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A new genus and six new species of the family Gomphodesmidae from the Udzungwa Mts are described, including Pogoro alopias Rosenmejer & Enghoff sp. nov., Pogoro siren Rosenmejer & Enghoff sp. nov., Pogoropsis prolixopes Rosenmejer & Enghoff gen. et sp. nov., Emphysemastix frampt Olsen & Enghoff sp. nov., Agrophogonus hamulus Olsen & Enghoff sp. nov., and Agrophogonus pusillokiellandi Olsen & Enghoff sp. nov. Emphysemastix dracarys Olsen & Enghoff sp. nov. from Iringa city is described. Descriptive notes are given for Pogoro scharffi Hoffman, 2005, and Agrophogonus mwanihana Hoffmann, 2005. General gomphodesmid gonopod morphology is described and illustrated. A key to Udzungwa gomphodesmids is presented, as well as revised keys to all species of Emphysemastix and Agrophogonus. All gomphodesmid species from the Udzungwa Mts are mapped.
A large, newly collected material of the genus Aquattuor from the Udzungwa Mts, Tanzania, has been studied. Two new species are described, viz., A. mollilobus sp. nov. from the Udzungwa Mts and A. nguruensis sp. nov. from the Nguru Mts, Tanzania. Aquattuor claudiahempae Enghoff & Frederiksen, 2015, is recorded as new for the Udzungwa fauna. Characters from the first pair of male legs are illustrated for all Aquattuor species and are shown to be useful for species discrimination. A new gonopodal difference between A. submajor Enghoff, 2015 and A. udzungwensis Enghoff, 2015 is described and illustrated. Intrageneric relationships and distribution patterns are discussed. A distribution map and a key to all Aquattuor species are presented.
Antoinettia, new genus (Coleoptera: Erotylidae: Erotylinae: Tritomini), is erected for three species: A. audbala (Skelley), new combination, A. huhnei Skelley, new species, and A. kovariki (Skelley), new combination. A genus complex involving Ischyrus Lacordaire, 1842, and Megischyrus Crotch, 1873, is defined and a preliminary key to neotropical genera of Tritomini with coarsely facetted eyes is presented.
The genus of Neotropical frogs Pithecopus includes 11 species occurring east of the Andes from southern Venezuela to northern Argentina. Recent genetic approaches pointed out an unusual genetic diversity among populations from localities in north-eastern Brazil recognized as P. nordestinus. In fact, one of these studies confirmed the hypothesis that the São Francisco River acted as an effective geographical barrier during vicariant events in the evolutionary history of P. nordestinus, resulting in two principal, highly divergent clades. Herein we formally describe this divergent clade as a new cryptic species of Pithecopus from north-eastern Brazil, the sister clade of P. nordestinus. It differs from other species of Pithecopus, except for P. azureus and P. nordestinus, by its small body size, lack of the reticulate pattern on flanks, smaller head width, and advertisement calls generally composed of a three-pulsed core.
A new cypridopsine genus, Cyprettadopsis gen. nov., described here, is principally characterized by the reduced caudal ramus, the strongly serrated claw G2 of the antenna (A2), the A2 subquadrate terminal segment, the undivided penultimate segment of the second thoracopod (T2), the morphology of the third thoracopod bearing a distinctly separated terminal segment, the complete septa on the posteroventral margin and the incomplete septa on the anterior margin of both valves. Based on a combination of these characters, a new tribe, Cyprettadopsini trib. nov., is created in the subfamily Cypridopsinae Kaufmann, 1900 to accommodate this new genus, and one new species, Cyprettadopsis sutura gen. et sp. nov., is described as the type species. Apart from the above generic characters, the following features are also typical of the new species: the tiny needlepoint-like pores along the anterior and ventral margins of both valves, the remarkably large β-seta on the mandibular palp and the considerably short d2 seta on the T2. The presence of marginal septa in the new genus is a distinctive character and constitutes the first record of this feature within Cypridopsinae. The taxonomically relevant characters in the new taxon and related taxa are briefly discussed.
A new species of Limnophilinae (Diptera: Limoniidae), Dicranophragma (Brachylimnophila) relictum Mederos sp. nov., is described. This is the third species of this genus recorded from the Iberian Peninsula. This new species was discovered in a cave in the Sant Llorenç del Munt i Serra de l’Obac Natural Park, Barcelona Province (Catalonia) and it is the first record of a species of this genus from a hypogeous environment. The absence of suitable ecological conditions for the survival of this new species, particularly in the vicinity of the cave, suggests that this population of D. relictum Mederos sp. nov. is isolated. Dicranophragma relictum Mederos sp. nov. is characterized by the following features: a general grey-to-greenish-grey coloration on its thorax and a dark-grey abdomen; wings more than four times longer than wide; pale-brown-to-yellowish tinted wings, with brown veins and a well-marked stigma; in the male genitalia the upper part of the aedeagus is acute when viewed laterally and is longer than the lower part. A key is provided for separating the three species of Dicranophragma known to be present in the Iberian Peninsula.
