Insecta Mundi
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Institute
- Extern (65)
0150
An additional 137 species and two tribes are added to the cerambycid fauna of Bolivia while 12 species are deleted. This brings the total number of species known from Bolivia to 1,561. Comments and statistics regarding the growth of knowledge on the Bolivian Cerambycid fauna and species endemicity are included.
0202
The vespid fauna of Greater Puerto Rico is reviewed (Hymenoptera: Vespidae). Three new species are described, Ancistrocerus isla Carpenter, Euodynerus jeitita Carpenter and Genaro, and Omicron aridum Carpenter and Genaro. Polistes crinitus americanus (Fabricius, 1775) and P. crinitus multicolor (Olivier, 1792) are both reduced to synonyms of nominotypical P. crinitus (Felton, 1765), revised status; Zeta abdominale hispaniolae (Bequaert and Salt, 1931) and Zeta abdominale ornatum (de Saussure, 1855) are both reduced to synonyms of nominotypical Zeta abdominale (Drury, 1770), revised status; and Zethus rufinodus monensis Bohart and Stange, 1965, and Zethus rufinodus virginicus Bohart and Stange, 1965, are both reduced to synonyms of nominotypical
Zethus rufinodus (Latreille, 1806), revised status. Parancistrocerus obliquus (Cresson, 1865) is newly recorded
from Puerto Rico. The presence of Pachodynerus guadulpensis (de Saussure, 1853) in Puerto Rico is confirmed. An analysis of the composition of the Puerto Rican vespid fauna is presented.
0161
0261
Collection methods and/or habitats sampled influence how many and which species are captured during entomological surveys. Here we compare Coleoptera catches among three survey activities, each using a single collection method, at the same study sites in Great Smoky Mountains National Park, USA. Activities included: short-term flight intercept trapping (FITs); sifting/Berlese funneling of leaf litter and extremely decayed downed coarse woody debris; and using emergence chambers containing coarse woody debris of various decay classes. In total, 2472 adult beetle specimens, representing 217 lowest identifiable taxa within 164 genera and 42 families, were collected during the FIT survey. Each survey activity yielded more than 2000 specimens, and a combined total of 413 species was collected. A combination of all surveys yielded the highest species richness when normalized for number of specimens indicating that variation of habitat and/or collection method significantly increases species richness. Of single surveys the FIT survey had the highest absolute species richness (217) and the highest richness when normalized for number of specimens. Species overlap among survey activities was low (Sorensen’s quotient of similarity was 0.20–0.27), which showed that each was about equally dissimilar from all others. Overlap of catch between FITs and emergence chambers was too low to justify substitution of emergence surveys with the FIT survey protocol used when attempting to collect saproxylic Coleoptera.
0236
Recent fieldwork on North Andros Island by the authors resulted in the collection of six species of Pterophoridae
(Lepidoptera), five of which were previously unrecorded for the Bahamas in published accounts. Three
additional species are noted for the Bahamian fauna based on specimens collected in the 1980s on other islands.
Representative specimens are illustrated from North Andros along with genitalic images for species where these
are not readily available in other publications. In addition, images of the larva and pupa are provided for a reared
species for which the life history was previously unknown.
0238
Snoqualmia, new genus, is described for two species of polydesmid millipeds from the northwestern
United States: Snoqualmia snoqualmie, new species, from Washington State, and S. idaho, new species,
from Idaho. Males of S. idaho possess unusually complex gonopods, perhaps the most complex to be found in the Order
Polydesmida. Snoqualmia is placed in context with other polydesmid genera known from North America. The
polydesmid fauna of North America is discussed, as well as characters of the gonopods of the family.
0235
0217
The taxonomic position of Onthophagus (Palaeonthophagus) lemuroides d’Orbigny, 1898 and Onthophagus
(Palaeonthophagus) fortigibber Reitter, 1909 is discussed (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini).
A key to the species is given. Photos of type specimens of the two taxa and significant chromatic varieties, and
drawings of aedeagi are presented.
0228
A taxonomic review of the ancora species group of Graphipterus Latreille (Coleoptera: Carabidae)
(2012)
The taxonomy of the ancora species group of Graphipterus Latreille (Coleoptera: Carabidae) is reviewed and seven species are recognized, all from southern Africa: Graphipterus ancora Dejean, Graphipterus cordiger Dejean, Graphipterus distinctus Péringuey (new status), Graphipterus fasciatus Chaudoir, Graphipterus fritschi Chaudoir, Graphipterus wahlbergi Boheman (new status), and Graphipterus westwoodi Brême (new status).
Diagnostic features are provided for each species and adult specimens of each species are illustrated.
0226
0225
Two new species of the genus Pahamunaya Schmid (Trichoptera: Polycentropodidae), P. talon sp. n. and P. spinifera sp. n., from Vietnam are described and illustrated. Examination of the holotype male of P. khoii Oláh and Johanson, in combination with an additional specimen of the same species, revealed new characters. New illustrations for this species are provided.
0224
0222
Significant new host and distribution records are presented for Hylesinus mexicanus (Wood) (Coleoptera: Curculionidae: Scolytinae), including its pest potential on cultivated olives. Hylesinus mexicanus is similar to H. fasciatus LeConte and information presented here suggests that the distinctiveness of these two species needs to be re-examined.
0229
Currently, the genus Chimarra Stephens (Trichoptera: Philopotamidae) is represented in the Oriental Region by 259 species. Of these, 61 species have been described or recorded from Vietnam. In this paper, 9 new species from Vietnam are described and illustrated (Chimarra aculeata, C. carinata, C. corneola, C. insolita, C. mina, C. prominens, C. rostrata, C. undulata, and C. ungula). In addition, 3 new country records are noted (Chimarra areli Malicky and Mey, Chimarra pipake Malicky and Chantaramongkol, and Chimarra suthepensis Chantaramongkol and Malicky), and 1 new species group (minuta Group) is proposed and populated. An additional species group (georgensis “Group”), with 1 new species from Vietnam, but otherwise only known from Africa, is discussed, but not formally defined. A table listing all known Vietnamese species of Chimarra is included, along with discussion of variability in the anal veins of the forewing found within this genus, and its relevance for defining subgenera and species groups.
0239
The diplopod orders Callipodida and Polydesmida, and their respective families Abacionidae and
Xystodesmidae, are initially recorded from South Dakota as is Polydesmidae from North Dakota. Other new records of
indigenous taxa include Abacion Rafinesque, 1820/A. texense (Loomis, 1937) and Pleuroloma/P. flavipes, both by
Rafinesque, 1820, from South Dakota, and Pseudopolydesmus Attems, 1898/P. serratus (Say, 1821) from Alabama,
Connecticut, Delaware, New Hampshire, North Dakota, South Carolina, and the District of Columbia. New records of
Aniulus garius Chamberlin, 1912, A. (Hakiulus) d. diversifrons (Wood, 1867), and Oriulus venustus (Wood, 1864)
(Julida: Parajulidae) are provided for western Minnesota and/or eastern North Dakota. Published records from these
states are summarized, and the introduced taxa, Julidae/Cylindroiulus Verhoeff, 1894/C. caeruleocinctus (Wood, 1864)
and Paradoxosomatidae/Oxidus Cook, 1911/O. gracilis (C. L. Koch, 1847), are newly recorded from the Dakotas. The
distribution of P. serratus, which extends from Maine to South Carolina and the Florida panhandle, west to Texas, and
north to Fargo, North Dakota is described and discussed. This distribution exhibits a prominent southeastern lacuna
which we hypothesize suggests replacement by younger, more successful species, as postulated for a similar distributional
gap in Scytonotus granulatus (Say, 1821).
0221
The chilopod, Cryptops hortensis (Donovan, 1810) (Scolopendromorpha: Cryptopidae), and the diplopods, Pseudospirobolellus avernus (Butler, 1876) (Spirobolida: Pseudospirobolellidae) and Oxidus gracilis (C. L. Koch, 1847) (Polydesmida: Paradoxosomatidae), are newly recorded from Saba Island, Lesser Antilles, which also harbors one additional scolopendromorph and four more chilognath millipeds. Except for the plausibly native scolopendrid centipede, Scolopendra alternans Leach, 1813, all are human introductions. Concentrated sampling is needed in the cloud/elfin forest atop Mt. Scenery, where indigenous millipeds may reside, and with extraction techniques throughout the island, to potentially document the diplopod subclass Penicillata. Nine small Caribbean islands in addition to Saba have been incorrectly reported as lacking diplopod records because publications citing them were overlooked by past authors. Works documenting myriapods from small Caribbean islands are consolidated.
