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This paper presents an improved diagnosis and definition of the genus Stenaelurillus Simon, 1886, with new taxonomic and faunistic data for 23 species. The genera Microheros Wesołowska & Cumming, 1999 and Mashonarus Wesołowska & Cumming, 2002 are synonymized with Stenaelurillus. Six new species are described: Stenaelurillus bandama sp. nov. (♂♀, from Côte d’Ivoire), S. belihuloya sp. nov. (♂, from Sri Lanka), S. jocquei sp. nov. (♂♀, from Cameroon), S. pseudoguttatus sp. nov. (♂, from Namibia), S. senegalensis sp. nov. (♂♀, from Senegal), and Stenaelurillus siyamae sp. nov. (♀, from Sudan). Lectotypes are designated for two species: S. albopunctatus Caporiacco, 1949 (♂) from Kenya and S. werneri Simon, 1906 (♀) from South Sudan. Six new combinations are proposed: Aelurillus ambiguus (Denis, 1966), comb. nov. (ex Stenaelurillus); Evarcha werneri (Simon, 1906), comb. nov. (ex Stenaelurillus); Phlegra davidi (Caleb, Mungkung & Mathai, 2015), comb. nov. (ex Mashonarus); Stenaelurillus brandbergensis (Wesołowska, 2006), comb. nov. (ex Mashonarus); Stenaelurillus guttatus (Wesołowska & Cumming, 2002), comb. nov. (ex Mashonarus); and S. termitophagus (Wesołowska & Cumming, 1999), comb. nov. (ex Microheros). Two species names are synonymized: Evarcha elegans Wesołowska & Russell-Smith, 2000 with E. werneri comb. nov.; and Aelurillus sahariensis Berland & Millot, 1941 with Stenaelurillus nigricaudus Simon
We describe five new species of fungi of the order Laboulbeniales Lindau growing on millipedes and belonging to the genera Diplopodomyces W.Rossi & Balazuc and Troglomyces S.Colla. Three new species of Diplopodomyces, viz. Diplopodomyces coronatus Santam., Enghoff & Reboleira sp. nov. living on Serboiulus spelaeophilus Gulicka, 1967 from Bulgarian caves, Diplopodomyces liguliphorus Santam., Enghoff & Reboleira sp. nov. on an unidentified species of Spirobolida from Sri Lanka, and Diplopodomyces ramosus Santam., Enghoff & Reboleira sp. nov. on Pachyiulus spp. from Turkey, Macedonia and Serbia; and two new species of Troglomyces, viz. Troglomyces dioicus Santam., Enghoff & Reboleira sp. nov. on Nepalmatoiulus sp. from Myanmar, and Troglomyces tetralabiatus Santam., Enghoff & Reboleira sp. nov. on Caucaseuma Strasser, 1970 and Heterocaucaseuma Antić & Makarov, 2016 from caves in Western Caucasus. Troglomyces dioicus sp. nov. is the first dioecious species described in the genus Troglomyces. Keys for all hitherto known species of Diplopodomyces and Troglomyces are presented, as is a discussion of the status of both genera. Additional records for Diplopodomyces lusitanipodos Santam., Enghoff & Reboleira and Troglomyces manfrediae S.Colla are also included.
An unexpected new subgenus and species of Campodeidae (Diplura), Plusiocampa (Pentachaetocampa) inopinata subgen. et sp. nov., a troglobitic species found in Schallsinger Höhle in an isolated karstic region in southwestern Germany is described. The new taxon shows two unique characters for the genus Plusiocampa: five dorsal femoral macrosetae and the presence of g1-glandular setae in females. Two other Plusiocampa species have been studied and taxonomic remarks made for them; both are also cave dwelling species from Germany: Plusiocampa dobati Condé in Dobat, 1975 studied from eight caves in the Swabian Alb, and one unnamed species of Plusiocampa (Plusiocampa) from four caves in the Franconian Alb. The biogeographical and taxonomic affinities among Plusiocampa species of Central Europe are discussed. The distribution of Plusiocampa species in Central Europe runs alongside the frontier of the Pleistocene glaciations, with non-troglomorphic Plusiocampa species adjacent to the glacial limits and troglomorphic Plusiocampa species below. Worthy of note is the presence only in the northeast of the Central Alps of two relict Plusiocampinae species, the already known Hystrichocampa pelletieri Condé, 1948 and the new species P. (P.) inopinata subgen. et sp. nov.