A new genus and species, Richerius marqueti gen. et sp. nov., of a crab of the family Hymenosomatidae MacLeay, 1838 are described from the inland waters of New Caledonia based on several specimens collected in two streams at altitudes of 180 m and 500 m, respectively. Richerius marqueti gen. et sp. nov. was compared to the other freshwater species known in New Caledonia, Odiomaris pilosus (A. Milne-Edwards, 1873), and to species of Amarinus Lucas, 1980, a genus comprising many freshwater species in New Zealand, Australia, Indonesia, the Philippines, and Papua New Guinea, but never recorded in New Caledonia. The barcode fragment of the COI mitochondrial gene was sequenced for seven specimens of R. marqueti gen. et sp. nov., and all sequences were deposited in GenBank. A brief and updated review of the New Caledonian marine and freshwater hymenosmatid fauna is provided.
A new genus, Janzena (Erebidae), and 37 new species of Noctuoidea (Lepidoptera) are described from Florida as well as two species elevated from synonymy to species status. Bleptina biformata, Bleptina extincta, Bleptina flavivena, Bleptina verticalis, Lascoria coma, Janzena pyraliformis, Sigela sordes, Sigela minuta,Sigela incisa, Sigela subincisa, Sigela rosea, Sigela lynx, Dyspyralis ocala, Metalectra nigrior, Metalectra dixoni, Melipotis florida, Doryodes acta, Doryodes unica, Doryodes fulva, Toxonprucha killamae, Zale lafontainei,Zale vargoi, Zale clandestina, Athyrma fakahatchee, Antiblemma perva, Antiblemma carolae, Paectes hercules,Meganola georgei, Litoprosopus linea, Tripudia calusa, Catabenoides insularis, Neogalea caracara, Condica collaris, Homophoberia australis, Diastema leo, Pyreferra slotteni, and Leucania elephas are described as new. Hemeroplanis floccalis (Zeller), revived status, is raised to species status from the synonymy of Hemeroplanis scopulepes (Haworth),and Euscirrhopterus argentata (Druce), revived status, is raised to species status from the synonymy of Euscirrhopterus poeyi Grote. The genus Araeopteron Hampson is restricted to the Old World and Araeopteron vilhelmina (Dyar) is transferred to the genus Sigela Hulst, new combination. The barcode index number (BIN) is provided for each species, when available.
A new genus and species of Sternaspidae (Annelida: Polychaeta) from the deep eastern Atlantic
(2020)
Based on specimens recently collected in sediments from 2700 m depth off Mauritania (Northwest Africa; type locality) and from 2700–4400 m depth off Angola (Southwest Africa), a new genus and species, Mauretanaspis longichaeta gen. et spec. nov., is described. The new genus and species are characterized by a unique combination of characters: ventro-caudal shield covered by firmly adhering sediment, lateral margins strongly bent and merging into integument; introvert hooks tapering; eight pre-shield segments; absence of peg chaetae; exceptionally long posteriormost lateral chaetae equaling body length; posterior shield chaetae equaling shield length. A comparative table of characters for all currently recognised sternaspid genera and a key to all species with ventro-caudal shield covered by firmly adhering sediment are provided.
A new genus of Baetidae is described from Southeast Asia, Procerobaetis gen. nov. It has a wide distribution reaching from Indonesia (Sumatra) to the Philippines. Two new species are described from Indonesia, P. leptobranchius gen. et sp. nov. and P. petersorum gen. et sp. nov., and one new species from the Philippines, P. freitagi gen. et sp. nov. Procerobaetis gen. nov. is characterized by having seven pairs of elongate, apically pointed gills. At least gills I and II are very slender with strongly extended points, which is unique in Baetidae. Similar gills were described from Leptophlebiidae. Procerobaetis gen. nov. is further characterized by having long, slender legs with extended, slender and slightly bent claws. The antennae posess remarkable spines at the outer, lateral margin, which are maximally developed on segments IX–XI of the flagellum. No spines are present on the posterior margins of abdominal tergites I–VI. COI sequences were obtained from all three of the new species. The genetic distances (Kimura 2-parameter) between these species are between 13% and 20%. Very limited genetic distances of 0% to 1% were found between specimens of the same species. The occurrence of two different species in the same area of Sumatra is discussed.
