Insecta Mundi
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528
This revision concerns a small group of Western Palaearctic Copris species (Coleoptera: Scarabaeoidea:
Scarabaeidae) distinguished by having three lateral teeth on the foretibae. According to the literature, this group consists of four taxa: Copris armeniacus Faldermann, 1835, C. felschei Reitter, 1892, C. pueli Mollandin de Boissy, 1905 and C. umbilicatus Abeille de Perrin, 1901. Copris armeniacus is herein deemed a species inquirenda, and C. felschei, for which a neotype is designated, is deemed valid. Copris umbilicatus is recorded for the fi rst time from Turkey. A key to all species of the genus Copris known from the Western Palaearctic is provided. Variability of the cephalic and pronotal armature, and morphology of the parameres are illustrated.
527
The melon fly, Bactrocera cucurbitae (Coquillett) (Diptera: Tephritidae), is a widespread, economically important tephritid fruit fly species. Bactrocera cucurbitae infests fruits and vegetables of a number of different plant species, with many host plants in the plant family Cucurbitaceae, but with additional hosts scattered across many other plant families. Although thought to be native to India, its distribution has spread throughout many countries in Oriental Asia, into a number of Pacifi c Island nations, and into Africa. The documented introductions into countries outside its native distribution show that this species could establish in other countries where it does not presently occur, particularly through the movement of infested fruit. As with other tephritid fruit fly species, establishment of B. cucurbitae can have signifi cant economic consequences, including damage and loss of food production, as well as requirements for implementation of costly quarantine treatments to permit export of commodities susceptible to infestation by B. cucurbitae and inspection of susceptible imported commodities. In order to avoid these adverse economic consequences, one needs to prevent the entry, establishment and spread of B. cucurbitae into a new habitat. To successfully achieve this, an accurate knowledge of the fly’s host plants is essential. Cognizant of this need, we prepared, and present here, a worldwide list of host plants for B. cucurbitae, with annotations on reported laboratory and fi eld infestation data. Overall, 136 plant taxa from 62 plant genera and 30 plant families are identifi ed as hosts of B. cucurbitae, based on reported fi eld infestation data. The predominant family, as expected, is Cucurbitaceae, with 56 plant taxa (41.2% of all host plant taxa) in which fi eld infestation by B. cucurbitae has been documented. The family with the 2nd highest number of documented infested plant taxa is Solanaceae, for which there are published fi eld infestation data for 20 plant taxa (14.7% of plant taxa for which there is documented fi eld infestation). Papers that list plants as hosts of B. cucurbitae based only on laboratory data, those that list plants as a host but do not report any fi eld infestation data, and those that report interception data add an additional 137 host plant taxa, representing a total of 80 genera and 39 plant families, 20 of which are additional plant families beyond those for which there is fi eld infestation data. These additional species must be considered “undetermined” hosts for which additional data are needed to document actual host status. This paper is a comprehensive documentation of host plants of the melon fly based on recorded infestations in laboratory and/or fi eld, interceptions at ports of entry, or “listing only” associations. Host records presented here will be used in vetting and developing the offi cial USDA list of host plants of the melon fly, which will be published by APHIS as a federal order.
526
525
The first myrmecophilous fl ea beetle genus (Myrmeconycha Konstantinov and Tishechkin, new genus) with four new species (M. erwini Konstantinov and Tishechkin, new species – Ecuador, M. gordoni Konstantinov and Tishechkin, new species – Brazil, M. pakaluki Konstantinov and Tishechkin, new species – Panama, and M. pheidole Konstantinov and Tishechkin, new species – Costa Rica) is described and illustrated. It is compared with fl ea beetles of the subtribe Disonychina (Coleoptera: Chrysomelidae: Galerucinae: Alticini) and may be easily differentiated based on the external and internal features, which include the waxy surface of the head and pronotum, reticulated surface of the pronotum, and four longitudinal ridges on each elytron.
