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The planthopper family Caliscelidae (Hemiptera: Auchenorrhyncha: Fulgoromorpha) and the tribe Adenissini are recorded here for the first time from the Socotra Archipelago (Yemen). A new genus, Oceatra gen. nov., is described for two new species from Socotra Island – Oceatra litoralis gen. et sp. nov. (type species) from coastal sand dunes and salt marshes, and O. scandens gen. et sp. nov. from evergreen montane woodland and shrub communities of the Hagher mountains. Oceatra gen. nov. is similar to Perissana Metcalf, 1952 but differs in the structure of the male style and phallobase. A key to distinguish the genera of the subtribe Adenissina is provided.
Adenomera is a genus of Neotropical leaf-litter frogs widely distributed in South America and regarded taxonomic-wise challenging. One of these is the open-habitat Adenomera diptyx from Paraguay, which may correspond to a species complex. An integrative analysis of morphological variation, in combination with acoustic and molecular data of several populations from Argentina, Paraguay, and Brazil resulted in the recharacterization of nominal A. diptyx and the description of a new, closely related species. Adenomera diptyx is recognized by its advertisement call given at a high repetition rate (176–299 per minute), and the dorsal color pattern consisting of a light mid-dorsal line and a dark brown interorbital bar with irregular black edges. In comparison with A. diptyx, the new species, Adenomera guarani sp. nov., produces its advertisement call at a lower repetition rate (73–147 per minute), and the dorsal color pattern consists of a light, broad, mid-dorsal stripe and a mask-like patch in the interorbital region. The redefinition of A. diptyx will contribute to future studies focusing on the taxonomic status of other genetic lineages tentatively assigned to this species complex, which could represent additional unnamed species in the open-habitat Adenomera clade.
A checklist with preliminary conservation assessments of native South American species of Acalypha is presented. This work is supported by the study of ca 6500 herbarium specimens and an in-depth literature review. As a result, 87 species (83 native and four introduced) and eight subspecies are accepted, and a further 395 names are considered synonyms. Geographical distribution, habitat, and altitudinal range for all species are also indicated. Brazil is the richest country in number of species of Acalypha (40), followed by Peru (32), Bolivia (29), Colombia and Ecuador—including Galapagos Islands—(24), Venezuela (18), Argentina (17), Paraguay (13), Guyana (8), Uruguay (5), French Guiana (4), and Suriname (3). The presence of the genus Acalypha in Chile is reported for the first time, alongside new country records of A. poiretii in Peru and A. venezuelica in Guatemala. The specimens previously identified as A. plicata from Colombia and Venezuela, are here considered belonging to A. cuspidata. The red list provided follows IUCN criteria and includes 39 species and three subspecies, 47% of total native species of Acalypha in South America: 16 species and one subspecies Critically Endangered (nine of them probably extinct), 15 species and two subspecies Endangered, and eight species Vulnerable.
The taxonomy of Loboscelidiinae in Vietnam is revised, with 16 new species being described: Loboscelidia bachmaensis sp. nov., L. barbata sp. nov., L. cilia sp. nov., L. convexa sp. nov., L. cucphuongensis sp. nov., L. cuneata sp. nov., L. do sp. nov., L. flavipes sp. nov., L. glabra sp. nov., L. komedai sp. nov., L. mediata sp. nov., L. parallela sp. nov., L. piriformis sp. nov., L. squamosa sp. nov., L. vang sp. nov. and L. vietnamensis sp. nov. In total, 24 species of Loboscelidia are recognized in the fauna of Vietnam. Keys to Indo-Chinese male and world female of Loboscelidia are provided. A brief observation of the foraging behavior of L. squamosa sp. nov. is also reported. Host-carriage and subsequent host egg burying are considered primary nesting behaviors of solitary wasps.
Viséan coiled nautiloids from North Africa are only poorly known. From the Mougoui Ayoun, Zrigat and Hamou-Rhanem formations of the eastern Anti-Atlas, we describe coiled nautiloids, which belong to the genera Rineceras, Stroboceras, Temnocheilus, Vestinautilus, Maccoyoceras, Endolobus, Epidomatoceras, Liroceras, Ephippioceras, and Solenochilus. The new species Temnocheilus imazighenorum sp. nov., Temnocheilus aubrechtovae sp. nov., Vestinautilus kesslerae sp. nov., Endolobus rota sp. nov., Epidomatoceras ebbighausenorum sp. nov., Liroceras vermis sp. nov., Liroceras karaouii sp. nov., Ephippioceras pygops sp. nov., Solenochilus lucynae sp. nov. and Solenochilus pohlei sp. nov. are described; six taxa are kept in open nomenclature. The assemblage is composed of the three superfamilies Trigonoceratoidea, Clydonautiloidea and Aipoceratoidea and shows a wide spectrum of conch morphologies, ranging from widely umbilicate compressed forms to involute compact forms, reflecting a broad ecological variation.