Recent samplings in cavities of the Iberian Peninsula led to the discovery of a new millipede species of the order Callipodida in one cave in the region of El Bierzo (León, Spain): Lusitanipus xanin sp. nov. In additional samplings (near that cave), more specimens of this new species were discovered in surface habitats. The species has various distinctive characteristics, such as its green colour, but the morphology of the gonopod has several similarities with that of Lusitanipus alternans (Verhoeff, 1893), indicating a close relationship between them. The diagnosis of the genus Lusitanipus Mauriès, 1978 has to be changed to accommodate the new species. The presence of crests in two different sizes is no longer a diagnostic character of the genus Lusitanipus, but of the species L. alternans. Lusitanipus xanin sp. nov. is the fourth species of the order Callipodida for the Iberian Peninsula, and the second species for the genus. Lusitanipus xanin sp. nov. seems to be an epigean but troglophile species.
A new microscopic cnidarian is described, infecting five of thirty (16.6%) specimens of Corydoras schwartzi caught in the Purus River, State of Amazonas, Brazil. Histological analysis showed that cyst development occurred in the serosa layer of the intestine. Mature myxospores are ovoid in body shape in frontal view, 22.4 ± 0.3 μm in total length and 16.3 ± 0.1 μm in width. Internally, two aubergine-shaped, elongate symmetrical polar capsules occupy more than half the length of the spore, 14.3 ± 0.2 μm in length and 6.5 ± 0.1 μm in width. Ultrastructural analysis provided evidence of five polar filament coils inside the polar capsule and binucleated sporoplasm containing a moderate number of sporoplasmosomes. The valvogenic cells abutting each other form a sutural ridge and frequently a thin layer of homogeneous material separates the cells. The outer surface of the myxospore valves is smooth, with no evidence of formation of ridges in the valves. Immature myxospores in various stages of development were observed. This study is the first report of a myxosporean parasitizing C. schwartzi and the first report of a myxosporean infection in the intestine of an ornamental fish from South America.
The branchial parasitic isopod Pleurocryptella altalis sp. nov. (Bopyridae: Pseudioninae) is described from the squat lobster host Munidopsis petalorhyncha Baba, 2005. The new species is morphologically similar to Pleurocryptella formosa Bonnier, 1900 and P. wolffi Bourdon, 1972b but can be distinguished based on male characters (differences in head, pleon and uropods) and female characters (differences in barbula, pleopods and pleotelson). The parasite specimens (a female and male pair) were collected with the squat lobster host at a depth of 5060–5130 m from the Kuril-Kamchatka Trench, representing the deepest record for any of the 850+ described bopyrid isopod species and for any record of an infested host. Dichotomous identification keys to females and males of Pleurocryptella species and subspecies are provided.
A new species of praying mantis, Vates phoenix sp. nov. (Mantidae, Vatinae), is described from localities within Rio de Janeiro and São Paulo states, Brazil. This is the first record of Vates from the Atlantic Rainforest biome. The new species is unique among its congeners in having, among other features, strongly reduced cuticular projections above the lateral ocelli, structures otherwise well developed and produced in all other species of Vates. Remarks on the natural history and biogeography of Vates, in relation to this new finding, are further discussed.
We review the species of Bidessus of Madagascar and describe Bidessus anjozorobe sp. nov. from material collected in Anjozorobe forest. Anjozorobe is part of the Anjozorobe-Angavo Protected Area, which is an important corridor of transition forest between typical eastern humid forests and the residual sub-humid forest of the Central Highlands. Bidessus longistriga Régimbart, 1895 and Bidessus perexiguus Kolbe, 1883 are widespread but endemic low-altitude species on Madagascar. Bidessus nesioticus Guignot, 1956 is an alpine species described from near the peak of the Ankaratra mountain massifs at 2500 m a.s.l. We recollected the species for the first time since its description, in Ankaratra and in a new area above 2000 m a.s.l. in the Andringitra mountain further south. Bidessus cf. nero Gschwendtner, 1933 is tentatively recorded for Madagascar for the first time but further studies are needed to test the status of mainland and insular populations. Bidessus apicidens Biström & Sanfilippo, 1986 has not been recollected on Madagascar since 1970. All species are endemic to Madagascar except potentially Bidessus cf. ceratus and Bidessus cf. nero described from Uganda and the Democratic Republic of the Congo, respectively. The older records of the two non-endemic species Bidessus complicatus Sharp, 1904 and Bidessus ovoideus Régimbart, 1895 on Madagascar could not be verified.
Chaetopterus is a globally distributed genus of marine Annelida with a long history of taxonomic confusion. Here, we describe Chaetopterus bruneli sp. nov. from a depth of 350 m in the St. Lawrence Estuary, eastern Canada. The new species represents the northernmost record for Chaetopterus in the western Atlantic to date. The similar European species Chaetopterus norvegicus M. Sars, 1835 is resurrected from long-standing synonymy and redescribed from type material, and a lectotype is designated.