0245
Based on two “uni-ocellate” females, the world’s first introductions of the milliped order Stemmiulida are recorded from Florida, United States (US). One individual was collected in 1976 in Gainesville, Alachua County (Co.)., in northcentral peninsular Florida, and the other was taken in 1991 some 408 km (255 mi) to the south-southeast in Pompano Beach, Broward Co. The absence of further individuals and additional samples suggests that the introductions did not result in viable populations, and stemmiulidans are not presently established in the state; the Gainesville site was reinvestigated in 2012 without finding additional specimens. New records from Mexico include the first from Chiapas, Oaxaca, Tabasco, Yucatan, San Luis Potosí, and Tamaulipas states, with the northernmost ordinal locality now becoming Rancho del Cielo, northwest of Gómez Farias, in the last. A northward range expansion of about 460 km (288 mi) from the previous limit, Xalapa, Veracruz, the site lies a mere 40 km (25 mi) south of the Tropic of Cancer and only some 320 km (200 mi) south of the Rio Grande and the US border at McAllen, Hidalgo Co., Texas. Indigenous Stemmiulida are not expected in the forested Rio Grande Valley of southernmost Texas, but their occurrence in the adjoining Mexican state renders such a discovery more plausible than before.
0207
The milliped genus Euryurus Koch, 1847, and the species, E. leachii (Gray, 1832) (Polydesmida: Euryuridae), are recorded from three sites on the northern part of Crowley’s Ridge (Cross, Lee, and Poinsett counties), Arkansas, where the only prior familial records are of Auturus evides (Bollman, 1887). Coupled with the published locality of E. leachii in Phillips Co., at the southern extremity of the Ridge, the only known occurrences of both the genus and species in Arkansas and west of the Mississippi River are in this physiographic feature. The Arkansas population is geographically peripheral but anatomically intermediate between the two recognized subspecies, E. l. leachii and E. l. fraternus Hoffman, 1978, and we do not assign it to a race. Molecular investigations seem necessary to resolve relationships in the “E. leachii complex.”
0234
With the discovery of Mitocybe auriportae Cook and Loomis, 1928 (Platydesmida: Andrognathidae) in Alameda County (Co.), east of San Francisco Bay, a potential overall distribution in coastal California is projected based on those of partly congruent diplopods. The area extends from northern Mendocino to central Monterey cos. and inland to central Lake, Yolo, and Santa Clara cos.
0210
Evidence is presented that the subspecies Chrysobothris thoracica guadeloupensis Descarpentries, 1981
(Coleoptera: Buprestidae) should be recognized at the species level. Character evidence is provided to separate C.
guadeloupensis, new status, from C. thoracica Fabricius, 1798. Both species are illustrated with habitus photographs
and images of the male genitalia.
0243
A new species of Culcua Walker (Diptera: Stratiomyidae), C. lingafelteri Woodley, new species, is described from northern Vietnam. It is diagnosed relative to other species using the recent revision of the genus by Rozkošný and Kozánek (2007). This is the first species of Culcua reported from Vietnam.
0231
Touroultia, a new genus of Onciderini Thomson, 1860 (Coleoptera: Cerambycidae: Lamiinae) is described and illustrated. Five new species of Onciderini are also described and illustrated: Jamesia ramirezi from Costa Rica; Peritrox marcelae from French Guiana; Touroultia swifti from Ecuador; Touroultia lordi from French Guiana; Trestoncideres santossilvai from Brazil. Keys to the known species of Peritrox Bates, 1865; Touroultia gen. nov.; and Trestoncideres Martins and Galileo, 1990 are provided. The following new synonymies are proposed: Calliphenges Waterhouse, 1880 (Colobotheini) = Malthonea Thomson, 1864 (Desmiphorini); Paraclytemnestra Breuning, 1974 (Onciderini) = Jamesia Jekel, 1861 (Onciderini); Orteguaza Lane, 1958 (Apomecynini) = Clavidesmus Dillon and Dillon, 1946 (Onciderini). The following new combinations are proposed: Clavidesmus funerarius (Lane, 1958) (Onciderini); Clavidesmus lichenigerus (Lane, 1958) (Onciderini); Ischiocentra insulata (Rodrigues and Mermudes, 2011); Malthonea cuprascens (Waterhouse, 1880) (Desmiphorini); Touroultia obscurella (Bates, 1865) (Onciderini). The following species is restored to original combination: Jamesia lineata Fisher, 1926 (Onciderini). The following 13 new country records are reported: Ataxia hovorei Lingafelter and Nearns, 2007 (Pteropliini) (Haiti); Carterica soror Belon, 1896 (Colobotheini) (Ecuador); Colobothea lunulata Lucas, 1859 (Colobotheini) (Colombia); Curius punctatus (Fisher, 1932) (Curiini) (Haiti); Cyclopeplus lacordairei Thomson, 1868 (Anisocerini) (Colombia); Iarucanga mimica (Bates, 1866) (Hemilophini) (Ecuador); Pirangoclytus latithorax (Martins and Galileo, 2008) (Clytini) (Costa Rica); Porangonycha princeps (Bates, 1872) (Hemilophini) (Colombia); Trestonia lateapicata Martins and Galileo, 2010 (Onciderini) (Brazil); Tulcus dimidiatus (Bates, 1865) (Onciderini) (Colombia); Unaporanga cincta Martins and Galileo, 2007 (Hemilophini) (Colombia); Zeale dubia Galileo and Martins, 1997 (Hemilophini) (Colombia); Zonotylus interruptus (Olivier, 1790) (Trachyderini) (Colombia).
0250
Larvae of thirty one species of antlions (Neuroptera: Myrmeleontidae) belonging to eleven genera live in the protection of cave mouths or large rock overhangs in Australia. New taxa proposed here include the following three new genera: Australeon, Newleon, Speleon. The following twelve new species are described: Froggattisca kakadu; Froggattisca rennerensis; Newleon fragilis; Speleon cavernicolus; Speleon pilliga; Speleon yallingup; Stenoleon xanthopsis; Xantholeon cavernicolus; Xantholeon kakadu; Xantholeon newi; Xantholeon pallens; Xantholeon pentlandensis. Two cave species are transferred into a new genus becoming new combinations: Australeon illustris (Gerstaecker), Australeon manselli (New and Matsura). The small non-cave species previously placed in Stenoleon Tillyard are transferred to Bandidus Navás, becoming B. gradostriatus (New), B. copleyensis (New), B. grandithecus (New), and B. navasi (New), new combinations. All known species of six genera (Stenoleon Tillyard, Xantholeon Tillyard, Eophanes Banks and three new genera) are known only in cave mouths. Also, species of Heoclisis Navás, Froggattisca Esben Petersen, Glenoleon Banks, Heoclisis Navás and Myrmeleon Linnaeus contain species living in cave mouths. Two main types of caves are found in Australia; those with loose organic material and those with loose inorganic material. The cave habitat is divided into four zones and several subzones. Many species are restricted to one zone or another but species of Stenoleon may overlap zones. Discussions of the species and some of their biological requirements are provided. One new parasite record is given, an undetermined species of Echthrobacella Girault (Hymenoptera: Encyrtidae) reared from the larvae of Speleon yallingup Miller and Stange. Diagnoses or descriptions are given for all of these cave species including distributional data. Keys to the cave mouth inhabiting antlions of Australia (adults and larvae) are provided including a revised key to the Xantholeon Tillyard.
0249
We provide an illustrated list of species belonging to the genus Symmachia Hübner, [1819] (Lepidoptera: Riodinidae: Symmachiini) collected during an expedition conducted between September 10-21, 2011 in the northern part of the Parque Nacional da Serra do Divisor, Acre, Brazil, a remote region of Amazon rainforest. A total of 46 individuals were collected belonging to 15 species. For all recorded species, drawings of male genitalia and behavioral information are provided to support future studies on the taxonomy of the group. A new species, Symmachia divisora Dolibaina and Leite, sp. nov., is described, and a revised status for Symmachia hetaerina hesione Stichel, 1910, stat. rev., is proposed.
0248
The genus Gromphas Brullé (Coleoptera: Scarabaeidae) comprises four species, of which G. aeruginosa (Perty) and G. amazonica Bates are known to occur in Peru. This paper presents a revised description of Gromphas as well as illustrated diagnoses and distributional and ecological data on the Peruvian species.
