Insecta Mundi
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767
Several taxonomic and nomenclatural issues are reviewed, clarified, and resolved for multiple genera of the Erotylinae (Coleoptera: Erotylidae). Generic-group names discussed: Brachymerus Dejean, 1836, Cypherotylus Crotch, 1873, Cytorea Laporte, 1840, Erotylus Fabricius, 1775, Eudaemonius Lewis, 1887, Eutriplax Lewis,1887, Gibbifer Voet, 1806, Neobarytopus Alvarenga, 1965, Neomorphoides Alvarenga, 1977, Ogcotriplax Heller,1920, Paratritoma Gorham, 1888, Platichna Thomson, 1863, Pseudochrysomela Voet, 1806, Pseudotriplax Heller,1920, Triplax Herbst, 1793, Tritomapara Alvarenga, 1970, Typocephalus Hope,1841, and Xestus Wollaston, 1864. Reviewing these issues resulted in a several nomenclatural actions. Eutriplax Lewis,1887,was found to be an unnecessary replacement name for Eudaemonius Lewis,1887.The genus name is reverted to Eudaemonius,
resulting in one new combination: Eudaemonius quinquepustulatus (Li and Ren, 2006).
The Neotropical Tritomapara Alvarenga,1970,was found to be a new objective synonym of Paratritoma Gorham,1888, which is a synonym of Triplax Herbst,1793,leading to the following eight new combinations:
Triplax atricaudata (Kuhnt,1910),Triplax brasiliensis (Guérin,1946),Triplax bruchi (Kuhnt,1910),Triplax caduca (Gorham,1888),Triplax dimidiata (Gorham,1888),Triplax melanoderes (Kuhnt,1910),Triplax triplacoides
(Crotch,1876), and Triplax vivida (Gorham,1888). Erotylus tibialis Duponchel, 1825, is recognized as the valid type species for Brachymerus Dejean 1836, which moves the name Brachymerus to a different genus-group taxon and renders Neomorphoides Alvarenga, 1977, a new synonym. This revalidates Neobarytopus Alvarenga, 1965, as originally proposed. These genus-group names are presently subgenera in Iphiclus Dejean, 1836, and the move creates 23 new combinations in Iphiclus(Brachymerus) Dejean, 1836: I. (B.) amazonus (Crotch, 1876), I. (B.) atriventris (Mader,1943), I. (B.) bicolor(Lacordaire,1842), I. (B.) clavicornis (Olivier,1792), I. (B.) columbiae (Crotch,1876), I. (B.) costaricensis (Mader,1943), I. (B.) disconigrum (Mader,1942), I. (B.) dorsonotatus (Lacordaire, 1842), I. (B.) fulviventris (Gorham,1888), I. (B.) humeropictus (Mader,1943), I. (B.) lateripunctatus (Crotch,1876), I. (B.) melanopus (Gorham,1888), I. (B.) neglectus (Guérin,1956), I. (B.) nigritarsis (Mader,1942), I. (B.) nigriventris (Crotch, 1876), I. (B.) nigropectus (Mader,1942), I. (B.) posticenigrum (Mader,1942), I. (B.) pyrrhocephalus (Erichson,1847), I. (B.) rubripennis (Lacordaire,1842), I. (B.) signaticollis (Kuhnt,1910), I. (B.) simplex (Lacordaire,1842), I. (B.) spilotus (Gorham,1888), I. (B.) tibialis (Duponchel,1825); and, 75 new combinations in Iphiclus (Neobarytopus) Alvarenga, 1965: I. (N.) adustus (Duponchel,1825), I. (N.) alboniger (Guérin,1956), I. (N.) amictus (Erichson,1847), I. (N.) andicola (Kirsch,1867), I. (N.) assequens (Mader,1942), I. (N.) bajulus (Lacordaire,1842), I. (N.) batesi (Gorham, 1889),I. (N.) bellulus (Lacordaire,1842), I. (N.) bicinctus (Olivier,1807), I. (N.) bistrifoliatus (Gorham,1889), I. (N.) bizonatus (Crotch,1876), I. (N.) bremei (Guérin-Méneville,1841), I. (N.) brongniarti (Lacordaire,1842), I. (N.) brunneostriolatus (Kuhnt, 1910), I. (N.) cerasinus (Lacordaire,1842), I. (N.) conformis (Lacordaire,1842), I. (N.)distinctus (Duponchel, 1825), I. (N.) divisus (Guérin,1956), I. (N.) dorsalis (Olivier,1792), I. (N.) eburneus (Crotch,1876), I. (N.) elegans (Mader,1942), I. (N.) epipleuralis (Crotch,1876), I. (N.) erichsoni (Lacordaire,1842), I. (N.) flavofasciatus (Duponchel,1825), I. (N.) flavosignatus (Duponchel,1825), I. (N.) fragmentatus (Gorham,1888), I.(N.) friedei (Mader,1938), I. (N.) geometra (Lacordaire,1842), I. (N.) hebriacus (Lacordaire,1842), I. (N.) hexastictus (Crotch,1876), I. (N.) incas (Gorham,1889), I. (N.) iris (Guérin,1956), I. (N.) jacinthoi (Alvarenga,1977), I.(N.) laceratus (Mader,1938), I. (N.) lugens (Lacordaire,1842), I. (N.) lunaris (Guérin,1956), I. (N.) luteozonatus (Crotch, 1876), I. (N.) miles (Mader,1942), I. (N.) mirus (Mader,1942), I. (N.) musicalis (Lacordaire,1842), I. (N.) neophyta (Lacordaire, 1842), I. (N.) nigripennis (Demay,1838), I. (N.) nigropictus (Lacordaire,1842), I. (N.) nigrofasciatus (Mader,1942), I. (N.) nitidulus (Oliver,1807), I. (N.) obsoletesignatus (Crotch, 1876), I. (N.) octoguttatus (Olivier,1807), I. (N.) octopustulatus (Guérin,1956), I. (N.) odyneroides (Crotch, 1876), I. (N.) ornatus (Kuhnt,1909), I. (N.) pantherinus (Kuhnt, 1909), I. (N.) pauper (Guérin,1956), I. (N.) peraffinis (Crotch,1876), I. (N.) perplexus (Mader,1942), I. (N.) peruvianus (Mader,1942), I. (N.) planipennis (Kuhnt,1909), I. (N.) puncticollis (Kirsch,1876), I. (N.) quadrifasciatus (Kirsch,1865), I. (N.) quinquefasciatus (Lacordaire,1842), I. (N.) ramosus (Olivier,1807), I. (N.) regularis (Erichson, 1848), I. (N.) rhomboidalis (Guérin,1956), I. (N.) rufipennis (Panzer,1798), I. (N.) salamandra (Erichson,1847), I. (N.) spectabilis (Lacordaire,1842), I. (N.) stramineus (Lacordaire,1842), I. (N.) subsanguineus (Crotch,1876), I. (N.) superbus (Mader,1942), I. (N.) tigrinatus (Guérin,1956), I. (N.) tricinctus (Duponchel,1825), I. (N.) trifasciatus (Olivier,1807), I. (N.) tripartitus (Lacordaire,1842), I. (N.) ucayalensis (Gorham,1889), I. (N.) venezuelae (Crotch, 1876), I. (N.) westwoodi (Guérin-Méneville,1841).
The works of Voet (1766–1778,1806) do not follow binominal nomenclature and are therefore unavailable by the International Code of Zoological Nomenclature, Article 11.4.Thus, Voet’s (1806) generic names “Pseudochrysomela” and “Gibbifer”, and the species names proposed in each, are unavailable. Removing them from nomenclatural considerations resulted in the following nomenclatural acts: the resurrection of Erotylus rufipennis Panzer,1798, now Iphiclus (Neobarytopus) rufipennis (Panzer) new combination; the proposal of a new name, Iphiclus (Brachymerus) fabricii Skelley for Erotylus rufipennis Fabricius, 1801, not Erotylus rufipennis Panzer, 1798; new combinations for the two species, Cypherotylus adrianae (Alvarenga, 1976) and Cypherotylus borgmeieri (Alvarenga, 1976); and revalidated status for the five species names, Erotylus variegatus Fabricius,
1781, Barytopus gronovii (Herbst, 1783), Prepopharus notatus (Olivier, 1792), Iphiclus (Iphiclus) sedecimguttatus (Olivier, 1792), and Cypherotylus duponcheli Arrow, 1937.