Crioprosopus baldwini Eya (Coleoptera: Cerambycidae: Cerambycinae: Trachyderini), sp. nov., from Costa Rica and Panama, is described. An amendment to the key to species of Crioprosopus Audinet-Serville, 1834, as presented in Eya (2015), is provided along with illustrations of the key characteristics to differentiate C. baldwini from other species.
A new species, Cryptomonas uralensis Martynenko, Gusev, Kulizin & Guseva sp. nov., is described from western slopes of the Ural Mountains (Russia) based on morphological and molecular data. Phylogenetic relationships inferred from nuclear SSU and LSU rDNA sequences show that the new species forms a clade with C. tetrapyrenoidosa Skuja emend. Hoef-Emden & Melkonian. Comparison of secondary structures of nuclear rDNA ITS2, including analysis of Compensatory Base Changes (CBC), confirms the separation between C. uralensis sp. nov. and C. tetrapyrenoidosa. Cell morphology and sizes of C. uralensis sp. nov. are very similar to C. tetrapyrenoidosa and C. pyrenoidifera, and C. uralensis sp. nov. may thus represent a species that can only be reliably identified using molecular data.
Diolcus thomasi Eger new species (Hemiptera: Heteroptera: Scutelleridae: Pachycorinae), is described from Cayman Brac and Little Cayman Island and compared to D. chrysorrhoeus (Fabricius), its closest congener. Nesogenes boscii (Fabricius) is removed from the Elvisurinae and placed in the Pachycorinae. The relationship of N. boscii to D. thomasi and D. chrysorrhoeus is discussed.
Oncotophasma aurantiaviridiata Murcia, Cadena-Castañeda, and Silva new species (Phasmida: Diapheromeridae)from the Colombian Andes is described. The new species is distinguished by its peculiar coloration, which highlights greenish and orange tones. It is also the first record of the genus for the eastern slope of the Colombian Andes. The previous distribution recorded for the genus is the Chocó biogeographic region from Costa Rica to Colombia, with species being found in the Colombian areas of influence of this biogeographic region, such as the western and central slopes of the Andes.
A new species of tortoise beetle from the north-central United States, Parorectis arenaria new species (Coleoptera: Chrysomelidae: Cassidinae: Cassidini), is described and illustrated. Comparative remarks and a key are provided to distinguish the new species from the three other species comprising the genus. Adults and larvae of the new species feed on Physalis L. (Solanaceae). The circum-foraminal ridge and antennal groove of the prothorax are described in detail. The pars stridens (file) of the stridulatory apparatus on the male cranium of the new species is illustrated. The pars stridens is present only in males of Parorectis Spaeth species. A patch of spicules is located centrally on the dorsal surface of the cranium in both males and females of Parorectis. The spicule patch is believed to function as a head-to-body binding patch.
Pteroplatus antonkozlovi Santos-Silva and Botero, sp. nov. (Coleoptera: Cerambycidae) is described from Panama. Neocompsa thelgema Martins,1971 is recorded from Panama, and chromatic variation is discussed.The female of Icimauna aysa Martins and Galileo,1991 is described. Some corrections in a recently published work on Acanthoderini are provided, and the combination Scythropopsis pupillata (Bates,1880) is established as having precedence over the simultaneously published combination Aegomorphus pupillatus (Bates,1880).
A new species of snail-eating snakes of the genus Pareas Wagler, 1830 is described from the eastern Himalayas. The species Pareas kaduri sp. nov. differs from all known species of the genus in bearing the following suite of characters: SVL 455–550 mm, TaL/TL 0.184–0.207, brown dorsum with black transverse bands throughout the body, 15 dorsal scale rows throughout the body and mid-dorsal vertebral scale rows enlarged, 8 rows keeled in males, loreal not touching orbit, ventrals 160–183, subcaudals 65–70 in males, 52 in one female specimen, hemipenis short, unilobed and 6–7 maxillary teeth. Molecular data for mitochondrial 16S rRNA and cytochrome b genes further attest the distinctness of the new species, which was recovered as a member of the Pareas hamptoni clade. Our work brings the total number of species recognized within the genus Pareas to 20.
A new symbiotic species of liljeborgiid amphipods, Liljeborgia associata sp. nov., is described from the burrows of the spoon worm Urechis unicinctus (Drasche, 1880) (Annelida: Polychaeta: Echiura: Urechidae) in the southern part of Peter the Great Bay and Posjeta Bay in the Sea of Japan. The new species is mostly similar and probably related to Liljeborgia geminata Barnard, 1969, known from the Californian coasts of the USA, and Liljeborgia serratoides Tzvetkova, 1967, described from Posjeta Bay in the Sea of Japan, but can be clearly distinguished from all congeners by morphological features of mouthparts, appendages and telson. The new species is only the fourth in the family Liljeborgiidae to be described from the Russian coast of the northwestern Pacific and the first in association with spoon worms (Echiura).
