time and locality as a female Myzomorphus amabilis (Tippmann, 1960) is likely the previously unknown male of the species. Both sexes are illustrated.
The previously published list of Coleoptera holotypes in the Museo de Historia Natural, Noel Kempff Mercado (MNKM), Santa Cruz, Bolivia is updated to include those deposited in the Museo since then though the end of 2015. Literature citations for the original descriptions of each listed holotype are also provided along with summary comments regarding family composition and authorship of included species. Photographs of collector, authors, habitats, and major Bolivian type repositories are provided.
One individual referable to Calliopiidae G.O. Sars, 1893 was collected from a chemically reduced habitat, the hydrothermal vent systems in Okinawa Trough, and was identified as a new genus and species belonging to this family after a morphological examination. A formal description of this new species and a discussion of the relationship of the new genus within Calliopiidae are presented.
The genus Paharia Distant, 1905 is reviewed based on the description of a new species, Paharia oorschoti sp. nov., and redescription of the allied Paharia putoni (Distant, 1892), both from Turkey. The relationships among Paharia, Subpsaltria Chen, 1943 and Tibicina Kolenati, 1857 of the tribe Tibicinini Distant, 1905 are discussed. The morphology of the exuviae of Pa. oorschoti sp. nov. and S. yangi Chen 1943 is described and compared. Tibicina insidiosa Boulard, 1977 is transferred to Paharia to become Paharia insidiosa comb. nov. A key to all species of Paharia is provided.
All extant species of the planthopper genus Limois Stål (Hemiptera: Fulgoromorpha: Fulgoridae) were studied. One new species, Limois sordida sp. nov., is described and illustrated from China. Six known species are re-described and photos and illustrations of male genitalia are provided. A key to all extant species of this genus is also given.
Taxonomic updates and descriptions of four new species from Yunnan, China are provided: three new species in the genus Pitambara Distant, 1906: P. triremiprocta Wang & Soulier-Perkins, sp. nov., P. impudica Wang & Bourgoin, sp. nov., P. tricorne Wang & Wang, sp. nov., and one new species in the genus Serida Walker, 1857: Serida parenthesisflexuosa Wang & Soulier-Perkins, sp. nov. A new identification key to Pitambara species is provided, as well as to the species of the genus Lacusa Stål, 1862. Lacusa yunnanensis Chou & Huang, 1985 stat. rev. is not considered as a synonym of the species L. fuscofasciata (Stål, 1854) anymore and Lacusa orientalis (Liang, 2000) is transferred to the genus Acothrura Melichar, 1915 as Acothrura orientalis (Liang, 2000) comb. nov.
Five new species of Meta Koch, 1836 (Araneae: Tetragnathidae) from Gaoligong Mountains, China
(2020)
Five new species of the genus Meta C.L. Koch, 1836 from the Gongligong Mountains, Yunnan are described: Meta hamata sp. nov. (♂♀), M. longlingensis sp. nov. (♂), M. tangi sp. nov. (♂♀), M. yani sp. nov. (♂♀) and M. yinae sp. nov. (♂♀). Detailed descriptions of somatic features and genitalic characters, photos of the body and genital organs, line drawings of the copulatory organs and distribution maps are provided.
Neopanorpa van der Weele, 1909 is the second largest genus in Panorpidae, and over 170 Oriental species have been reported hitherto. In this paper, we describe two new species: Neopanorpa luojishana sp. nov. and Neopanorpa xingmini sp. nov. from Southwest China. We also report the first discovery of the male of Neopanorpa ocellaris (Navás, 1908) from Guangxi and Guizhou, China, a species recorded from Sikkim previously. Neopanorpa brevivalvae Chou & Wang, 1988 is synonymized with Neopanorpa lungtaushana Cheng, 1957. In addition, Neopanorpa furcula nom. nov. is proposed for Neopanorpa furcata Zhou, 2005, a junior homonym preoccupied by Neopanorpa furcata (Hardwicke, 1825). The biogeographical implications of some Neopanorpa species are discussed in brief.
On eight species of the spider genus Synagelides Strand, 1906 from China (Araneae: Salticidae)
(2020)
Six new species of Synagelides Strand, 1906 are diagnosed and described: S. bohdanowiczi sp. nov. (♂♀), S. leigongensis sp. nov. (♂♀), S. logunovi sp. nov. (♂♀), S. subgambosus sp. nov. (♂♀), S. wuliangensis sp. nov. (♂♀) and S. xingdouensis sp. nov. (♂♀). The female of S. forkiforma Yang, Zhu & Song, 2007 and the male of S. longus Song & Chai, 1992 are described for the first time. Photos of the habitus and copulatory organs, as well as a distributional map, are provided.
Three species of the genus Prorops Waterson, 1923 occur in Madagascar. Prorops nasuta Waterson, 1923 is recorded for the first time from Madagascar and two new species are described and illustrated: P. sparsa sp. nov. and P. impotens sp. nov., both based on the morphology of males and females. A brief discussion of the status of the genus, illustrations, and a key to Madagascan species of Prorops are provided.