0265
The multicicatrices fluted scale, Crypticerya multicicatrices Kondo and Unruh (Hemiptera: Coccoidea: Monophlebidae) is reported from the islands of San Andres and Providencia, Colombia, as a recent invasive species. This scale insect is polyphagous, and attacks numerous plants of economic importance such as avocado, breadfruit, mango, papaya and tropical ornamental plants. A compiled list of 95 host plant species of C. multicicatrices is given. A diagnosis of the adult female of C. multicicatrices and a revised taxonomic key to the species of the tribe Iceryini (Hemiptera: Monophlebidae) known from South America is provided. The pink hibiscus mealybug, Maconellicoccus hirsutus (Green) (Hemiptera: Coccoidea: Pseudococcidae), also is reported for the first time from the island of San Andres, where it is found commonly on Hibiscus spp. and Malvaviscus arboreus Cav. The need for the implementation of a classical biological control program in the archipelago in order to control invasive scale insect pests is discussed.
0273
Five new species of Hydroptila (Trichoptera: Hydroptilidae), H. murtlei sp. n., H. criokera sp. n, H. auriscuspa sp. n., H. santarosa sp. n., and H. ebroensis sp. n., from Florida are described and illustrated. We provide an annotated list of the 76 species known to occur in the state, including 10 species that represent new state records, one of which is also a new country record. New illustrations are presented for those species which were difficult to identify or exhibited new range extensions into Florida, namely: Hydroptila acadia Ross, H. ajax Ross, H. icona Mosely, H. latosa Ross, H. lloganae Blickle, H. maculata (Banks), H. novicola Blickle and Morse, H. wakulla Denning, Oxyethira arizona Ross, O. lumipollex Kelley and Harris, and O. simulatrix Flint.
0267
Beetles of the genus Paederus sensu stricto Fabricius, 1775 (Coleoptera: Staphylinidae) are often noticed because of their potency in inducing a dermal lesion, so-called linear dermatitis. This genus, which is placed in the tribe Paederini and subfamily Paederinae of Staphylinidae, currently comprises 490 species worldwide. Our study presents a short review of the former records of Paederus spp. in Iran plus some unpublished data. Field collections were done during March-October yearly (1997-2007) in northern and southern Iran and April-June from central, eastern, western and north-western Iran (2008-2009). The present study adds four species to the Iranian fauna of the genus Paederus, which are P. brevipennis Lacordaire, 1835, P. basalis Bernhauer, 1914, P. pubescens Cameron, 1914 and P. schoenherri Czwalina, 1899. Paederus brevipennis and P. schoenherri are the first members of the subgenus Harpopaederus Scheerpeltz, 1957, ever reported from Iran. Considering previous reports, museum-deposited materials and our findings, 14 species and subspecies of the genus Paederus, which are grouped in five subgenera, occur in Iran. These subgenera are Eopaederus Scheerpeltz, Harpopaederus Scheerpeltz, Heteropaederus Scheerpeltz, Paederus Fabricius and Poederomorphus des Cottes; however P. duplex spectabilis Bernhauer, 1913 is not yet attributed to any of the 13 so-far defined subgenera.
0269
Systematic revision of the genus Orodaliscoides Schmidt, 1913 (Coleoptera: Scarabaeidae: Aphodiinae)
(2012)
The species belonging to the genus Orodaliscoides Schmidt (Coleoptera: Scarabaeidae: Aphodiinae) are redescribed and figured. The following new combinations are proposed: Orodaliscoides fimbripes (Brown, 1928) and Orodaliscoides giulianii (Gordon, 1977).
0271
Amorbia concavana (Zeller) (Lepidoptera: Tortricidae) is reported in Florida, USA. Male and female specimens are figured, and new host records are given. The species is compared to other Nearctic species of Amorbia Clemens. Florida specimens are more similar to Cuban than to Central American material with respect to male genitalia.
0270
0272
A list of scale insects (Hemiptera: Coccoidea) intercepted on imported succulent plants at the plant quarantine of Korea from 2006 to 2010 is provided. A total of 15 species belonging to four families are listed. Of the 15 species that were intercepted, some species are potential pests that could gain entry and establish in Korea through the importation of plant material. Current information on intercepted scale insects is required to alert inspectors at quarantine sites to look carefully at succulent plants to prevent the introduction of an exotic species.
0275
We review the variegated mud-loving beetle fauna of the southeastern United States (Coleoptera: Heteroceridae), with an emphasis on Mississippi and Alabama. A key is presented to all species known to occur in the southeastern US, and includes several extra-limital species. Descriptions, illustrations and distribution maps are presented for each species. One new species, Tropicus nigrellus, is described and a lectotype is designated for Heterocerus schwarzi Horn. Our molecular data suggest that many previously recognized generic concepts are unnatural. As a result, the following generic synonymies are proposed: Culmus Pacheco 1964, Damfius Pacheco 1964, Efflagitatus Pacheco 1964, Erus Pacheco 1964, Filiolus Pacheco 1964, Gradus Pacheco 1964, Lanternarius Pacheco 1964, Lapsus Pacheco 1964, Neoheterocerus Pacheco 1964, Olmedous Pacheco 1964 and Peditatus Pacheco 1964 are synonyms of Heterocerus Fabricius; the genera Centariatus Pacheco 1964, Explorator Pacheco 1964, and Microaugyles Pacheco 1964 are synonyms of Augyles Schiödte. New combinations proposed and used in this paper include: Augyles auromicans (Kiesenwetter 1851, Heterocerus), Heterocerus parrotus (Pacheco 1964, Lanternarius), Heterocerus sandersoni (Pacheco 1964, Neoheterocerus) and Heterocerus selanderi (Pacheco 1964, Efflagitatus), Heterocerus texanus (Pacheco 1964, Peditatus). Other new combinations for North American species formed as a result of these generic reconfigurations, but not used in this paper, include: Heterocerus inciertus (Pacheco 1964, Damfius), Heterocerus longilobulus (Pacheco 1964, Neoheterocerus), Heterocerus sinuosus (Pacheco 1964, Lanternarius), Augyles canadensis (Fall 1920, Heterocerus), Augyles compactus (Fall 1937, Heterocerus), Augyles moleculus (Fall 1920, Heterocerus) and Augyles mundulus (Fall 1920, Heterocerus). New combinations for South American species suggested by molecular data, all originally described in the genus Efflagitatus, include: Heterocerus boliviensis (Pacheco 1964), Heterocerus freudei (Pacheco 1973), Heterocerus furmidus (Pacheco 1964), Heterocerus ingeniosus (Pacheco 1964), Heterocerus meridianus (Pacheco 1975), Heterocerus reticulatus (Pacheco 1964), Heterocerus solitarius (Pacheco 1973), Heterocerus splendidus (Pacheco 1964), Heterocerus tortuosus (Pacheco 1973), Heterocerus woodruffi (Pacheco 1975). No new combinations involving synonymy within the genera Culmus, Erus, Filiolus, Gradus, Lapsus, and Olmedous were proposed because all included species were originally described as Heterocerus. Introduction
0274
The purpose of this paper is to reassess the taxonomy of Phanaeus MacLeay (Coleoptera: Scarabaeidae) to accommodate new taxa and changes in taxonomic opinion since the publication of Edmonds’ 1994 revision of the genus. The two subgenera and 13 species groups established by Edmonds (1994) remain unchanged. A revised set of keys with accompanying comments and illustrations separates 54 recognized valid species. Seven recently described valid species are incorporated into the revised classification: Phanaeus blackalleri Delgado-Castillo, 1991; P. bordoni Arnaud, 1996; P. changdiazi Kohlmann and Solís, 2001; P. lecourti Arnaud, 2000; P. martinezorum Arnaud, 2000; P. yecoraensis Edmonds, 2004; and P. zapotecus Edmonds, 2006. The new name Phanaeus sororibispinus Edmonds and Zidek replaces Phanaeus alvarengai Arnaud, 1984, a primary junior homonym of P. alvarengai Pereira and d’Andretta, 1955. Three subspecies recognized in 1994 are elevated to species rank, new status: Phanaeus texensis Edmonds, 1994; P. pilatei Harold, 1863; and P. guatemalensis Harold, 1871. Phanaeus obliquans Bates, 1887 is removed from synonymy and given new status as a valid species. Twelve new junior subjective synonyms (bold) are recognized: P. tridens balthasari Arnaud, 2002 (of P. tridens Castelnau, 1840); P. dzidoi Arnaud, 2000 (of P. palaeno Blanchard, 1843); P. genieri Arnaud, 2002 (of P. amethystinus Harold, 1863); P. prasinus jolyi Arnaud, 2001 (of P. prasinus Harold, 1868); P. kirbyi ledezmai Arnaud, 2002 (of P. kirbyi Vigors, 1825); P. achilles lydiae Arnaud, 2000 (of P. achilles Boheman, 1858); P. chalcomelas grossii Arnaud, 2001 (of P. chalcomelas [Perty, 1830]); P pyrois malyi Arnaud, 2002 (of P. pyrois Bates, 1887); P. tridens moroni Arnaud, 2001 (of P. tridens Castenau, 1840); P. lecourti peruanus Arnaud, 2000 (of P. lecourti Arnaud, 2000); P. endymion porioni Arnaud, 2001 (of P. endymion Harold, 1863); P. pseudofurcosus Balthasar, 1939 (of P. tridens Castelnau, 1840); and P. prasinus trinidadensis Arnaud, 2001 (of P. prasinus Harold, 1868). “Phanaeus viridicollis” Olsoufieff, 1924 (sensu Arnaud 2002) is an unavailable name here considered a color variant of P. pyrois Bates, 1887.