We describe a new species of the genus Zhangixalus Li, Jiang, Ren & Jiang, 2019 from Ha Giang Province, Vietnam based on morphological and molecular data. In the molecular phylogenetic analyses, the new species is nested in the Zhangixalus duboisi (Ohler, Marquis, Swan & Grosjean, 2000) group, where it is sister to Z. duboisi with a genetic distance of 2.51%. The new species, Zhangixalus franki sp. nov., differs from its congeners by a combination of the following morphological characters: size large, SVL 77.9–85.8 mm in males; finger webbing formula I1-1II0-1III½-0IV; dorsal surface of head and body green with dark brown spots; ventral surface grey or dark grey, white stripe along edge of jaw, insertion of limbs, and along lateral ridges of fore and hind limbs and flank, separating upper green part from lower grey part. The new species occurs in evergreen montane tropical forests at an elevation of ca 1300 m a.s.l. The new discovery brings the total number of known species in the genus Zhangixalus to 38 and the species number reported from Vietnam to nine.
Over the past decade, a previously unrecorded spittlebug of the genus Clastoptera Germar (Hemiptera: Cercopoidea: Clastopteridae) has been observed in abundance on oaks (Quercus L. spp., Fagaceae) in several Florida counties. We describe this spittlebug as a new species, Clastoptera querci Thompson, Halbert and Rothschild, new species, provide information on its life history, host plants and distribution, and place it in the context of other members of the genus. Clastoptera spp. can transmit Xylella fastidiosa Wells et al., a bacterial pathogen that causes bacterial leaf scorch associated with oak decline. Thus C. querci should be monitored as a possible vector of X. fastidiosa in oaks.
This paper describes a novel species of the genus Heterophoxus Shoemaker, 1925 from Brazil. The material examined was collected during the Mini Biological Trawl Project, off southeast and southern Brazil's coast off the states of Rio de Janeiro, Santa Catarina, and Rio Grande do Sul, ranging in depth from 25 to 128 meters. The new species, H. shoemakeri sp. nov., is similar to H. videns K.H. Barnard, 1930; however, the former species can be distinguished by: pereopod 7 basis posterior margin serrate with acute teeth, merus posterior margin not produced; epimeral plate 3 posteroventral corner with a longer acute projection, posterior margin almost straight; uropod 3 outer ramus article 2 clearly notable in both sexes. For Brazilian waters, this is the second record of the genus and the first species described with type material from the country. Besides that, we provide an identification key for the world species of Heterophoxus and a distribution map for the new species.
Nothochodaeus marsupialis Paulsen, new species (Coleoptera: Ochodaeidae), is described from Aceh Province, Indonesia, on the island of Sumatra. The species possesses deep abdominal cavities behind a shield-like central process, as well as rows of conical tubercles laterally on the second and third ventrites. These structures have not been found in any other members of the genus or family.
The Colaspis suilla species group (Coleoptera: Chrysomelidae: Eumolpinae) is defined relative to other species of the genus Colaspis Fabricius occurring in the United States. The group is composed of five species, of which three from Florida are described as new species: C. ansa Riley from the Florida Panhandle, C. skelleyi Riley from Central Florida, and C. thomasi Riley from the southern Lake Wales Ridge. Colaspis suilla borealis Blake is reduced to a full synonym of C. suilla Fabricius, new synonymy. Comparative remarks, habitus images, images of male and female genitalia, range maps, specimen data, and a key to species are presented.
Indian spider species currently assigned to Storena Walckenaer, 1805 are revised mostly based on the type material available in the National Zoological Collection, Zoological Survey of India, Kolkata. A new genus, Laminion gen. nov. is proposed to include four species; three are transferred from Storena: Laminion arakuensis (Patel & Reddy, 1989) gen. et comb. nov., Laminion birenifer (Gravely, 1921) gen. et comb. nov. and Laminion debasrae (Biswas & Biswas, 1992) gen. et comb. nov., whereas the fourth species is from Suffasia Jocqué, 1991: Laminion gujaratensis (Tikader & Patel, 1975) gen. et comb. nov. The species Storena tikaderi Patel & Reddy, 1989 syn. nov. is synonymised with L. birenifer gen. et comb. nov. Storena dibangensis Biswas & Biswas, 2006 and Storena indica Tikader & Patel, 1975 are transferred to Mallinella Strand, 1906. All the type material examined are imaged and redescribed. In addition, images of the type material of Storenomorpha joyaus (Tikader, 1970) are presented.