The Charipinae Dalla Torre & Kieffer, 1910 present in the Palaearctic region are revised; 2410 specimens have been identified, belonging to 75 species: 52 to Alloxysta, one to Apocharips, six to Dilyta and 16 to Phaenoglyphis. For 33 species, new country-level distribution records are provided. Two new species are here described: Alloxysta palearctica Ferrer-Suay & Pujade-Villar sp. nov. and Alloxysta pascuali Ferrer-Suay sp. nov. A diagnosis for these species is included and their diagnostic features are shown in different figures. A key to identify all the species of Charipinae in the Palaearctic region is also given.
The Siriella brevicaudata species group from the West Indo-Pacific, defined and designated by Murano & Fukuoka (2008), previously contained five nominal species. In this study we describe five new species in the brevicaudata group: S. bassi sp. nov. from the Bass Strait, southern Australia, S. occulta sp. nov. from the Arabian Gulf, S. muranoi sp. nov. from the coast of Northern Territory, Australia, S. tabaniocula sp. nov. from Ningaloo Reef of Western Australia and Lodestone Reef off Queensland, and S. talbotae sp. nov. from Lizard Island, Queensland, Australia. Furthermore, Siriella hanseni W.M. Tattersall, 1922 from India and S. vincenti W.M. Tattersall, 1927 from South Australia are redescribed based on re-examination of their type material. A re-examination of specimens subsequently attributed to these two species from other geographical regions showed that these were misidentifications, partly representing three of the new species described herein. Siriella gibbosa (Ledoyer, 1970), which was previously synonymized with S. brevicaudata Paulson, 1875 by Bačescu, is revalidated and included within the brevicaudata group. Siriella lacertilis Talbot, 2009, from Lizard Island, is placed within the brevicaudata group. Diagnostic features for all the members of the group and the group itself are updated. As a result of the present study, the brevicaudata group now comprises 12 valid species.
Five species of Batillipes Richters, 1909 were collected from subtidal sediments of the Portuguese coast. Two of them, B. algharbensis sp. nov. and B. lusitanus sp. nov., are new to science. Batillipes algharbensis sp. nov. differs from all the other Batillipes species in having the middle toes 3 on the fourth feet longer than middle toes 4 and by the presence of rounded lateral body projections between legs III and IV. Batillipes lusitanus sp. nov. has the middle toes of the fourth feet equal in length, but it exhibits a dorsal cuticular ornamentation, constituted by large pillars, similar to the cuticle of B. adriaticus Grimaldi de Zio, Morone De Lucia, D’Addabbo Gallo & Grimaldi, 1979 and B. roscoffensis Kristensen, 1978. However, contrary to B. adriaticus, the caudal apparatus of B. lusitanus sp. nov. is a roundish cuticular expansion and B. roscoffensis lacks caudal apparatus. Batillipes adriaticus and B. phreaticus Renaud-Debyser, 1959 are new records for Portugal. Based on the examination of specimens of B. phreaticus collected at the Portuguese coast and their comparison with type material of this species and also of B. littoralis Renaud-Debyser, 1959, the toe arrangement patterns in species of Batillipes are clarified and a new identification key to species of this genus is provided.