0260
Coleoptera species composition and succession in downed woody debris habitats are poorly known in eastern North America. A photoeclector emergence chamber was used to concentrate Coleoptera that emerged from various decay classes of fine and coarse woody debris (FWD and CWD, respectively) collected in primary and secondary forest sites in Great Smoky Mountains National Park, Tennessee, USA. A total of 5673 adult beetle specimens, representing 305 lowest identifiable taxa within 227 genera and 51 families, was collected. One hundred fifteen species (38%) were represented by single individuals. Many more specimens and species were collected from CWD (4129 and 247, respectively) than from FWD (1544 and 162, respectively), but species richness accumulation curves were not significantly different. Many more specimens but an equal number of species were collected from primary forest (3347 and 207, respectively) than from secondary forest (2326 and 207, respectively). Species accumulation curves indicated higher richness in secondary forests. Based on a subset of 71 species represented by 10 or more specimens, 27 species were associated with fresh fine woody debris, 11 species with weathered fine woody debris, four with coarse woody debris decay class I, 14 with coarse woody debris decay class II, and eight with coarse woody debris decay class III–IV. Sixteen species were associated with secondary forests, whereas 28 species were associated with primary forests. Coarse woody debris decay class II taken in primary forests had highest absolute species richness with 156 species. In coarse woody debris species overlap decreased with increased difference in decay indicating faunal succession. Published works related to the study of the ecology of downed woody material are briefly summarized. Recommendations on developing a database of legacy trees for future researchers are given. Notes on the biology and photographs of the 71 species represented by 10 or more specimens are given to provide an atlas of eastern U.S. beetle species most commonly encountered in these habitats.
0259
The community within extremely decayed downed coarse woody debris, here referred to as decay class V (CWD5), has never been systematically sampled. The presumption has been that rotten wood is eventually overrun by surrounding soil and litter inhabitants. Leaf litter and CWD5 were sampled for Coleoptera with a sifting/ Berlese technique at three primary and three secondary forest sites in Great Smoky Mountains National Park, Tennessee, USA, during fall 2006 and spring 2007. A total of 4261 adult beetle specimens, representing 216 lowest identifiable taxa within 159 genera and 28 families, were collected. Sixty-six species (31%) were represented by single individuals. Many more specimens (3471) and species (170) were collected from leaf litter than from CWD5 (790 and 111, respectively) but species accumulation curves showed that species richness was not significantly different between the two habitats. Eight species were significantly associated with CWD5, and 40 species were significantly associated with leaf litter. Species richness was significantly higher in secondary forest than primary forest, but more species were significantly associated with primary than secondary forest. Species richness was significantly higher in spring than fall. Notes on the biology and photographs of the 59 species represented by 10 or more specimens are given to provide an atlas of common eastern U.S. beetle species found in these habitats. Overall CWD5 is a distinct but overlooked habitat that may harbor numerous undescribed species or species considered rare.
0263
This report contains the first inclusive phylogenetic analysis and taxonomic structure for the genera presently included within the scarabaeid tribe Dynastini. The study was based upon morphological, biogeographic and molecular data and yielded direct support for the recognition of three subtribes in the Dynastini: Dynastina MacLeay, New Status (Dynastes Kirby, Augosoma Burmeister, Megasoma Kirby, Golofa Hope), Xylotrupina Hope, New Status (Xylotrupes Hope, Allomyrina Arrow, Trypoxylus Minck, Xyloscaptes Prell) and Chalcosomina Rowland and Miller, New Subtribe (Chalcosoma Hope, Haploscapanes Arrow, Beckius Dechambre, Eupatorus Burmeister, Pachyoryctes Arrow). The results provide indirect but significant support for the origin of the tribe Dynastini to be of an age no later than the early Late Cretaceous. Molecular and morphological evidence suggests that Eupatorus as constituted in current taxonomic literature is not monophyletic, and taxonomic alternatives are discussed to address this deficiency. A plausible explanation is also provided for the perplexing use by F. W. Hope of the family-group names Xylotrupidae and Dynastidae.
0254
Surveys for adult Cylindera (s. str.) lunalonga (Schaupp) (Coleoptera: Carabidae: Cicindelinae) were conducted between 2001 and 2011 at over 80 sites throughout the species’ historic range in the San Joaquin Valley and Sierra Nevada Mountains of California. Previously considered extirpated from the Valley, these surveys resulted in finding adults at 18 sites, several with large populations (>50 individuals). As suggested by historic records, our studies confirmed that the Valley populations of Cy. lunalonga occur in what were historically wetland sites, but are now lowland agricultural croplands. Adults were always associated with wet, muddy soil within and along the edges of irrigation ditches. A comparison of morphology, behavior, habitat, and conservation is made with the only known extant Sierra Nevada population.
0251
Surveys during the summer of 2004 and August 2009 on Prince of Wales Island, Alaska, USA resulted in collection of 1064 adult spiders representing 84 species. Barcoding of spiders collected in 2009 resulted in DNA barcode data for 212 specimens representing 63 species. DNA barcode data were then used to facilitate the identification of otherwise unidentifiable juvenile and female specimens as well as to investigate phylogenetically four lineages with large branch lengths between specimens. Using morphological and DNA barcode identifications provided a more complete list of identified specimens than was possible using morphological data alone.
0257
A re-description and new records of Onthophagus viriditinctus Reitter, 1892 (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini), an uncommon species from Iran, are provided. The taxonomic position and some nomenclatural problems are discussed. The placement in the subgenus Exonthophagus Kabakov, 2006 is proposed. Images of the male, female, aedeagus and drawings of lamella copulatrix of Onthophagus viriditinctus and Onthophagus haroldi Ballion, 1871, the only other species included in the subgenus, are supplied. A key for distinguishing the two species is provided.
0252
Reticulitermes nelsonae, a new species of subterranean termite (Isoptera, Rhinotermitidae) is described from Sapelo Island, Georgia, United States of America, with specimens also found in North Carolina and Florida. The adult and soldier castes are described and illustrated to distinguish R. nelsonae from the four described Reticulitermes spp. known to occur in the southeastern United States.
0253
Important missing specimen data are provided for the original description of Ozophora woodruffi Slater (2005: 245) (Hemiptera: Lygaeidae), along with additional comparative relationships. Because of the missing type information, according to ICZN rules (1999), the species became a nomen nudum. This paper now serves to validate the name, and authorship becomes Slater (2012).
0241
A comprehensive revision of the Subfamily Parandrinae (Coleoptera, Cerambycidae) from the Afrotropical Region is presented. Two new genera are described: Adlbauerandra and Meridiandra. The known species from the Afrotropical Region are excluded from Parandra Latreille, 1802, which resulted in the following new combinations: Acutandra beninensis (Murray, 1862), A. comoriana (Fairmaire, 1895), A. gabonica (Thomson, 1858), Adlbauerandra morettoi (Adlbauer, 2004) and Meridiandra capicola (Thomson, 1861). Eighteen new species are described: Acutandra amieti, A. barclayi, A. camiadei, A. dasilvai, A. delahayei, A. gaetani, A. garnieri, A. grobbelaarae, A. hugoi, A. jolyi, A. leduci, A. leonardi, A. lucasi, A. noellae, A. oremansi, A. plenevauxae, A. quentini, and A. vingerhoedti. The species Parandra comoriana Fairmaire, 1895 is revalidated and a lectotype is designated. Parandra beninensis Murray, 1862 and Parandra conradti Kolbe, 1893 are revalidated. A lectotype is designated for Parandra gabonica Thomson, 1858 as the designation by Quentin and Villiers (1975) was considered as invalid. Keys are presented to separate genera and all species of African Parandrinae from each other. Illustrations are provided for all the species including many special characters used in the keys.