765
The tribal allocation of Xalitla Lane, 1959 (Cerambycidae: Cerambycinae) is discussed. Xalitla limoni Santos-Silva and Skillman, new species,is described from Mexico (Jalisco). Xalitla lezamai Galileo and Martins,2008 is determined to be synonymous with X. genuina Martins,1970 and formally placed in synonymy. A key to the species of Xalitla, which includes the new species and synonymy, is provided.
760
Crioprosopus baldwini Eya (Coleoptera: Cerambycidae: Cerambycinae: Trachyderini), sp. nov., from Costa Rica and Panama, is described. An amendment to the key to species of Crioprosopus Audinet-Serville, 1834, as presented in Eya (2015), is provided along with illustrations of the key characteristics to differentiate C. baldwini from other species.
754
Two new genera of Acanthocinini (Coleoptera: Cerambycidae), Luctithonus Lingafelter and Duocristala Lingafelter, are described from Hispaniola. Two new species of Luctithonus are described: Luctithonus aski Lingafelter and L. duartensis Lingafelter. A third species, L. pantherinus (Zayas), is newly recorded from Hispaniola and the Dominican Republic (new country record), and transferred from Sternidius Haldeman as a new combination. Additional new species of Lamiinae are described from Hispaniola: Eugamandus albipumilus Lingafelter; Leptostylopsis opuntiae Lingafelter; and Lethes turnbowi Lingafelter. Keys to tribes of Lamiinae, genera of Acanthocinini, and species of Luctithonus in Hispaniola are included.
752
748
The following new species of Eupogonius LeConte, 1852 (Coleoptera: Cerambycidae: Lamiinae) are described: E. tlanchinolensis Wappes and Santos-Silva (Mexico, Hidalgo); E. albofasciatus Wappes and Santos- Silva (Mexico, Puebla); E. sonorensis Wappes and Santos-Silva (Mexico, Sonora); E. guerrerensis Wappes and Santos-Silva (Mexico, Guerrero); E. boteroi Wappes and Santos-Silva (Mexico, Guerrero); E. nascimentoi Wappes and Santo-Silva (Mexico, Jalisco and Colima); and E. monzoni Wappes and Santos-Silva (Guatemala, Alta Verapaz). Additionally, a detailed description of the female of Eupogonius fulvovestitus Schaeffer, 1905 is provided for the first time, along with notes on the likely host of the species. New state records in Mexico are provided for Eupogonius comus Bates, 1885, and E. stellatus Chemsak and Noguera, 1995. Other taxonomic or nomenclatural actions included herein are: Eupogonius knabi Fisher, 1925 is transferred to Atelodesmis Chevrolat, 1841, new combination; the gender of the species-group name in Eupogonius azteca Martins, Santos-Silva and Galileo, 2015 is commented on; notes on the geographical distribution of Eupogonius affinis Breuning, 1942, and the problematic morphology of E. infimus (Thomson, 1868) are presented; Eupogonius subaeneus Bates, 1872, and E. marmoratus Fisher, 1925 are revalidated, and E. columbianus Breuning, 1942 is a new synonym of E. subaeneus”.
742
In the course of working on new species of North American Phyllophaga Harris, 1827 (Coleoptera: Scarabaeidae: Melolonthinae) some synonyms have been found and are proposed here. New synonymies: Phyllophaga knausii (Schaeffer, 1907) is synonymized with Phyllophaga sociata (Horn, 1878); Phyllophaga chippewa Saylor, 1939 is synonymized with Phyllophaga rugosa (Melsheimer, 1845); and Phyllophaga falta Sanderson, 1950 is synonymized with Phyllophaga bipartita (Horn, 1887). Lectotypes are here designated for the following species: Listrochelus knausii Schaeffer, Listrochelus sociatus Horn, and Lachnosterna bipartita Horn. A neotype for Ancylonycha rugosa Melsheimer is here designated from the Horn Collection.