The Nearctic species of Fornax Laporte are reviewed. Three species are redescribed, and six new species are described mainly from southeastern United States. The new species are F. appalachiensis, F. convexicollis, F. floridana, F. lucidicollis, F. melsheimeri and F. parallelicollis. A neotype is designated for Hylochares bicolor Melsheimer, with notes on its purpose. Each species is diagnosed and imaged. A new identification key is provided for all known species of Fornax present in the Nearctic region, replacing previous Nearctic species keys of Muona (2000) and Otto (2017).
This paper examines the described species of Cissidium and adds 77 new ones based on a study of material in the Manchester Museum, UK, the Natural History Museum, London, UK and in the possession of Dr Yoshihiro Sawada, Japan. The new taxa are: Cissidium advena sp. nov., C. amazonicum sp. nov., C. amieuense sp. nov., C. angulatum sp. nov., C. angusi sp. nov., C. apterum sp. nov., C. arcuatum sp. nov., C. aristophanousi sp. nov., C. besucheti sp. nov., C. bifoveolatum sp. nov., C. bilineatum sp. nov., C. bomjesus sp. nov., C. brachypterum sp. nov., C. ceylonicum sp. nov., C. clareae sp. nov., C. cryptophagoides sp. nov., C. davaoense sp. nov., C. deanei sp. nov., C. delicatum sp. nov., C. dewhurst sp. nov., C. dividuum sp. nov., C. dybasi sp. nov., C. eciton sp. nov., C. ecuadoriense sp. nov., C. faustum sp. nov., C. flavum sp. nov., C. franzi sp. nov., C. fraternum sp. nov., C. glabratum sp. nov., C. greensladei sp. nov., C. houailou sp. nov., C. ibicarense sp. nov., C. impressum sp. nov., C. inexspectatum sp. nov., C. insulare sp. nov., C. kolombangaricum sp. nov., C. lamington sp. nov., C. lawrencei sp. nov., C. loebli sp. nov., C. logunovi sp. nov., C. losbanos sp. nov., C. lucidulum sp. nov., C. luctuosum sp. nov., C. mahleri sp. nov., C. mindanao sp. nov., C. misellum sp. nov., C. modestum sp. nov., C. montanum sp. nov., C. murphyi sp. nov., C. mussardi sp. nov., C. noumeae sp. nov., C. obscenum sp. nov., C. obscurum sp. nov., C. obsoletum sp. nov., C. opacum sp. nov., C. orami sp. nov., C. peruviense sp. nov., C. pilosellum sp. nov., C. pinense sp. nov., C. plaumanni sp. nov., C. puncticolle sp. nov., C. riparium sp. nov., C. robustum sp. nov., C. rogeri sp. nov., C. seditiosum sp. nov., C. semicalvum sp. nov., C. similatum sp. nov., C. steeli sp. nov., C. subangulatum sp. nov., C. subfaustum sp. nov., C. sueae sp. nov., C. trangse sp. nov., C. upiense sp. nov., C. visendum sp. nov., C. waginicum sp. nov., C. werneri sp. nov. and C. yoruba sp. nov. The division of the genus is discussed and five species groups are established. The majority of the new species are the first records of Cissidium for the 28 countries represented.
The genus Rudolfina Roháček, 1987 is revised and redefined with the description of the following nine new species, all from the New World: R. bucki sp. nov. (Mexico), R. exuberata sp. nov. (widespread, from USA to Brazil), R. howdeni sp. nov. (Mexico), R. megepandria sp. nov. (Mexico), R. newtoni sp. nov. (Mexico), R. pauca sp. nov. (Guatemala, Mexico), R. pilosa sp. nov. (Mexico), R. remiforma sp. nov. (Mexico) and R. tumida sp. nov. (Mexico, USA). Rudolfina is compared to closely related genera in the Archiceroptera genus complex, which in turn is recognized as part of a large, mostly Neotropical clade including Robustagramma Marshall & Cui, 2005, Pterogramma Spuler, 1924, Aptilotella Duda, 1924, Bitheca Marshall, 1987, Bromeloecia Spuler, 1924 and Archiceroptera Papp, 1977.