0247
The species of the "modestus" group of the genus Psalidognathus Gray (Cerambycidae: Prioninae) (currently with four species and three subspecies), which are recognized mainly by the antennomeres without apical spines, are studied. Psalidognathus thomsoni Lameere, 1885 is revalidated as a species distinct from P. modestus Fries, 1833. Psalidognathus erythrocerus reichei Quentin and Villiers, 1983 and P. erythrocerus pubescens Quentin and Villiers, 1983 are considered as different species from P. erythrocerus Reiche, 1840. Psalidognathus colombianus Demelt, 1989 and Psalidognathus modestus chocoensis Salazar, 2005 are synonymized with P. modestus, syn. nov. Psalidognathus rufescens, although belonging to “friendii” group, is redescribed and some remarks are added. Species redescribed: P. erythrocerus; P. modestus; P. reichei; P. rufescens; P. onorei Quentin and Villiers, 1983; and P. thomsoni. Psalidognathus cerberus sp. nov. is described from Colombia and Ecuador. A provisional key is provided to males of species of "modestus" group.
0233
0264
Aplagiognathus Thomson, 1861 (Coleoptera, Cerambycidae, Prioninae, Macrotomini) and its two species, A. spinosus (Newman, 1840) and A. hybostoma Bates, 1879, are redescribed and figured. As the type of A. spinosus is apparently lost, a neotype for the species is designated herein as is a lectotype for A. hybostoma. Keys to the North and Central American genera of Macrotomini (excluding the West Indies) and to Aplagiognathus species are provided. Details on the numerous changes in the nomenclatural history of the genus are also chronicled.
0246
The tribe Krisnini (Hemiptera: Cicadellidae) is presently known in the New World from three species from Puerto Rico and one species from Dominican Amber, all described in the Old World genus Krisna Baker. The three species from Puerto Rico are being placed in Lipokrisna, new genus, with Krisna insularis Oman as the type species, becoming Lipokrisna insularis (Oman), new combination. The other species in this genus are L. montana (Caldwell) and L. aesta (DeLong) both new combinations. The one species from Dominican amber is placed in the Genus Archiokrisna, new genus, with Krisna garciamarquezi Dietrich as the type species, becoming Archiokrisna garciamarquezi (Dietrich), new combination. The genus, Neokrisna, new genus, is described for six new species from the Dominican Republic, with Neokrisna oncora, new species, as the type species. The other new species in the genus are N. breva, N. decliva, N. libera, N. longula, and N. stena. A key to the species of Neokrisna is included. The three genera are compared with each other and the old World genus Krisna.
0242
0214
We conducted a morphological and mtDNA analysis of Cicindela decemnotata Say (Coleoptera: Carabidae: Cicindelinae) populations from throughout its geographic range to determine the extent of variation within the species and to assess the validity of subspecific names. The morphological study included an analysis of traditional subspecific characters including elytral color and maculations. These results provided evidence for the recognition of four subspecies of C. decemnotata, three of which are new: 1. C. d. decemnotata Say usually with green to dark green dorsal coloration and complete elytral maculations; it is widely distributed from Canada south to northern New Mexico and west into southern Utah and Idaho; 2. C. d. meriwetheri n. ssp. is distinguished by its bright green to green dorsal coloration, elytral maculation characterized by a thin middle band, a lack of humeral maculations, and a small number of genal setae; it has a restricted distribution from eastern Washington north to south central British Columbia; 3. C. d. bonnevillensis n. ssp. is distinguished by a combination of green to green-purple dorsal coloration and its greatly reduced elytral maculations; it is restricted to the area of ancient Lake Bonneville in north central Utah; 4. C. d. montevolans n. ssp. is distinguished by a predominately red-purple dorsal color and greatly reduced elytral maculations; its distribution is restricted to high elevations of the Bear River Mountains of northeastern Utah and southeastern Idaho. We also analyzed the mitochondrial haplotypes for cob and cox1 genes for one to six individuals from each of the six populations. This molecular analysis indicated recently diverged but discrete groups within C. decemnotata that are compatible with the subspecies distinctions postulated from morphology. These shallow molecular divergences within C. decemnotata are best explained by rapid phylogenetic radiation in the recent geological past in the wake of postglacial recession.
0230
The American species belonging to the genus Gonaphodiellus Schmidt (Coleoptera: Scarabaeidae: Aphodiinae) are redescribed and figured. Two new genera are described: Gonaphodioides and Gonaphodiopsis. Fourteen new species are described: Gonaphodiellus arcanus, G. cerropuntanus, G. manantlanicus, G. martinpierai, G. omiltemicus, G. sacatepequezis, Gonaphodioides newtoni, G. ratcliffei, G. skelleyi, Gonaphodiopsis deloyai, G. hypogea, G. montesdeocai, G. pachecoi, G. teopiscaensis. The lectotype of Aphodius columbicus (Harold, 1880) is designated. The following new combinations are proposed: Gonaphodiellus nigrinus (Schmidt, 1916), Gonaphodioides acutecernans (Balthasar, 1960), Gonaphodioides chapini (Hinton, 1934), Gonaphodioides columbicus (Harold, 1880), Gonaphodioides sincerus (Petrovitz, 1973). Aphodius (Gonaphodiellus) xalapensis Galante, Stebnicka and Verdú, 2003 is placed as a junior synonym of Gonaphodiellus bimaculosus (Schmidt, 1909), new synonymy. Aphodius ataenioides Hinton, 1938 and Aphodius (Blackburneus) castanescens Petrovitz, 1973 are placed as junior synonyms of Gonaphodioides chapini (Hinton, 1934), new synonymies.
0227
Two species of the weevil genus Compsus Schoenherr (Coleoptera: Curculionidae: Entiminae) from Colombia are redescribed: C. obliquatus Hustache and C.viridivittatus (Guérin-Méneville). A key by Hustache in 1938, to 33 of the 34 recognized species of Colombian Compsus then known, is modified to include the one additional species. Habitus illustrations of males and females of the two species and illustrations of selected parts of the male and female genitalia are included. Nearly all of the specimens of these two species were collected on various species or varieties of citrus, indicating their potential as citrus pests in the future.
0213
A preliminary checklist of the Cerambycidae, Disteniidae, and Vesperidae (Coleoptera) of Peru is presented. Within Cerambycidae, we record five subfamilies, 55 tribes, 345 genera and subgenera, and 714 species. Within Disteniidae, we record one tribe, six genera, and 11 species. We also record one subfamily, one tribe, one genus, and two species within Vesperidae. Four new country records are recorded: one species in the tribe Anacolini (Cerambycidae: Prioninae): Cycloprionus flavus Tippmann, 1953; and three species in the tribe Onciderini (Cerambycidae: Lamiinae): Cacostola simplex (Pascoe, 1859); Marensis simplex (Bates, 1865); Trachysomus cavigibba Martins, 1975. In addition, 161 species recorded are known only from Peru.
0258
The insect fauna within inflorescences of Heliconia bourgaeana Petersen (Zingiberales: Heliconiaceae) was evaluated in Parque Metlac, Fortín de las Flores, Veracruz, Mexico between May and October 1995. Floral bracts were present in May and some persisted to October, despite much destruction in August by a grackle, Quiscalus mexicanus (JF Gmelin) (Icteridae). Flowers were abundant in the bracts in May-June, after which their number declined as fruits matured. Insects that fed on the flowers were most abundant in May-June; several of them could be pollinators; the immature stages of most of these insects were absent from Heliconia Linnaeus. The floral bracts contained water enriched by decomposition of the flowers, and this provided nutrition for aquatic organisms. Syrphid larvae (Diptera) contributed the largest biomass among the aquatic insects, and their distribution among bracts was more uniform in time than that of other aquatic insects. Larvae of Culicidae and Psychodidae (Diptera) were more variable in density and were more abundant after decomposition of the flowers. The most abundant aquatic predators of culicid larvae were larvae of Toxorhynchites Theobald (Culicidae). The most abundant amphibious predators of dipterous larvae were adults of four species of Platydracus Thomson (Staphylinidae), one of them yet undescribed. To capture its prey, the Platydracus adult would immerse its head and thorax, with open mandibles, to snap at passing dipterous larvae.
0209
Chalcosicya maya, new species, (Coleoptera: Chrysomelidae: Eumolpinae) is described and the species
key of Blake (1951) is modified to accommodate it. This is the first known mainland species of this previously
Antillean genus. Sclerotized rods in the apical segment of the ovipositor of Chalcosicya Blake and related genera
are shown to be useful systematic characters within the eumolpine tribe Adoxini. Relationships with other genera
suggest that Chalcosicya belongs to a clade derived from ancestors with a western Tethyian distribution.