741
Seven new species of Cacostola Fairmaire and Germain, 1859 (Coleoptera: Cerambycidae: Lamiinae: Onciderini) are described: C. plotkini Wappes and Santos-Silva from Costa Rica; C. galenae Wappes and Santos- Silva from Panama; C. opitzi Wappes and Santos-Silva from Bolivia; C. howdenae Wappes and Santos-Silva from Guatemala; C. thomasorum Wappes and Santos-Silva from Brazil; C. nearnsi Wappes and Santos-Silva from Bolivia and Panama; and C. rothschildi Wappes and Santos-Silva from Argentina and Brazil. Notes on C. simplex (Pascoe, 1859), and C. mexicana (Breuning, 1943) are also included.
737
736
Systematic, faunistic and ecological aspects of the six families and 34 species and subspecies in the order Ephemeroptera currently recorded from Cuba are reviewed based primarily on a reference collection located at the Universidad de Oriente (Santiago de Cuba), collections at the Institute of Ecology and Systematics (Havana) and historic literature. A key to nymphs is included with photographs of significant features of many species. An annotated list of species is presented with comments on type localities, species ecology and distribution. The morpho- ecological types of the nymphs are updated according to current taxonomic changes, and indicator species of organic contamination are analyzed according to the BMWP-Cub index. Based on present data, mayflies are best collected between January and June although many species are present throughout the year, and almost half of the species are widely distributed. Possible routes of penetration from the continents toward Cuba are from South America through the arc of islands formed by the Lesser Antilles, from Central and South America through the peninsula of Yucatan, and via an ancient landspan or island chain from northern South America (GAARlandia). With one exception, there is no evidence for dispersal of species from North America (through Florida) to Cuba (and then to the Antilles) or vice versa. The pattern of geographical distribution of Ephemeroptera inside Cuba is very similar to that of the orders Trichoptera and Odonata. The greatest number of species is found in the Eastern region and the fewest in the Central and Central-East regions. The high endemism (76.5%) is probably due to geographical isolation and processes that bring about this phenomenon together with the low vagility that characterizes the order.
735
731
Genomic sequencing and analysis of worldwide skipper butterfly (Lepidoptera: Hesperiidae) fauna points to imperfections in their current classification. Some tribes, subtribes and genera as they are circumscribed today are not monophyletic. Rationalizing genomic results from the perspective of phenotypic characters suggests two new tribes, two new subtribes and 50 new genera that are named here: Ceratrichiini Grishin, trib. n., Gretnini Grishin, trib. n., Falgina Grishin, subtr. n., Apaustina Grishin, subtr. n., Flattoides Grishin, gen. n., Aurivittia Grishin, gen. n., Viuria Grishin, gen. n., Clytius Grishin, gen. n., Incisus Grishin, gen. n., Perus Grishin, gen. n., Livida Grishin, gen. n., Festivia Grishin, gen. n., Hoodus Grishin, gen. n., Anaxas Grishin, gen. n., Chiothion Grishin, gen. n., Crenda Grishin, gen. n., Santa Grishin, gen. n., Canesia Grishin, gen. n., Bralus Grishin, gen. n., Ladda Grishin, gen. n., Willema Grishin, gen. n., Argemma Grishin, gen. n., Nervia Grishin, gen. n., Dotta Grishin, gen. n., Lissia Grishin, gen. n., Xanthonymus Grishin, gen. n., Cerba Grishin, gen. n., Avestia Grishin, gen. n., Zetka Grishin, gen. n., Turmosa Grishin, gen. n., Mielkeus Grishin, gen. n., Coolus Grishin, gen. n., Daron Grishin, gen. n., Barrolla Grishin, gen. n., Brownus Grishin, gen. n., Tava Grishin, gen. n., Rigga Grishin, gen. n., Haza Grishin, gen. n., Dubia Grishin, gen. n., Pares Grishin, gen. n., Chitta Grishin, gen. n., Artonia Grishin, gen. n., Lurida Grishin, gen. n., Corra Grishin, gen. n., Fidius Grishin, gen. n., Veadda Grishin, gen. n., Tricrista Grishin, gen. n., Viridina Grishin, gen. n., Alychna Grishin, gen. n., Ralis Grishin, gen. n., Testia Grishin, gen. n., Buzella Grishin, gen. n., Vernia Grishin, gen. n., and Lon Grishin, gen. n. In addition, the following taxonomic changes are suggested. Prada Evans is transferred from Hesperiinae to Trapezitinae. Echelatus Godman and Salvin, Systaspes Weeks, and Oenides Mabille are removed from synonymy and are treated as valid genera. The following genera are new junior