Spondias purpurea L. (Anacardiaceae), native to the Neotropical region, is cultivated in the Philippines for the edible fruits and the tree is economically significant. The adventive leaf beetle, Podontia quatuordecimpunctata (L.) (Chrysomelidae: Galerucinae: Alticini), has become a major defoliating pest of the tree in the country. The Philippines government has initiated study of the pest, now locally called the sineguelas leaf beetle (SLB). This paper reports the results of a one-year field study on the biology of SLB on S. purpurea (red sineguelas), in Batangas City, Philippines. The SLB eggs hatch in 5–7 days. The larval period is 14–16 days with 4 larval instars, the pre-pupal period is 2–3 days, and the pupal period is 15–22 days. The total life cycle from egg to adult emergence is completed within 36–48 days. Host-choice experiments revealed that different stages of SLB do not feed on carabao mango (Mangifera indica L.) or pili (Canarium ovatum Engl.), however, they fed a little but did not survive or reproduce on cashew (Anacardium occidentale L.). A predatory bug (Eocanthecona furcellata Wolff., Hemiptera: Pentatomidae) is a potential natural enemy. Unidentified fungi infecting the pupae and adults of SLB were also recorded.
ZooBank registration. urn:lsid:zoobank.org:pub:28A1186E-589B-481A-A868-DE8C1E994352
A catalog of the species of Trichodesma LeConte (Coleoptera: Ptinidae) for the world is provided, including synonyms and misspellings. Six additional genera, Anobiopsis Fall, Belemia Español, Nanodesma Zahradník, Nicobium LeConte, Trichobiopsis White, and Trichodesmina Español, are also cataloged for completeness. The species name, author, year, and page number of description are given for each species, as well as references for combinations by different authors. The type depository is given when known, and followed by a “?” when unknown but suspected, and a general distribution is given by region of the world followed by country. Authorship of the genus Trichodesma in Lepidoptera is transferred to Schaus.
ZooBank registration. urn:lsid:zoobank.org:pub:99BDA976-4B58-4D42-8CF5-4D3034B95C7B
The Egyptian fauna of mealybugs (Hemiptera: Coccoidea: Pseudococcidae) is reviewed and an illustrated key to the 30 genera and 54 species is provided. Phenacoccus madeirensis Green is reported for the first time in Egypt. A new genus, Ezzatacoccus Evans and Abd-Rabou, is described and illustrated with Amonostherium arabicum Ezzat, 1960 designated as its type species. Octococcus salicicola Priesner and Hosny, 1935 is reinstated as a valid taxon and transferred to Misericoccus Ferris, new combination. Ripersia cressae Hall is transferred to Maconellicoccus Ezzat, new combination and Planococcus lindingeri (Bodenheimer) is transferred back to Formicococcus Takahashi, revised status.
ZooBank registration. urn:lsid:zoobank.org:pub:8CA7B000-E8D4-463D-95B0-431BA0A7BA57
A new species of Strepsiptera of the genus Paraxenos Saunders, 1872 (Xenidae) from the United Arab Emirates is described. It was recorded from the host species Bembix kohli Morice, 1897 and represents the first occurrence of Paraxenos from Bembix Fabricius, 1775 in the Afrotropical region. A detailed redescription of the female cephalothorax of Paraxenos hungaricus (Székessy, 1955) is provided, together with the first description of the male cephalotheca. The holotype of Paraxenos krombeini Kifune & Hirashima, 1987 was redescribed. Additionally, a key for parasites of Bembix among Paraxenos species is provided based on characters of the female cephalothorax and male cephalotheca. The distribution and conservation status of Paraxenos spp. on Bembix are also discussed.