0220
0237
0212
The Caribbean Islands (or the West Indies) are recognized as one of the leading global biodiversity hot
spots. This is based on data on species, genus, and family diversity for vascular plants and non-marine vertebrates. This
paper presents data on genus level endemicity for the most speciose (but less well publicised) group of terrestrial
animals: the beetles, with 205 genera (in 25 families) now recognized as being endemic (restricted) to the West Indies.
The predominant families with endemic genera are Cerambycidae (41), Chrysomelidae (28), Curculionidae (26), and
Staphylinidae (25). This high level of beetle generic endemicity can be extrapolated to suggest that a total of about
700 genera of all insects could be endemic to the West Indies. This far surpasses the total of 269 endemic genera of all
plants and non-marine vertebrates, and reinforces the biodiversity richness of the insect fauna of the West Indies.
0211
We provide the first records of six species of biting midges (Diptera: Ceratopogonidae) in the genus
Culicoides Latreille from Mexico: C. baueri Hoffman, C. castillae Fox, C. debilipalpis Lutz, C. iriartei Fox, C. leoni
Barbosa and C. pusilloides Wirth and Blanton. In addition, C. leopoldoi Ortiz is confirmed from Mexico, and new
records are included for 25 other species previously recorded in Mexico: C. arubae Fox and Hoffman, C. blantoni Vargas
and Wirth, C. crepuscularis Malloch, C. daedalus Macfie, C. diabolicus Hoffman, C. foxi Ortiz, C. furens (Poey), C.
gabaldoni Ortiz, C. haematopotus Malloch, C. hylas Macfie, C. insignis Lutz, C. jamaicensis Edwards, C. luteovenus
Root and Hoffman, C. neopulicaris Wirth, C. nigrigenus Wirth and Blanton, C. pampoikilus Macfie, C. panamensis
Barbosa, C. paraensis (Goeldi), C. phlebotomus (Williston), C. poikilonotus Macfie, C. pusillus Lutz, C. stigmalis Wirth,
and all three species in the C. (Monoculicoides) variipennis complex, C. variipennis (Coquillett), C. occidentalis Wirth
and Jones, and C. sonorensis Wirth and Jones.
0240
The mirine plant bug Tropidosteptes forestierae, new species (Hemiptera: Miridae) is described from
Collier County, Florida, where it was found causing serious injury to an extensive ornamental hedge of Florida swampprivet, Forestiera segregata (Jacq.) Krug and Urb. (Oleaceae). Adult male and female, fifth instar, and egg are described. Color images of the adults, nymph, egg, and injury; scanning photomicrographs of selected adult structures; and illustrations of male genitalia are provided. A key to help distinguish the 16 species of Tropidosteptes known to occur in the southeastern United States is given.
0215
0206
Among the four oriental genera of the tribe Helluonini, Omphra Dejean (Coleoptera: Carabidae), is unique for its endemism to the Indian subcontinent and aptery. High intraspecies variability in morphological characters and limited diagnostic information makes species differentiation of the genus Omphra a complicated task. The present study provides a description of a new species, Omphra drumonti n. sp. from the Western Ghats, redescriptions and a key to the species of Omphra, details of intraspecies variation, discussion of relationships between taxa and distributional patterns of the genus. Based on the distributional patterns in the Indian subcontinent and flightlessness of the genus, inability to cross the physical barrier of the Ganges–Brahmaputra delta between north and peninsular India is indicated as the reason for its absence in the northeastern Indian subcontinent and endemism to the lower Indian subcontinent.
0256
Two new species of Goera, G. zwicki and G. nozakii (Trichoptera, Goeridae), from the Philippines and one new species, G. meyi, from Vietnam are described and illustrated herein. In addition, type specimens of G. disparilis Banks, G. octospina Banks, G. tagalica Banks, and G. uniformis Banks are illustrated. All species of Goera known from both the Philippines and Vietnam are listed.
0244
The known distribution of a rarely collected omethid, Omethes marginatus LeConte (Coleoptera), includes Arkansas, Connecticut, District of Columbia, Indiana, Maryland, New Jersey, New York, North Carolina, Ohio, Pennsylvania, Tennessee, Virginia, and West Virginia. Collection and ecological notes on recent collections in Arkansas, Indiana, and Virginia are presented.
0232
0218
A new genus and species of armored scale insect (Hemiptera: Diaspididae), Protomorgania koebelei.
Dooley and Evans, is described and illustrated from specimens collected by Albert Koebele on Pittosporum sp.
(Pittosporaceae) in Australia around the year 1900. A key to the genera of armored scale insects similar to Protomorgania
and known to occur in Australia is provided.
0205
A biodiversity inventory of the Lepidoptera of Pico Bonito National Park and vicinity, in the Department of Atlantida of northern Honduras, was initiated in 2009 to obtain baseline data. We present a revised checklist of Honduran butterfly species (updated from the initial 1967 lists), as well as the first comprehensive list of Honduran moths. Our updated list includes 550 species of Papilionoidea, 311 Hesperioidea, and 1,441 moth species.
0268
Delphastus quinculus Gordon and Diomus seminulus (Mulsant) (Coleoptera: Coccinellidae) are reported for the first time feeding on eggs and first-instar nymphs of Crypticerya multicicatrices Kondo and Unruh (Hemiptera: Monophlebidae). Diagnosis and illustrations are provided for both species. Updated information on their biology, hosts and geographical distribution is also provided.
0223
Two species of the genus Sisyra Burmeister (Neuroptera: Sisyridae), S. cameroonensis, n. sp., and S. gruwelli, n. sp., are described from the African Republic of Cameroon. Sisyra pallida Meinander is synonymized with Sisyra delicata Smithers, new synonymy, after comparison of the types of the former with topotypic paratypes of the latter. Type material of Sisyra nilotica Tjeder appears to be lost. Examples of Sisyra are recorded from Nigeria, Ethiopia and Uganda. A second species of the endemic African genus, Sisyborina, Monserrat, S. scitula, n. sp., is described from Cameroon, Guinea, and Zambia.
0216
0219
Extreme economic effects globally of various populations of the whitefly Bemisia tabaci (Gennadius) (Hemiptera: Aleyrodidae) led to an in depth study of the morphology of that species as well as that of numerous other species of whiteflies in the genus Bemisia Quaintance and Baker and other similar appearing species. The data collected are presented here as illustrations of the puparia (fourth instar nymphal stages) and discussions of morphology as it relates to species and generic separations within this closely knit group of insects. A brief history of the pest outbreaks of B. tabaci is given and an overview of the important morphological characteristics of aleyrodine whiteflies is provided. Each of the eighty illustrations is accompanied by a discussion of the more important aspects of morphology.
0282
This addendum to “An annotated list of the centipedes (Chilopoda) in The National Collection of Arachnids (Colección Nacional de Arácnidos, CNAN), Instituto de Biología, Universidad Nacional Autónoma de México” (México City), is based on new samples deposited in the last three years. It updates the preliminary list of 197 samples determined to genus and/or species. In this paper a total of 132 samples were added: Scutigeromorpha, 27; and Scolopendromorpha, 105. It also provides new state distribution records for Dendrothereua linceci (Wood, 1867), Scolopendra morsitans Linnaeus, 1758, S. polymorpha Wood, 1861, Rhysida immarginata (Porat, 1876), Scolopocryptops melanostomus Newport, 1845, Newportia spinipes Pocock, 1896, and Ectonocryptoides quadrimeropus Shelley and Mercurio, 2005.
0266
Lingafelteria, a new genus of Onciderini Thomson, 1860 (Coleoptera: Cerambycidae: Lamiinae) is described and illustrated. Five new species of Onciderini are also described and illustrated: Cylicasta mariahelenae, Lingafelteria giuglarisi, Psyllotoxus dalensi, Psyllotoxus faurei from French Guiana; Trestonia solangeae from Bolivia. Keys to the known species of Psyllotoxus Thomson, 1868 are provided. Psyllotoxoides albomaculata Breuning, 1961 is redescribed; and the first known females of Strioderes peruanus Giorgi, 2001 and Tibiosioma martinsi Nearns and Swift, 2011 are described. The following eight new country records are reported: Peritrox marcelae Nearns and Tavakilian, 2012 (Brazil); Pseudobeta ferruginea Galileo and Martins, 1990 (French Guiana); Tibiosioma martinsi Nearns and Swift, 2011 (Brazil, Peru); Trestonia exotica Galileo and Martins, 1990 (French Guiana); Trestonia morrisi Martins and Galileo, 2005 (French Guiana); Tritania dilloni Chalumeau, 1990 (French Guiana, Suriname).