subjective synonyms: Tosta Evans of Eantis Boisduval; Turmada Evans of Neoxeniades Hayward, Arita Evans of Tigasis Godman, and Alera Mabille of Perichares Scudder. Eantis pallida (R. Felder) (not Achlyodes Hübner), Gindanes kelso (Evans) (not Onenses Godman and Salvin), Isoteinon abjecta (Snellen) (not Astictopterus C. and R. Felder), Neoxeniades ethoda (Hewitson) (not Xeniades Godman), Moeris anna (Mabille) (not Vidius Evans), and Molo pelta Evans (not Lychnuchus Hübner) are new genus-species combinations. The following are species-level taxa: Livida assecla (Mabille) (not a subspecies of Livida grandis (Mabille), formerly Pythonides Hübner) and Alychna zenus (E. Bell) (not a junior subjective synonym of Alychna exclamationis (Mabille), formerly Psoralis Mabille); and Barrolla molla E. Bell (formerly Vacerra Godman) is a junior subjective synonym of Barrolla barroni Evans (formerly Paratrytone Godman). All these changes to taxonomic status of names are propagated to all names currently treated as subspecies (for species), subgenera (for genera) and synonyms of these taxa. Finally, taxa not mentioned in this work are considered to remain at the ranks and in taxonomic groups they have been previously assigned to.
728
A new species of the genus Bomansius Lacroix (Coleoptera: Lucanidae), which had been illustrated in the literature but never formally named, is described from Aneityum Island, Vanuatu (former New Hebrides) under the name of Bomansius cheesmanae Kakinuma, new species. The generic redescription of Bomansius and the redescription of B. gabrieli Lacroix, 1978 are also given based on additional specimens. Bomansius is transferred to tribe Aegini Huang and Chen, 2013.
718
717
716
Nomenclatural and taxonomic changes are proposed for American Apomecynini (Coleoptera: Cerambycidae: Lamiinae). New synonymies: Parmenonta valida Thomson, 1868, type species of Parmenonta Thomson, 1868, is transferred to Adetus LeConte, 1852, making obligatory the synonymy of Parmenonta with Adetus; Adetus cylindricus Bates, 1866 = A. inaequalis (Thomson, 1868); Adetus leucostigma Bates, 1880 = Adetus binotatus (Thomson, 1868); Adetus tuberosus Galileo and Martins, 2003 = Typophaula melancholica Thomson, 1868; Adetus latericius Belon, 1902, and Adetus irregularis (Breuning, 1939) = Adetus nanus (Fairmaire and Germain, 1859). New records: Adetus binotatus for Chiapas, Guerrero, Tamaulipas, and Quintana Roo (Mexico), new state records; A. inaequalis for Amapá (Brazil), new state record; A. punctatus for Brazil (Rondônia), and Suriname, new country records; Adetus bacillarius Bates, 1885 for Mexico, new country record, and Brazil, new country record; Adetus insularis Breuning, 1940 for Mexico, new country record; Adetus nanus, for Brazil (Pará), new state record, Colombia and Venezuela, new country records. Adetus validus (Thomson, 1868) comb. nov. (from Parmenonta Thomson, 1868). New genera and new species: Adetus x-fasciatus Santos-Silva, Nascimento and Wappes, from Paraguay and Argentina; Adetus monteverdensis Santos-Silva, Nascimento and Wappes, from Costa Rica; Adetus pseudobacillarius Santos-Silva, Nascimento and Wappes, from Costa Rica; Adetus clinei Santos- Silva, Nascimento and Wappes, from Bolivia; Adetaptera Santos-Silva, Nascimento and Wappes, new genus, with A. albisetosus (Bates, 1880) comb. nov. designated as type species; Adetaptera schaffneri Santos-Silva, Nascimento and Wappes, from Mexico; Morrisia Santos-Silva, Nascimento and Wappes, new genus, for M. squamosa (Chemsak and Noguera, 1995) comb. nov., transferred from Adetus LeConte, 1852 and designated as type species, and M. pulchra Santos-Silva, Nascimento and Wappes, from Mexico; Skillmania Santos-Silva, Nascimento and Wappes, new genus, with S. obrienorum Santos-Silva, Nascimento and Wappes comb. nov., from Mexico, designated as type species. New combinations: The following 15 species are transferred from Parmenonta to Adetaptera: A. albosticta (Galileo and Martins, 2003), A. chapadensis (Martins and Galileo, 1999), A. fulvosticta (Bates, 1885), A. insularis (Fisher, 1930), A. laevepunctata (Breuning, 1940), A. lenticula (Galileo and Martins, 2006), A. maculata (Martins and Galileo, 1999), A. minor (Bates, 1880), A. ovatula (Bates, 1880), A. parallela (Lameere, 1893), A. punctigera (Germar, 1823), A. strandiella (Breuning, 1940), A. thomasi (Linsley and Chemsak, 1985), A. wickhami (Schaeffer, 1908), and A. dominicana (Galileo and Martins, 2004).