Eight new species of Itauara Müller, 1888 are described based on specimens collected in southeastern Brazil: I. bispinata sp. nov., I. caparao sp. nov., I. cipoensis sp. nov., I. cristata sp. nov., I. holzenthali sp. nov., I. mangaratiba sp. nov., I. robertsonae sp. nov., and I. rupicola sp. nov. In addition, we provided an updated distributional list of species of Itauara, with new records of I. jamesii Robertson & Holzenthal, 2011 and I. lucinda Robertson & Holzenthal, 2011 for Espírito Santo State (Brazil), I. plaumanni (Flint, 1974) for Paraná State (Brazil), and I. tusci Robertson & Holzenthal, 2011 for Minas Gerais State (Brazil). We performed an equal weighted parsimony analysis adding the new species to the dataset provided by Robertson & Holzenthal (2013), with modification of the interpretation of some morphological characters. The genus was recovered as monophyletic, but overall statistic support for clades was weak. Itauara is restricted to South America and has a disjunct distribution, with some species occurring in northern South America and others in southeastern South America. Probably, the diversification of Itauara in South America is related to the connections between the Amazon Forest and the Atlantic Forest through historical events at different times.
The chelodesmid genus Dibolostethus Hoffman, 2009, the sole member of the tribe Dibolostethini, is reviewed. The genus contains the type species D. sicarius Hoffman, 2009 known only from the Los Rios Province, Ecuador, and two new species from the Tropical Andes, D. inopinatus Means, Bouzan & Ivanov sp. nov. from the Morona-Santiago Province, Ecuador and D. kattani Means, Bouzan, Martínez-Torres & Ivanov sp. nov. from the Valle del Cauca Department, Colombia. We redescribe D. sicarius and provide a revised diagnosis of the genus, images of diagnostic morphological characters, and a key to the males of Dibolostethus. In addition, we provide a summary and a distribution map of the Chelodesmidae of the Tropical Andes Biodiversity Hotspot.
A new spionid polychaete, Prionospio expansa sp. nov., collected from the intertidal habitat of the Yellow Sea in Korea, is described. The new species is closely related to P. japonica Okuda, 1935 from Northeast Asia both morphologically and genetically. They share four pairs of branchiae which are cirriform and apinnate, whereas the new species differs from P. japonica in the length of the branchiae, expansion of the anteriormost body, and size of the first notopodial postchaetal lamellae. Herein, a detailed description and illustrations of P. expansa sp. nov. are provided, with molecular data for three gene fragments: mitochondrial cytochrome c oxidase subunit I, 16S ribosomal DNA (rDNA), and nuclear 18S rDNA. A phylogenetic analysis was conducted based on mitochondrial and nuclear gene fragments.
Iberian material originally identified as Hincksina flustroides is revised and five different species are now identified: the Atlantic species Hincksina flustroides is present to the NW of the Iberian Peninsula, whilst the Mediterranean species Hincksina synchysia is here reported for the first time in Iberian waters. Two new species of Hincksina are described, one from the Strait of Gibraltar area, and another from the Alboran Sea, whereas another colony from this area actually corresponds to Hincksina longispinosa. A new description of Electra angulata is given, based on the original and other historical material, as well as on newly collected colonies. The taxonomic position of the species in the genus Arbopercula is discussed, and the diagnosis of the genus itself is amended. Finally, a new species of Caberea is described from deep waters off the north Iberian coast.
A new genus, Plesioxylion Liu & Beaver gen. nov., is described for Amintinus gambianus Borowski, 2018 from West Africa with a more detailed description and new records of both sexes. We also provide a key to the ten Afrotropical genera in the tribe Xyloperthini Lesne, 1921 as the baseline information for a future study.
In this study, we revise the taxonomy of the endemic Mexican genus Ceratotrupes Jekel, 1865. It comprises three species, distributed in the Sierra Madre Occidental and Trans-Mexican Volcanic Belt (Ceratotrupes fronticornis (Erichson, 1847) and Ceratotrupes bolivari Halffter & Martínez, 1962), and the Sierra Madre del Sur (Ceratotrupes sturmi (Jekel, 1865)). We confirm C. mniszechi (Jekel, 1865) as a junior subjective synonym of C. sturmi. The examination of the holotype of C. sturmi and specimens recently collected in the Sierra Sur de Oaxaca (Oaxaca) and Omiltemi (Guerrero) allowed us to identify and describe a new species, Ceratotrupes gonzaloi sp. nov. from Omiltemi, Guerrero. We redescribe the morphology of C. sturmi and provide a diagnostic key to the four members of the genus Ceratotrupes. Photographs of the habitus and male genitalia of the four species, a distribution map, and observations on their ecology and biogeography are provided.