Features of the Maechidiini (Scarabaeidae: Melolonthinae) genera Maechidius Macleay, 1819, Epholcis Waterhouse, 1875 and Paramaechidius Frey, 1969 are critically revised and a new synonymy is proposed: Maechidius = Epholcis syn. nov. = Paramaechidius syn. nov. A key to and an annotated checklist of Maechidiini from the Indo-Australian transition zone are presented for the first time. Thirty-five new species are described, namely Maechidius aiyura sp. nov., M. alesbezdeki sp. nov., M. awu sp. nov., M. babyrousa sp. nov., M. bintang sp. nov., M. boessnecki sp. nov., M. brocki sp. nov., M. caperatus sp. nov., M. ciliatus sp. nov., M. crypticus sp. nov., M. dani sp. nov., M. deltouri sp. nov., M. dendrolagus sp. nov., M. hamatus sp. nov., M. kazantsevi sp. nov., M. konjo sp. nov., M. lapsus sp. nov., M. legalovi sp. nov., M. leucopsar sp. nov., M. longipes sp. nov., M. mailu sp. nov., M. maleo sp. nov., M. merdeka sp. nov., M. miklouhomaclayi sp. nov., M. nepenthephilus sp. nov., M. owenstanleyi sp. nov., M. riedeli sp. nov., M. similis sp. nov., M. skalei sp. nov., M. sougb sp. nov., M. suwawa sp. nov., M. trivialis sp. nov., M. ursus sp. nov., M. weigeli sp. nov. and M. yamdena sp. nov. Six new synonyms are proposed: Maechidius Macleay, 1819 = Epholcis Waterhouse, 1875 syn. nov. = Paramaechidius Frey, 1969 syn. nov., Maechidius esau Heller, 1914 = M. setosus Moser, 1920 syn. nov. = M. setosellus Frey, 1969 syn. nov., Maechidius heterosquamosus Heller, 1910 comb. rest. = Paramaechidius clypeatus Frey, 1969 syn. nov. and Maechidius paupianus Heller, 1910 = M. arrowi Frey, 1969 syn. nov. The first records of Maechidiini from the Tanimbar Islands (Yamdena), Sangihe Islands (Sangir) and Lesser Sunda Islands (Bali) are documented, of which the latter two are the northern- and westernmost known records of Maechidius and of the tribe Maechidiini. Lectotypes are designated for 23 species. Fifteen new combinations are proposed and the original combination to Maechidius is restored for four species. Ecological data are presented for the first time for selected Papuan and Wallacean species. Type material of Wallacean and Papuan Maechidiini is depicted for the first time. A key to species is given. In total, 78 species of Maechidiini are confirmed for the Indo-Australian transition zone.
Eight species of the genus Prodasineura Cowley, 1934 are recorded from Vietnam, including two newly described species: Prodasineura lancastrei sp. nov. and P. kong sp. nov. from north and central Vietnam. All species recorded are illustrated, including figures of morphological structures, and distribution maps and keys to the males and females are provided. We consider the record of P. laidlawi (Forster in Laidlaw, 1907) a misidentification and exclude it here.
For more than 100 years, behavioural biologists have extensively studied satin bowerbirds (Ptilonorhynchus violaceus) to understand the complexities of bower construction and courtship, but this research has not accounted for the ecologically important role the species plays as a seed disperser in eastern Australia. In this study, we have used a simple method involving pans of water and mirrors to collect scats from male satin bowerbirds on the Dorrigo Plateau of northeastern New South Wales. Based on seeds identified from scats collected using this method, we identify the fruits of 37 plant species in 22 families that are eaten by satin bowerbirds and show that 35 of these species germinated after passage through the digestive system of bowerbirds. This method was developed for bush regenerators to collect rainforest tree seeds that are otherwise inaccessible and/or difficult to germinate; it is applicable throughout the range of satin bowerbirds. We hope the method will be adopted by bush regenerators for harvesting seeds otherwise difficult to source, and by ornithologists studying the diets and seed dispersal of these ecologically significant birds.
A new giant species of the subfamily Rhynchoproctinae with strikingly bi-coloured, red-yellow legs contrasting with a black body is described: Alienostreptus bicoloripes sp. nov. from Vietnam. The new species is assigned to the genus Alienostreptus Pimvichai, Enghoff & Panha, 2010 hitherto comprising one species, A. alienus (Attems, 1936), and differing from other rhynchoproctine genera by having the femoral spine duplicated. Species of this genus share three synapomorphic characters of the subfamily, viz (1) anterior coxal fold forms deep concavity in posterior view, (2) posterior coxal fold very low, and (3) posterior coxal fold with mesal flap. A superficially very similar colourful species from Borneo, also with bi-coloured legs but clearly not belonging to Alienostreptus due to the position of ventral soft pads on male legs, is documented based on photographs.