The described Afrotropical species of the genus Hexophthalma Karsch, 1879 (under the genus name Sicarius Walckenaer, 1847), of the spider family Sicariidae Keyserling, 1880, were recently reviewed. In the present paper the Afrotropical species of the genus Hexophthalma are revisited. After a thorough examination of all the available specimens from nine major collections, the species H. testacea (Purcell, 1908) is here synonymized with H. hahni (Karsch, 1878), three new species are described – H. binfordae sp. nov., H. goanikontesensis sp. nov. (both from Namibia) and H. leroyi sp. nov. (from South Africa) – and the male of H. dolichocephala (Lawrence, 1928) is described for the first time. The distribution of the species is also revised and a new updated key to the species is compiled.
Homollea Arènes (Rubiaceae, subfamily Ixoroideae, tribe Pavetteae) is a genus of shrubs and small trees endemic to western and northern Madagascar. The genus comprises five species occurring in dry deciduous forest, often in limestone areas. The five species are narrow endemics and their conservation status is either Endangered (4 species) or Critically Endangered (1 species). Homollea is characterized by few-flowered, pseudo-axillary, pedunculate inflorescences, well-developed calyces with the lobes much longer than the tube, laterally flattened seeds with a shallow, elongated to linear hilum and entire endosperm, ovules arising from the upper margin of the placenta, and, pollen grains with supratectal elements in the shape of microgemmae. Until now, three species were known and their descriptions are amended. Two further species, H. furtiva De Block sp. nov. and H. septentrionalis De Block sp. nov., are described as new for science. The five species are dealt with in detail: descriptions, distribution maps, conservation assessments, illustrations, lists of exsiccatae and an identification key are given.
A new monotypic genus of flatid planthoppers (Hemiptera: Fulgoromorpha: Flatidae), Medleria gen. nov., is described for Medleria caudata gen. et sp. nov. (type species) from the island of Socotra (Yemen). Habitus, male and female external and internal genital structures of the new species are illustrated and compared with similar taxa. Medleria caudata gen. et sp. nov. is probably endemic to Socotra where it is known to date from a small area of the Dixam mountain plateau only.
We describe two new species of the annelid genus Trilobodrilus Remane, 1925 (Dinophilidae Verill, 1892) from an intertidal and a subtidal location in San Diego, California. These two species show morphological and molecular divergences between each other and the previously described, geographically distant species. Intertidal T. windansea sp. nov. differs from subtidal T. ellenscrippsae sp. nov. most remarkably in the number and pattern of ciliary tufts and bands on the prostomium and along the body length, besides showing ca. 15% difference in gene fragments of COI and CytB. Trilobodrilus windansea sp. nov., though nesting with T. ellenscrippsae sp. nov. in the molecular phylogenetic analyses, morphologically resembles the Japanese T. itoi Kajihara, Ikoma, Yamasaki & Hiruta, 2015 most closely, but still differs from this species in the higher number of apical ciliary tufts, an additional ciliary row posterior to the second ciliary band, and by lacking a forth ciliary band and segmentally arranged lateral ciliary tufts. Trilobodrilus ellenscrippsae sp. nov. is morphologically most similar to the Japanese T. nipponicus Uchida & Okuda, 1943, but is much shorter, has more apical ciliary tufts, and less regularly arranged lateral ciliary tufts along the body. All species differ significantly in all compared gene fragments, and no obvious correlation was found between habitat and the species morphology or relationships.
The skeleton and the relationships of Libanopycnodus wenzi gen. et sp. nov. and of Sigmapycnodus giganteus gen. et sp. nov., two pycnodontiform fishes from the marine Cenomanian (Late Cretaceous) of Lebanon, are studied in detail. Both species belong to the family Pycnodontidae, as shown by their parietal brush-like process. Some osteological characters (bifid cloacal scale, large well visible dilatator fossa surrounded by the dermosphenotic and the dermopterotic, etc.) indicate that they are members of the subfamily Pycnodontinae. Libanopycnodus gen. nov. has the preopercle much deeper than the exposed region of the hyomandibula-dermohyomandibula and is the most primitive genus of this lineage. Sigmapycnodus gen. nov. is one of the most advanced members of the subfamily. It shares with Oropycnodus Poyato-Ariza & Wenz, 2002 and Pycnodus Agassiz, 1833 an apomorphy not present in the less specialized Pycnodontinae, the arcocentra in hypercomplex contact.