The railway cutting near Oberrödinghausen at the northern margin of the Rhenish Mountains is the cardinal section for the investigation of Early Tournaisian (Early Carboniferous; Mississippian) ammonoids. The ammonoids from the Hangenberg Limestone (= Gattendorfia Limestone) of this and neighbouring outcrops are revised here, using the historical collections as well as undescribed new material. The ammonoid assemblages are composed of a total of 67 species, which occur in four successive ammonoid zones. The assemblages are composed of predominant prionoceratids (Order Goniatitina) with the twenty genera Mimimitoceras (two species), Globimitoceras (one species), Paragattendorfia (two species), Kornia (three species), Stockumites (eleven species), Acutimitoceras (two species), Costimitoceras (one species), Nicimitoceras (four species), Imitoceras (one species), Voehringerites (one species), Gattendorfia (eight species), Zadelsdorfia (two species), Kazakhstania (one species), Gattenpleura (one species), Weyerella (three species), Hasselbachia (three species), Paprothites (five species), Pseudarietites (three species), Rodingites (two species), Paralytoceras (one species) as well as subordinate eocanitids (Order Prolecanitida) with the genera Eocanites (eight species) and Nomismocanites (one species). The new genera Rodingites gen. nov. and Nomismocanites gen. nov. as well as the new species Mimimitoceras perditum sp. nov., Kornia fibula sp. nov., Kornia acia sp. nov., Stockumites parallelus sp. nov., Stockumites voehringeri sp. nov., Acutimitoceras ucatum sp. nov., Acutimitoceras paracutum sp. nov., Imitoceras initium sp. nov., Gattendorfia rhenana sp. nov., Gattendorfia bella sp. nov., Gattendorfia valdevoluta sp. nov., Gattendorfia schmidti sp. nov., Gattendorfia corpulenta sp. nov., Gattendorfia immodica sp. nov., Zadelsdorfia oblita sp. nov., Weyerella lenis sp. nov., Hasselbachia erronea sp. nov., Paprothites beckeri sp. nov., Paprothites kullmanni sp. nov., Eocanites delicatus sp. nov. and Nomismocanites raritas gen. et sp. nov. are described from Oberrödinghausen. Mimimitoceras mina sp. nov., Stockumites marocensis sp. nov., Zadelsdorfia zana sp. nov. and Kazakhstania kana sp. nov. are newly named for material from the Anti-Atlas of Morocco.
The early Tournaisian (Early Carboniferous; Mississippian) ammonoids from the classical abandoned limestone quarry of Gattendorf (Upper Franconia) are revised, using the historical collections as well as so far undescribed material. The ammonoid assemblage is composed of prionoceratid ammonoids of the six genera Mimimitoceras, Paragattendorfia, Stockumites, Acutimitoceras, Gattendorfia and Gattenpleura, which indicate a stratigraphic position near the Devonian–Carboniferous boundary in the earliest Carboniferous. The new species Stockumites hofensis sp. nov. and S. nonaginta sp. nov. are described.