The present study took advantage of the availability of high resolution ADS40 digital imagery to 1) systematically resample the vegetation of the Lord Howe Island Group (LHIG, excluding Ball's Pyramid); 2) conduct a numerical analysis of the floristic data; 3) map vegetation extent and the distribution of vegetation communities and 4) compare the resultant classification and mapping with those of Pickard (1983). In July 2013, a total of 86 full floristic and 105 rapid floristic sites were sampled across the island, based on a stratified random sampling design. A hierarchical agglomerative clustering strategy (Flexible UPGMA) and Bray-Curtis dissimilarity coefficient with default beta, along with nearest neighbour analysis to identify anomalous site allocations, was used to analyze the floristic data. In total 33 vegetation communities were delineated and mapped: 19 mapping units from the full floristic analysis; 7 variants identified within five of the above 19 groups; 3 mapping units from analysis of canopyonly floristic data; and 4 mapping units recognised in previous studies that are mapped but were not sampled in this survey. The resultant list of vegetation communities and non-vegetation mapping units, along with their equivalence to Pickard (1983) and DECC (2007) units, is provided. 222 plant taxa were recorded in the survey, including 47 exotic taxa. Weeds are a common component of some communities, particularly coastal strandline communities, the shrublands of the southern mountains and regenerating vegetation on landslips. The threatened plants Xylosma parvifolia, Lepidorrhachis mooreana, and Geniostoma huttonii were recorded in floristic sites. Two communities, Gnarled Mossy Cloud Forest and Lagunaria Swamp Forest are listed as Critically Endangered Ecological Communities under the NSW Biodiversity Conservation Act 2016 (BC Act). Coastal Saltmarsh, an Endangered Ecological Community listed under the BC Act, also occurs on the main island. An assessment of the conservation status of the vegetation communities recognised in the present study is provided. A number of communities warrant further consideration for listing as threatened ecological communities. Significant improvement in vegetation community attribution and spatial resolution was possible with the highresolution digital imagery, however Pickard's 1983 classification and mapping was found to be a comprehensive and accurate description of the island’s vegetation considering the imagery available at the time. This project has resulted in greatly improved accuracy of vegetation mapping linework for the LHIG. The Pickard (1983) vegetation map comprised 321 individual polygons, whereas the new map includes 1 840 polygons of sufficient accuracy to support detailed environmental planning programs, particularly within the Settlement area where spatial accuracy in the delineation of native vegetation is critical. The new vegetation communities were applied to the updated linework to complete the project. Detailed profiles were compiled of each vegetation community for which enough information was available. These profiles should prove useful in field identification of vegetation types and the assessment of their conservation status.
Neogene (Siwalik-aged) deposits from India and Pakistan have yielded many vertebrate fossils, of which most were named during the 19th century, including numerous geoemydid turtles. In contrast to many other faunal components from the Siwaliks, geoemydids have not undergone taxonomic revision for more than a century and most fossils have therefore been believed to correspond to recent taxa. In this study, we conduct a taxonomic revision of all previously described geoemydid material from the Siwalik-age. We propose that all specimens of 'Clemmys' from the Siwaliks of Punjab, Pakistan should be identified as Melanochelys sivalensis comb. nov.; that Melanochelys tricarinata var. sivalensis represents a valid species, for which we propose the replacement name Melanochelys tapani to avoid homonymy; that specimens originally identified as Batagur cautleyi and Pangshura flaviventer cannot be identified beyond the generic level; and that many fragmentary palatochelydians cannot be identified to any particular species or genus due to the lack of preserved diagnostic osteological characters. With a few exceptions, the Siwalik fauna mostly corresponds in its distribution to that of the recent fauna, indicating a certain amount of geographic stasis. However, as the stratigraphic provenance of most material is poor, it is not possible to discern meaningful temporal patterns.
The catalogue of species-group names of nine coprine genera (Coleoptera: Scarabaeidae: Scarabaeinae: Coprini), published earlier this year, is updated and revised. Presented are species-group taxa overlooked in compiling the catalogue and new species-group taxa described in 2020, i.e. after the cut-off date of the catalogue.
Close-up photographs of nest entry, nest closure and prey transport taken on sandy coastal back dunes in Santa Barbara County, CA by Alice J. Abela substantiate and enhance written descriptions of these nesting behavior components in Miscophus californicus (Ashmead) [=M. laticeps (Ashmead)] (Hymenoptera: Crabronidae). Dictynidae (Dictyna Sundevall or Emblyna Chamberlin) is introduced as a new host family and host spider leg amputation is revealed for the first time for this small miscophine wasp.
We report here the results from field trips to collect Odonata in the northeastern parts of Kelantan state and the north of Terengganu state, Peninsular Malaysia. Eighty four species were collected, and four of these are new records for the state Kelantan and 10 are new records for the state of Terengganu. Notable records obtained from the field trips were Euphaea masoni, Coeliccia sameerae, Pseudagrion ?lalakense, Leptogomphus tioman and Macromia cupricincta. Checklists for Kelantan (140 species) and Terengganu (132 species) are given in an appendix.