Three new species of Entomobrya Rondani, 1861 from China are described: E. leviseta sp. nov. and E. polychaeta sp. nov. from Shaanxi Province and E. dingi sp. nov. from Yunnan Province. This is the first report of Entomobrya from Shaanxi Province. Entomobrya leviseta sp. nov. is characterised by prelabral smooth chaetae on the labrum; E. polychaeta sp. nov. by three pairs of longitudinal dark blue stripes from Th. II to Abd. III and eight lateral mac on Abd. III; and E. dingi sp. nov. by only a little pigment on the body and 5 central mac on And. II & III. A key to all Chinese species of Entomobrya is given.
The concept of the jumping spider genus Pochytoides Berland & Millot, 1941 is reviewed, based on the examination of described and undescribed species. Pochytoides is elevated from the subgeneric to the generic rank and a short diagnosis and description of the genus are presented. Redescriptions or descriptions of all species are provided together with a key to the species. Two new combinations are proposed: Pochytoides perezi (Berland & Millot, 1941) comb. nov. and P. poissoni (Berland & Millot, 1941) comb. nov. (both from Pochyta). Pochyta remyi Berland & Millot, 1941 originally placed in the subgenus Pochytoides is excluded; new combination Thiratoscirtus remyi (Berland & Millot, 1941) comb. nov. is proposed for it (but its generic status is uncertain). Six new species are described: Pochytoides monticola sp. nov., P. obstipa sp. nov., P. lamottei sp. nov., P. patellaris sp. nov., P. securis sp. nov. and P. spiniger sp. nov. The genus has a West African distribution.
The Palaearctic species of Ismaridae Thomson, 1858 are reviewed. Thirteen species of Ismaridae are recognized from the Palaearctic. Five species are described as new: Ismarus brevis Kim & Lee sp. nov. from the Russian Far East and South Korea; I. distinctus Kim, Notton & Ødegaard sp. nov. from Norway and the United Kingdom; I. excavatus Kim & Lee sp. nov. from China, Japan and South Korea; I. similis Kim, Notton & Lee sp. nov. from the United Kingdom and I. tripotini Kim & Lee sp. nov. from South Korea. Ismarus apicalis Kolyada & Chemyreva, 2016 is newly recorded from China, France, Japan and South Korea; I. dorsiger (Haliday, 1831) from France, Montenegro, Norway, South Korea and Switzerland; I. flavicornis (Thomson, 1858) from Bulgaria and Norway; I. grandis Alekseev, 1978, I. halidayi Förster, 1850 and I. multiporus Kolyada & Chemyreva from Japan and South Korea; I. rugulosus Förster, 1850 from Austria and I. spinalis Kolyada & Chemyreva, 2016 from China, Japan and South Korea. An identification key to all species found in the Palaearctic region is presented.
A new classification of Ophiuroidea, considering family rank and above, is presented. The new family and superfamily taxa in O’Hara et al. (2017) were proposed to ensure a better readability of the new phylogeny but are unavailable under the provisions of the ICZN. Here, the morphological diagnoses to all 33 families and five superfamilies are provided. Ten new families, Ophiosphalmidae fam. nov., Ophiomusaidae fam. nov., Ophiocamacidae fam. nov., Ophiopteridae fam. nov., Clarkcomidae fam. nov., Ophiopezidae fam. nov., Ophiernidae fam. nov., Amphilimnidae fam. nov., Ophiothamnidae fam. nov. and Ophiopholidae fam. nov., are described. The family Ophiobyrsidae Matsumoto, 1915, not yet discovered in the previous publication, is added, based on new molecular data. A new phylogenetic reconstruction is presented. Definitions of difficult-to-apply morphological characters are given.