Several taxonomic groups within Empidoidea Latreille, 1809 have been subject to unclear phylogenetic assignments along with multiple parallel hypotheses causing difficulties in classification and morphological identification. This study reviews the internal classification of the Ragadidae and includes a diagnosis and description of all included subfamilies and genera based on the results of an analysis of morphological characters using maximum parsimony. Illustration of important characters and a key to all genera in the family is given. The genus Hormopeza Zetterstedt, 1838 is found to be most closely related to Anthepiscopus Becker, 1891 and Iteaphila Zetterstedt, 1838, and the subfamily Iteaphilinae Wahlberg & Johanson, 2018 is therefore expanded to also include that genus. Hormopeza is consequently excluded from Ragadinae Sinclair, 2016. This study provides diagnoses, descriptions and keys in a contribution to a thorough classification of the empidoid groups and increased ease in morphological recognition.
Populus primaveralepensis A.Vázquez, Muñiz-Castro & Zuno sp. nov., a new species from relict gallery cloud forest in Bosque La Primavera Biosphere Reserve (Mexico), is described and illustrated. The new species belongs to P. subsect. Tomentosae Hart., and is morphologically similar to P. luziarum A.Vázquez, Muñiz-Castro & Padilla-Lepe, but differs from it in having taller trees without root suckers, white and ringed young stems and branches, a branching angle of ca 45º, leaves with higher blade to petiole ratio, leafs frequently elliptic or ovate to widely ovate (vs widely ovate to ovatedeltoid), denser inflorescences, and shorter capsules. The conservation status of the species was assessed as Critically Endangered (CR).
The fossil sirenian material from the upper Oligocene Linz Sands of Upper Austria is reviewed and re-described in detail following a recent approach on the invalidity of the genus Halitherium Kaup, 1838. This morphological study provides the first evidence for the synonymy of “Halitherium” cristolii Fitzinger 1842, “H.” abeli Spillmann, 1959 and “H.” pergense (Toula, 1899), supporting the hypothesis that only a single species inhabited the late Oligocene shores of present-day Upper Austria. In the course of the taxonomic revision of the “Halitherium” species-complex, this taxon is now assigned to the new genus Lentiarenium Voss gen. nov. It represents a more derived sirenian compared to Eocene and early Oligocene taxa distributed across Central Europe and North Africa, which is in accordance with the stratigraphical data. An updated inventory list of all identifiable and referable skeletal material is provided, including a detailed synonymy list for the new taxonomic combination.
In this paper the primary types of Centris bees described by the British entomologist Theodore Dru Alison Cockerell deposited in the Natural History Museum (London) and the Oxford University Museum of Natural History (Oxford) in the United Kingdom, as well as in the United States National Museum (Washington), American Museum of Natural History (New York), the Academy of Natural Sciences of Drexel University (Philadelphia), and in the California Academy of Sciences (San Francisco) in the United States were studied. To stabilize the application of the name C. lepeletieri (= C. haemorrhoidalis (Fabricius)), a lectotype is designated. The study of the primary types allow proposing the revalidation of C. cisnerosi nom. rev. from the synonymy of C. agilis Smith, C. nitida geminata nom. rev. from C. facialis Mocsáry, C. rufulina nom. rev. from C. varia (Erichson), C. semilabrosa nom. rev. from C. terminata Smith and C. triangulifera nom. rev. from C. labrosa Friese. Centris bakeri syn. nov., C. bimaculata carrikeri syn. nov., C. fusciventris matoensis syn. nov., C. heterodonta syn. nov. and C. elegans grenadensis syn. nov. are proposed as a new junior synonyms of C. varia, C. claripennis Friese nom. rev., C. caurensis, C. dentata Smith and C. elegans Smith, respectively. Centris ruae is withdrawn from the synonymy of C. transversa Pérez and proposed as a new junior synonym of C. nitida Smith. In addition, a lectotype for C. buchholzi Herbst (= C. wilmattae) is designated. Information on the repository of the lectotype of C. lepeletieri and images of most primary types studied here are also provided.
In this paper, the species of Centris of the “hyptidis group” are revisited, proposing to recognize them as members of Anisoctenodes subgen. nov., a new subgenus supported by morphological and molecular data. The species included in this new taxon are C. hyptidis Ducke, 1908 (type-species), C. hyptidoides Roig-Alsina, 2000, C. thelyopsis Vivallo & Melo, 2009 and C. anisitsi (Schrottky, 1908), transferring this latter from Centris (Xanthemisia) Moure, 1945. An updated key, information on the type depository, a distribution map, photographs of both sexes as well as of the diagnostic characters of the new subgenus are also provided.
This paper presents the faunistic results of three short field excursions conducted in spring and early summer of 2018 in Kosovo, considering dragonfly fauna one of the most understudied countries of Europe. This study presents first systematic dragonfly research in Kosovo. Within a total of 13 field days between end of April and end of June 2018, 60 sites were surveyed and 44 dragonfly species were found. Significant results include the first documented report of 15 species for Kosovo. New data on several other species with a broader European concern or generally rare on the West Balkan peninsula, i.e. Coenagrion ornatum, Anax ephippiger, Caliaeschna microstigma, Cordulegaster heros, C. bidentata, Somatochlora flavomaculata, and Sympetrum flaveolum, are also presented. The overview of all visited sites is included. Altogether, 47 dragonfly species are now reported for Kosovo.