The number of currently described species of Afrotropical parasitoid wasps does not reflect the true species diversity. One of the most severely understudied parasitoid wasp groups is Ceraphronoidea. In this first study on Afrotropical mainland Ceraphronoidea in more than 20 years, which is also the first ever taxonomic monograph focusing on Ceraphronidae, we describe 88 new species of Ceraphronidae (85 new species) and Megaspilidae (3 new species) from Kakamega Forest (Kenya), Mt. Kilimanjaro (Tanzania) and Ivindo NP (Gabon): Aphanogmus abaluhya sp. nov., A. ashitakai sp. nov., A. idakho sp. nov., A. ikhongamurwi sp. nov., A. isiukhu sp. nov., A. kakamegaensis sp. nov., A. lateritorum sp. nov., A. mangimelii sp. nov., A. mariae sp. nov., A. mashariki sp. nov., A. nehbergi sp. nov., A. njia sp. nov., A. vestrii sp. nov., A. yala sp. nov. (all clavicornis species group), A. dimidiatus sp. nov., A. fraterculus sp. nov., A. guenteri sp. nov., A. kakakili sp. nov., A. kisiwa sp. nov., A. maua sp. nov., A. morriconei sp. nov., A. ndefu sp. nov., A. ngai sp. nov., A. nikii sp. nov., A. pilosicoxa sp. nov., A. rafikii sp. nov., A. robustus sp. nov., A. simbai sp. nov., A. taji sp. nov., A. ukanda sp. nov. (all fumipennis species group), A. campanula sp. nov., A. kikuyu sp. nov., A. pagoda sp. nov. (all tenuicornis species group), Ceraphron banda sp. nov., C. brashi sp. nov., C. breviharpis sp. nov., C. breviscapus sp. nov., C. buyangu sp. nov., C. chemositi sp. nov., C. cingulum sp. nov., C. clavatumeris sp. nov., C. digiti sp. nov., C. eaerendili sp. nov., C. ekero sp. nov., C. ellae sp. nov., C. eulbergi sp. nov., C. herreni sp. nov., C. hitagarciai sp. nov., C. insolitus sp. nov., C. isecheno sp. nov., C. isukha sp. nov., C. ivindoensis sp. nov., C. kaharabu sp. nov., C. kaimosiensis sp. nov., C. kakamegaensis sp. nov., C. kidole sp. nov., C. kimathii sp. nov., C. lirhanda sp. nov., C. longiharpis sp. nov., C. longisetae sp. nov., C. longumerunus sp. nov., C. maathaiae sp. nov., C. malava sp. nov., C. mamamutere sp. nov., C. metapleuralis sp. nov., C. mikoi sp. nov., C. mwekaensis sp. nov., C. nandi sp. nov., C. nzoia sp. nov., C. onesimusi sp. nov., C. pilosiharpis sp. nov., C. pleurosulcus sp. nov., C. reinholdi sp. nov., C. salazar sp. nov., C. sataoi sp. nov., C. semira sp. nov., C. sungura sp. nov., C. tenuimeris sp. nov., C. tiriki sp. nov., C. trietschae sp. nov., Cyoceraphron dhahabudorsalis sp. nov., C. harpe sp. nov., C. invisibilis sp. nov., C. kahawia sp. nov., C. njano sp. nov. (all Ceraphronidae), Conostigmus kijiko sp. nov., C. koleo sp. nov., and Dendrocerus wachagga sp. nov. (all Megaspilidae). In addition, we describe four species of Aphanogmus and five species of Ceraphron without formal naming. A neotype is designated for Dendrocerus anneckei Dessart, 1985 (Megaspilidae). With these new species we more than double the number described from the Afrotropical mainland (65 vs 153). The species numbers found allow us to estimate the real worldwide species number of Ceraphronoidea as being roughly 12 000–21 000, i.e., 16–29 times the number of the currently described species (~730, including the species described herein). This study is meant to highlight that it is necessary and also possible to study the parasitoid wasps of tropical regions and provide momentum for exploring the diversity of small and diverse insect groups in the Afrotropics and elsewhere while also providing the basic knowledge that is much needed for protecting biodiversity and understanding evolution and the networks of life on earth. All described species are diagnosed and illustrated, with focus on the male genitalia. Furthermore, we provide an identification key to males of Afrotropical Ceraphronidae.
We provide a floristic account for the Begoniaceae of Peru. The family is represented in Peru by 76 species, which are all members of the genus Begonia and represent eighteen sections. Twelve new species are described: B. condorensis Jara & Moonlight sp. nov., B. deltoides Moonlight sp. nov., B. huancabambae Moonlight sp. nov., B. imbrexiformis Moonlight sp. nov., B. longinqua Moonlight sp. nov., B. longitepala Moonlight sp. nov., B. nunezii Moonlight sp. nov., B. occultata J.P.Allen & Moonlight sp. nov., B. pedemontana Moonlight sp. nov., B. serratistipula Moonlight sp. nov., B. vargasii Moonlight sp. nov. and B. yuracyacuensis Moonlight sp. nov. We also provide four new records for the country: B. andina Rusby, B. brandbygeana L.B.Sm. & Wassh., B. neoharlingii L.B.Sm. & Wassh. and B. unilateralis Rusby. We provide an identification key to all species. The ecology, distribution, and conservation status of all Peruvian Begonia species are discussed, including provisional IUCN threat assessments. Most species are illustrated by either historical illustrations, contemporary photographic plates, or line drawings. Twenty-six names are newly synonymised including fifteen previously accepted species, and we designate eighty-three lectotypes, four neotypes, and four epitypes.