The genus Camafroneta gen. nov. is established to accomodate Camafroneta oku gen. et sp. nov., a new species of mynoglenine spider from Cameroon, West Africa, with unusual complex male genitalia. Phylogenetic analyses suggest that the new species does not belong to a known genus of mynoglenine spiders. Another three new species of Afroneta Holm, 1968 from East Africa are described: Afroneta flavescens sp. nov., Afroneta serrata sp. nov. and Afroneta sarahae sp. nov. We also describe the missing sex (♂) of Afroneta elgonensis Merrett, 2004.
An update of the current knowledge of Anacharis Dalman, 1823 for the Palaearctic and Indomalayan regions is given. The previously known Palaearctic species Anacharis antennata Belizin, 1951, Anacharis eucharoides (Dalman, 1818), Anacharis immunis Walker, 1835 and Anacharis parapsidalis Belizin, 1951 are redescribed. Three new species are described: Anacharis fergussoni sp. nov. from Europe, Anacharis norvegica sp. nov. from Norway and Anacharis belizini sp. nov. from Thailand, the first recorded Indomalayan species for the genus. Anacharis gracilipes Ionescu, 1969, is synonymized with A. eucharoides, while Anacharis flavidicornis Kieffer, 1910 is transferred to the genus Aegilips Haliday, 1835, resulting in Aegilips flavidicornis (Kieffer, 1910) comb. nov. Diagnostic characters and data about the biology, distribution and affinities with other species of Anacharis are discussed. An identification key for the Palaearctic and Indomalayan species of Anacharis is given.
A new genus and species of owlfly from eastern and southern Africa (Neuroptera: Ascalaphidae)
(2018)
The genus Dorsomitus Tjeder, 1992, is considered a nomen nudum. Dorsomitus gen. nov. is described and validated here. A new combination, Dorsomitus neavei (Kimmins, 1949) gen. et comb. nov. is proposed, Dorsomitus tjederi gen. et sp. nov. is described, and Disparomitus neavei Kimmins, 1949, is designated as type species of the genus Dorsomitus gen. nov.
The recently described family Chummidae, now the sister clade of Macrobuninae, so far only known from two South African species, is extended with seven new species, six from the southern part of South Africa and one from Lesotho: Chumma bicolor sp. nov. (♀), C. foliata sp. nov. (♂♀), C. interfluvialis sp. nov. (♂♀), C. lesotho sp. nov. (♀), C. striata sp. nov. (♂♀), C. subridens sp. nov. (♂♀) and C. tsitsikamma sp. nov. ♂. A key to the species is provided. Although Chumma is part of a clade containing the Macrobuninae, it is argued that the family name Chummidae should remain valid.
The Late Jurassic–Early Cretaceous Slottsmøya Member of the Agardhfjellet Formation in central Spitsbergen has yielded two new species of asteroids and two species of ophiuroids, one of which is described as new. Polarasterias janusensis Rousseau & Gale gen. et sp. nov. is a forcipulatid neoasteroid with elongated arms, small disc and very broad ambulacral grooves with narrow adambulacrals. Savignaster septemtrionalis Rousseau & Gale sp. nov. is a pterasterid with welldeveloped interradial chevrons. The Spitsbergen specimens are the first described articulated material of Savignaster and reveal the overall arrangement of the ambulacral groove ossicles. Ophiogaleus sp. is an ophiacanthid with relatively long jaws and lateral arm plates, with a coarsely reticulate outer surface. Here again, we report the first articulated skeletons of this genus, providing unprecedented insights into the disc morphology. Ophioculina hoybergia Rousseau & Thuy gen. et sp. nov. is an ophiopyrgid with a well-developed arm comb and tentacle pores reduced to within-plate perforations starting at median arm segments. These new finds are important additions to the asterozoan fossil record with regard to their good degree of articulation and the high latitudinal position of the localities. They significantly add to the set of exhaustively known fossil asterozoan taxa which play a key role in the phylogenetic analysis and reconstruction of evolutionary history.