0324
We provide new records of biting and predaceous midges (Diptera: Ceratopogonidae) from Guadeloupe in the subfamily Ceratopogoninae, including descriptions and illustrations of three new predaceous species in the genera, Parabezzia Malloch, Stilobezzia Kieffer and Palpomyia Meigen, respectively, and the first records of the New World predaceous genus, Amerohelea Grogan and Wirth, from the Caribbean region. We also provide the first Guadeloupe records of the biting midges, Culicoides (Anilomyia) decor (Williston), C. (Avaritia) pusillus Lutz, C. (Drymodesmyia) bredini Wirth and Blanton, C. (D.) poikilonotus Macfie, C. (Haematomyidium) hoffmani Fox, C. (Hoffmania) insignis Lutz, C. rangeli Ortiz and Mirsa and C. trilineatus Fox, and the predaceous midges, Brachypogon (Brachypogon) bifidus Spinelli and Grogan, B. (B.) telesfordi Spinelli and Grogan, B. (B.) woodruffi Spinelli and Grogan, Monohelea maya Felippe-Bauer, Huerta and Ibáñez-Bernal, Stilobezzia (Stilobezzia) diminuta Lane and Forattini, S. (S.) thomsenae Wirth, Amerohelea galindoi Grogan and Wirth, Bezzia (Bezzia) flinti Spinelli and Wirth, B. (Homobezzia) venustula (Williston) and Palpomyia insularis Spinelli and Grogan.
0299
0322
The Bolivian species of Polyrhaphis Audinet-Serville, 1835, (Coleoptera, Cerambycidae, Lamiinae) are reviewed and illustrated, with P. skillmani new species described. A key is presented to the six species recorded from Bolivia (P. angustata Buquet, 1853; P. argentina Lane, 1978; P. gracilis Bates, 1862; P. pilosa Lane, 1965; P. spinosa (Drury, 1773); and P. skillmani Wappes and Santos-Silva, new species). Their collection localities, based on recently identified specimens examined by the authors, are plotted to show the distribution of species, and displayed next to an ecoregion map of Bolivia to illustrate biogeographical information for Polyrhaphis.
0284
The elytra of Balcus signatus Broun (Coleoptera: Cleridae: Clerinae) from New Zealand have pale markings. Such markings, most prominently found in females, represent intraspecific variations of Balcus violaceus (Fabricius). Accordingly, Balcus signatus Brown is synonymized with Notoxus violaceus Fabricius, new synonymy. Four habitus figures of Balcus violaceus (Fabricius) are presented to display the range of elytral color variation in the species.
0305
Thirty-two new species of Perilypus Spinola (Coleoptera: Cleridae: Clerinae) are described; they are Perilypus ancorus, P. angustatus, P. aquilus, P. arenaceus, P. caligneus, P. cartagoensis, P. collatus, P. comosus, P. concisus, P. copanensis, P. copiosus, P. diutius, P. divaricatus, P. elimatus, P. flavoapicalis, P. galenae, P. hamus, P. hornito, P. infussus, P. iodus, P. lateralis, P. latissimus, P. licinus, P. limbus, P. miculus, P. odous, P. orophus, P. patulus, P. punctus, turnbowi, P. violaceus, and P. yasuniensis. Included in this work are 58 line drawings and 32 color habitus photographs of primary types. To facilitate species identification the species included herein are linked to a key to Perilypus species provided in a previous review of the genus.
0296
The two species of Sufetula Walker (Lepidoptera: Crambidae) occurring in Florida are reviewed based on adult specimens. Sufetula carbonalis n. sp. is described, Sufetula diminutalis (Walker) is diagnosed, and they are differentiated from related Caribbean species and similar sympatric Crambidae. Both are occasional root pests of ornamental palms. Unusual structural characters suggest that Sufetula is misclassified in Spilomelinae.
0304
Crossidius grahami Morris and Wappes new species (Coleoptera: Cerambycidae) is described from the Ohoopee Dunes of southern Georgia. Comments are included on the new species’ biology and disjunct distribution compared to other species of Crossidius LeConte. Illustrations include: dorsal and lateral views of the primary types of C. grahami, its natural habitat, the known host plant (Chrysoma pauciflosculosa (Michx.) Greene (Asteraceae)), pupal chamber, and dorsal views of both sexes of Crossidius humeralis quadrivittata Penrose, 1974, considered its closest anatomical counterpart and nearest geographical relative.
0303
We describe the rediscovery of the Florida scorpionfly, Panorpa floridana Byers (Mecoptera: Panorpidae), at Gold Head Branch State Park, Clay County, Florida, based upon a single, living, adult specimen photographed on 4 November 2010. The hardcopy photographic prints and electronic digital images of this panorpid are the first vouchers for P. floridana in 28 years, the only observation of a living specimen, and the sixth individual known of this seemingly rare Florida endemic.
0312
Dilatitibialis Duverger (61 species) (Coleoptera: Coccinelidae: Coccidulinae; Hyperaspidini) is discussed, species described, illustrations provided, and a key to all recognized taxa included. Cleothera cognata Mulsant, Cleothera cruciferae Mulsant, Cleothera fuscomaculata Mulsant, Cleothera gaynoni Mulsant, Cleothera glyphica Mulsant, Cleothera jucunda Mulsant, Cleothera luteola Mulsant, Cleothera mulsanti Kirsch, Cleothera oseryi Mulsant, Cleothera poortmanni Mulsant, Cleothera scenica Mulsant, Cleothera semicincta Weise, Cleothera tropicalis Mulsant, Hinda guttipennis Weise, Hyperaspis carolinae Crotch, Hyperaspis ceciliae Crotch, Hyperaspis dilatata Crotch, Hyperaspis florifera Vogel, Hyperaspis gravabilis Brèthes, Hyperaspis hybridula Crotch, Hyperaspis laterinotata Brèthes, Hyperaspis silvani Crotch, and Hyperaspis suzannae Crotch are transferred to Dilatitibialis, becoming new combinations. Lectotypes are designated for D. boliviana, D. cognata, D. florifera, D. fuscomaculata, D. gaynoni, D. glyphica, D. gravabilis, D. guttipennis, D. luteola, D. jucunda, D. mulsanti. D. poortmanni, D. retigera, D. scenica, D. semicincta, and D. staudingeri. A total of 38 new species of Dilatitibialis are described: Dilatitibialis annie, D. carmen, D. cindy, D. connie, D. crystal, D. dawn, D. diana, D. edith, D. edna, D. elaine, D. ellen, D. emily, D. ethel, D. fallax, D. florence, D. gladys, D. grace, D. josephine, D. kim, D. lillian, D. lois, D. marjorie, D. norma, D. paula, D. peggy, D. phyllis, D. rita, D. robin, D. rosa, D. shannon, D. sheila, D. sherry, D. sylvia, D. thelma, D. tiffany, D. tina, D. tracy, and D. wendy. Corrections are made to titles of previous Parts of this series, as follows: South American Coccinellidae, Part XII (Gordon 2007) is changed to Part XIII; South American Coccinellidae, Part XII (Gordon et al. 2013) is changed to Part XIV.
0289
Annotated world bibliography of host fruits of Bactrocera latifrons (Hendel) (Diptera: Tephritidae)
(2013)
Bactrocera latifrons (Hendel) (Diptera: Tephritidae) infests fruits and vegetables of a number of different plant species, with host plants primarily found in the plant families Solanaceae and Cucurbitaceae. Although B. latifrons is of primarily Asian distribution (e.g., Pakistan, India, Sri Lanka, Burma, China [Fujian, Yunnan, Hong Kong, Hainan], Thailand, Laos, Vietnam, Malaysia, Singapore, Taiwan, and Brunei), its range has expanded through introductions into Hawaii, Okinawa, Tanzania, and Kenya, and poses a threat of introduction into other countries where it does not presently occur. As with other tephritid fruit fly species, establishment of B. latifrons can have significant economic consequences, including damage and loss of food production, as well as requirements for implementation of costly quarantine treatments to permit export of commodities susceptible to infestation by B. latifrons. In order to avoid these adverse economic consequences, one needs to prevent the entry, establishment and spread of B. latifrons into a new habitat. To successfully achieve this, an accurate knowledge of the fly’s host plants is essential. Cognizant of this need, we prepared, and present here, a worldwide list of host plants for B. latifrons, with annotations on reported laboratory and field infestation data. Overall, a total of 59 plant species from 14 plant families are identified as hosts of B. latifrons, based on reported field infestation data.