524
A new species of Autostichidae, Eraina beechei nov. sp., is described from specimens collected in the Andean mountains of Ñuble, Biobío. Diagnostic characters for the species are presented; photographs of adults, illustrations of male and female genitalia, and a key for identifi cation of Eraina species are provided.
523
The species of Dipropus Germar (Coleoptera: Elateridae) of the west-central region of North America are taxonomically reviewed. Historical records of D. approximatus (Candèze), D. ferreus (LeConte), D. simplex (LeConte) and D. soleatus (Say) in the region are based on misidentifi cations. Dipropus pericu new species is described from Baja California Sur. Dipropus reinae new species, D. sonora new species and D. yaqui new species are described from southern Arizona, New Mexico and Sonora. Dipropus warneri new species is described from Arizona, New Mexico and Texas. Dipropus sus (Candèze) is reported from Morelos, Sonora and Sinaloa as new state records. Keys to the species of the region are provided. The historical value of a specimen of Elater soleatus Say in the Dejean collection is discussed and its lectotype designated. Ischiodontus oblitus Candèze is treated as an objective synonym of Dipropus soleatus.
522
Ptyoiulus Cook 1895, the dominant parajulid diplopod genus in the eastern United States (US), comprises two species – P. impressus (Say 1821), with a slanted, fl ared, circumferentially entire, and marginally serrate apical calyx on the anterior gonopod coxal process, and P. montanus (Cope 1869), n. comb., with a smooth, upright, cupulate calyx that is open caudad and coaxial with the process’ stem. The genus occupies a broad area between the Mississippi River and Atlantic Ocean extending from southern New England, Ontario, and Michigan to the Florida Panhandle and four small disjunct ones – from Montreal, Québec, to northern Vermont, along southwestern Lake Michigan in Wisconsin and Illinois; northeastern/eastcentral Arkansas, primarily in Crowley’s Ridge physiographic feature and beside the “bootheel” of Missouri; and a point locality in northeastern Louisiana just south of the Arkansas line. A male from Chester County (Co.), Pennsylvania, is designated as the neotype of Julus impressus, as is one from Durham Co., North Carolina, for J. montanus. As both species inhabit Montgomery Co., Virginia, the type locality of J. montanus, we exercise the right of first reviser, conserve the latter name, and assign it to the species with the smooth, cupulate, and coaxial calyx. We also exercise first reviser rights and assign Parajulus ectenes Bollman 1887 to this form, thereby relegating it to synonymy under Ptyoiulus montanus. Other new synonymies include Ptyoiulus georgiensis Chamberlin 1943 under P. impressus and P. coveanus Chamberlin 1943 under P. montanus. Both Ptyoiulus and P. impressus are projected for Delaware and Rhode Island and newly reported from Québec, Connecticut, District of Columbia, Maryland, Mississippi, South Carolina, Vermont, West Virginia, and Wisconsin, and the genus and species, respectively, are newly documented from Louisiana and Arkansas; P. montanus is newly cited from Alabama, Arkansas, Georgia, Mississippi, and South Carolina. Ptyoiulus impressus occupies every state except perhaps Louisiana and is the only species in areas that were inundated during the Cretaceous and glaciated during the Pleistocene; by contrast, P. montanus inhabits a relatively narrow east/west transect through the center of the generic range. Their distribution patterns suggest an old species, montanus, being actively displaced by the younger and more successful impressus. The decurvature of the epiproct in uroblaniulinines appears to increase with age and developmental stage. A key is presented to parajulid familygroup taxa in the US and Canada east of the Rocky Mountains.
521
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519
Cnestus mutilatus (Blandford) (Coleoptera: Curculionidae: Scolytinae) is reported from Pennsylvania for the fi rst time, new state record. Specimens were collected using baited Lindgren funnels as early as 2013. Within Pennsylvania, C. mutilatus is now reported from Berks, Bucks, Lehigh, Montgomery, and York Counties.