714
Genera of Cryptognathini (Coleoptera: Coccinellidae) are discussed and a key to all recognized genera is provided. Cryptognatha is revised, and species of this genus are keyed. New species, authored by González and Hanley, are Cryptognatha pam, C. kellie, C. hannah, C. whitney, C. karla, C. celia, C. shelia, C. gayle, C. della and C. vicki. The following new synonymies are proposed: Cryptognatha simillima Sicard = Cryptognatha gemellata Mulsant, Cryptognatha fryii Crotch = Cryptognatha pudibunda Mulsant, Cryptognatha bryanti Brèthes = Cryptognatha pudibunda Mulsant. Lectotypes are here designated for Cryptognatha amicta Gorham, C. weisei Brèthes, C. pudibunda Mulsant and C. fryii Crotch.
703
Four new species in the genus Amphicnaeia (Coleoptera: Cerambycidae: Lamiinae) are described: A. panamensis Wappes, Santos-Silva and Galileo and A. fuscofasciata Wappes, Santos-Silva and Galileo from Panama; A. bezarki Wappes, Santos-Silva and Galileo from Venezuela; and A. rileyi Wappes, Santos-Silva and Galileo from Costa Rica and Panama. Amphicnaeia affinis Breuning, 1940 is placed in synonymy with A. lineata Bates, 1866, and the species newly recorded from the Brazilian state of Minas Gerais. Amphicnaeia cordigera Aurivillius, 1920 is transferred to Rosalba Thomson, 1864, resulting in a new combination, and Rosalba rufescens Breuning, 1940, is found to be a junior synonym of the former. The holotypes of A. vitticollis Breuning, 1940, and A. villosula (Thomson, 1868) are illustrated for the first time.
701
A new genus and new species of Onciderini Thomson, 1860 (Coleoptera: Cerambycidae: Lamiinae) are described and illustrated: Velozideres buntyni, from Ecuador. The following two new synonymies in Onciderini are proposed: Oncideres paurosoma Noguera, 1993 = Microcanus minor (Bates, 1885); Oncioderes piauiensis Martins and Galileo, 2013 = Oncioderes rondoniae Martins and Galileo, 1990. The following seven new country records are reported: Clavidesmus chicae Giorgi, 1998 (Onciderini) (Argentina); Ecthoea quadricornis (Olivier, 1795) (Onciderini) (Venezuela); Oncideres ophthalmalis Dillon and Dillon, 1946 (Onciderini) (Costa Rica); Oncideres punctata Dillon and Dillon, 1946 (Onciderini) (El Salvador); Oncideres xavieri Galileo and Martins, 2010 (Onciderini) (Peru); Trestonia signifera Buquet, 1859 (Onciderini) (Brazil); and Oideterus crenatocerus (Galileo, 1987) (Cerambycidae: Prioninae: Anacolini) (Costa Rica).
699
Two new species of Onciderini Thomson, 1860 (Coleoptera: Cerambycidae: Lamiinae) are described and illustrated: Neocherentes adrianoi Nearns and Monné, from Brazil, and Neocherentes pergeri Nearns and Monné, from Bolivia. The male of Neocherentes dilloniorum Tippmann, 1960 is redescribed and the female is described for the first time. Neocherentes dilloniorum is excluded from the Brazilian fauna. A key to the known species of Neocherentes Tippmann, 1960 is provided.