0295
Aulacaspis difficilis (Cockerell) and Aulacaspis latissima (Cockerell), occurring on Elaeagnus glabra Thunb. and Distylium racemosum Siebold and Zucc. (Elaeagnaceae), are newly recorded in the Korean fauna of armored scales (Hemiptera: Diaspididae). The characters of these species are here redescribed with illustrative photographs and information on distribution and hosts. Also a key to species of Aulacaspis Cockerell is provided for correct species identification.
0300
Retournement or turning of the aedeagus about its longitudinal axis through about 180o during development is known in Chrysomeloidea (Coleoptera). This change in the orientation of the organ may be observed during the postembryonic development. This change produces certain morphological effects. By observing these morphological features in the imago the retournement may be inferred. Such morphological features in Curculionidae (Coleoptera) are here recorded. From this it has been inferred not only that retournement of the aedeagus is included in the ontogeny of curculionids, but also that the change of orientation of the organ occurs by the same mechanism as in Chrysomeloidea. These inferences attest the notion of a close phyletic relationship between the superfamilies Curculionoidea and Chrysomeloidea.
0298
We present a preliminary checklist of bees (Hymenoptera: Apoidea: Anthophila) compiled for St. Eustatius, an island located in the Lesser Antilles of the eastern Caribbean. The list has nine species, including six that have not been previously documented on St. Eustatius. One species is exotic to the Caribbean, one species is found only on St. Eustatius and St. Kitts, and five species occur elsewhere in the Lesser and Greater Antilles. Two of the collected specimens could not be assigned to a species; their geographical distributions are unknown.
0278
Genera of Brachiacanthini (Coleoptera: Coccinellidae: Hyperaspidinae) are discussed and a key to all recognized genera provided. Cyrea, new genus, is proposed, and Serratitibia, new genus, is erected and revised, Cleothera Mulsant and Hinda Mulsant are recognized as valid genera and revised. Helesius caseyi Sicard is transferred to Hinda and recognized as a synonym of Hinda designata Mulsant, new synonymy. Brachiacantha brethesi (Korschefsky), Cleothera abendrothi Kirsch, Cleothera ambigua Mulsant, Cleothera bisquatuorpustulata Mulsant, Cleothera decemsignata Mulsant, Cleothera gaillardi Mulsant, Cleothera humerata Mulsant, Cleothera tortuosa Mulsant, Cleothera traili Brèthes, Cleothera uncinata Mulsant, Hinda joeli Almeida and Milléo, Hinda modesta Weise, Hinda regularis Kirsch, Hyperaspis aliciae Crotch, and Hyperaspis fraudulenta Kirsch are transferred to Serratitibia, becoming new combinations. One new species of Hinda, H. ecuadorica, is described. A total of 73 new species of Serratitibia are described: Serratitibia amanda, S. andrea, S. angela, S. anna, S. ashley, S. barbara, S. barclayi, S. betty, S. beverly, S. bonnie, S. brenda, S. cheryl, S. christine, S. cynthia, S. debra, S. denise, S. donna, S. doris, S. elizabeth, S. evelyn, S. frances, S. gloria, S. heather, S. helen, S. irene, S. jacqueline, S. janet, S. janice, S. jean, S. jennifer, S. joan, S. joyce, S. judith, S. judy, S. julie, S. karen, S. katherine, S. kathleen, S. kathy, S. kelly, S. kimberly, S. laura, S. linda, S. lisa, S. loreto, S. lori, S. louise, S. margaret, S. marilyn, S. mary, S. martha, S. melissa, S. michelle, S. mildred, S. nancy, S. nicole, S. pamela, S. paprzycki, S. patricia, S. quincemil, S. rachel, S. rebecca, S. rose, S. ruby, S. ruth, S. sarah, S. satipoensis, S. shirley, S. stephanie, S. susan, S. tammy, S. teresa, and S. virginia. Lectotypes are here designated for Serratitibia lividipes, S. gaillardi, S. decemsignata, S. abendrothi, and S. ambigua.
0285
The new species Cephalocyclus majomaensis and Oscarinus cabreroi from Mexico are described and figured. While studying specimens from Mexico to complete a systematic revision of the Aphodiinae (Coleoptera: Scarabaeidae) of that region, we found a couple of species new to science. They belong to the genera Cephalocyclus Dellacasa, Gordon and Dellacasa, 1998 and Oscarinus Gordon and Skelley, 2007, and are described here.
0287
0288
0286
0297
0335
Chromosomes of four rare and localized Caribbean Prioninae (Coleoptera: Cerambycidae) species were analysed. All have 26, XY karyotypes. Those of Solenoptera canaliculata (Solenopterini) from Guadeloupe nearby islands Les Saintes and Marie-Galante and S. quadrilineata from Martinique look similar. They have a single pair of sub-metacentric autosomes. The karyotype of S. touroulti from St. Lucia has three sub-metacentric pairs. It appears closer to that of Hovorodon maxillosum (Mallodontini) from Marie-Galante which has ten sub-metacentric pairs. The CO1 gene sequence, taking two European species Aegosoma scabricorne (Prioninae: Aegosomatini) and Ergates faber (Prioninae: Ergatini) as external groups was analysed in S. canaliculata and S. quadrilineata. In spite of their karyotype similarity, their CO1 genes differ by a strong accumulation of mutations. Thus, either chromosomal or genetic data confirm the species status of the three closely related Solenoptera species. Ten different CO1 haplotypes are found among the 21 specimens of S. canaliculata studied from les Saintes and Marie-Galante. Both different haplotypes were found in each island and identical haplotypes were found in different islands. Hence, the gene flow was not interrupted. Biogeographical parameters favor the hypothesis that repeated passages between islands were made possible by floating trunks, principally from Les Saintes to Marie-Galante.
0336
Two new species of Deltosoma Thomson, 1864 are described from French Guiana: D. humeralis sp. nov. and D. fernandezi sp. nov. Deltosoma lacordairei Thomson, 1864 is redescribed based on examination of the holotype. A study of the aedeagi of these three species was conducted. Dorsal habitus photographs of both sexes, ventral photographs of males, and aedeagi photographs are provided.
0320
For the first time in Brazil, the weevil Cissoanthonomus tuberculipennis Hustache, 1939 (Coleoptera: Curculionidae) are reported preying on seeds of Cardiospermum grandiflorum Swartz (Sapindaceae). Observations are presented on oviposition and larval behavior, pupation site, and adult emergence. Photos of host plant, egg, larva, pupa and adult are provided.
0321
0327
The external morphology of the thorax and abdomen of four species of Neotropical Hesperiidae, belonging to different tribes, are described and illustrated. The morphological characters traditionally used in the classification of the family are reviewed and new information is added with emphasis on the characters usually neglected in the classification and identification of Hesperiidae and Lepidoptera. The use of these characters, along with those commonly used in literature for the identification and taxonomy of the family, is discussed, aiming to contribute to comparative studies of morphology and taxonomy of this group.
0323
0277
This paper presents the first comprehensive list of 2,688 species of Staphylinidae (Coleoptera) recorded from Brazil. The list is based on the taxonomic and ecological literature, and new records from some insect collections, and includes locality references for each species. In addition, Brazilian localities and the country-level distribution outside of Brazil are provided for each species. Brazilian localities are organized by state, and include the bibliographic reference and page number where each locality was reported. All localities are geo-referenced, organized by state, and listed in an Appendix.
0314
Presented is a checklist of the world Trogidae (Coleoptera: Scarabaeoidea) including synonyms, geographic distributions, type repositories where known, lists of valid species by genera and subgenera, citations of all papers containing original descriptions, and a supplemental literature section containing works on various other aspects of the family. The Literature Cited and Supplemental Literature sections combine to form a comprehensive bibliography.
0292
180 species of bark and ambrosia beetles (Curculionidae: Scolytinae and Platypodinae) are known to occur in Texas and Oklahoma. 175 species are known from Texas, 35 of which are reported here for the first time. 78 species are known from Oklahoma, 47 of which are new records for the state. Based on overall distribution patterns the largest group of species found in Texas and virtually all known from Oklahoma are widely distributed in eastern and southeastern North America, reaching their southwestern limits here. In the case of Texas other large elements include Neotropical elements shared with Mexico and a large number found in southwestern North America. New distribution and significant new host records are discussed. Distribution maps are included for most species and a checklist is provided as an appendix.