Insecta Mundi
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560
Chiquitano gen. nov. Chiquitano volcanesensis sp. nov., Compsibidion achiraensis sp. nov. and Compsibidion amboroensis sp. nov. (Coleoptera: Cerambycidae: Cerambycinae: Neoibidionini) are described from Bolivia. Notes on Rhysium Pascoe, 1866 and Rhysium bimaculatum Pascoe, 1866 are provided, and Brechmoidion separatum Martins and Galileo, 2007 is transferred to Rhysium. Keys to species of Compsibidion Thomson, 1864, Brechmoidion Martins, 1969 and Rhysium Pascoe, 1866 are also provided.
570
A detailed study of the holotype of Sphecomyrma canadensis Wilson, 1985 (Hymenoptera: Formicidae) from Canadian amber has led to the conclusion that the specimen belongs to a new genus, here named Boltonimecia gen.n. Since the taxonomy of stem-group ants is not well understood, in order to find the taxonomic position of this genus, it is necessary to review the classifi cation of stem-group ants in a study of their relation to crown-group ants. In the absence of data for traditional taxonomic approaches, a statistical study was done based on a morphometric analysis of antennae. Scape elongation is believed to play an important role in the evolution of eusociality in ants; however, this hypothesis has never been confirmed statistically. The statistical analysis presented herein lends support to the view that antennal morphology reliably distinguishes stem-group ants from crown-group ants, to determine whether a species belongs to one or the other group. This, in turn, may indicate a relationship exists between eusociality and scape elongation. A review of Cretaceous records of ants is made and the higher classification of Formicidae with definitions of stem and crown groups is proposed. Newly obtained data are discussed focusing particularly on the origin, evolution and diversity of ants.
569
A primitive subfamily of false click beetles (Coleoptera: Eucnemidae: Phlegoninae) distributed primarily in the Neotropical region is revised. Euryphlegon gen. nov. is described from Belize in Central America. New species include: Phlegon chiriquiensis sp. nov. (Panama), Phlegon panamensis sp. nov. (Panama), Euryphlegon jacqueschassaini sp. nov. (Panama) and Euryphlegon parallelus sp. nov. (Belize). Phlegon herculeanus (Lacordaire) stat. res. is resurrected from synonymy with Phlegon buqueti Laporte. One new combination is proposed: Euryphlegon degallieri (Muona) (Phlegon). Based on a number of observed external character traits, Euryphlegon is placed in Orodotini Muona, 1993 within Macraulacinae Fleutiaux, 1922. Identifi cation keys are provided for species of Phlegon and Euryphlegon in the Neotropical region. The relationships among Phlegon, Euryphlegon, Euryptychus LeConte and other groups within Echthrogasterini Cobos, 1964 and Orodotini are discussed.
568
New Bolivian Rhinotragini (Coleoptera, Cerambycidae, Cerambycinae) are described: three species of Phygopoda Thomson, 1864 (P. longiscopifera sp. nov., P. boliviensis sp. nov. and P. chaquensis sp. nov.); and one species of Phygopoides Peñaherrera-Leiva and Tavakilian, 2007 (P. maxwelli sp. nov.). Two Brazilian species of Neophygopoda Melzer, 1933 are transferred to the genus Phygopoda: P. exilis (Melzer, 1933) comb. nov. and P. agdae (Martins, Galileo and Santos-Silva, 2015) comb. nov. All the species are illustrated, and a key to the Bolivian species of Phygopoda and host flower records are provided.
558
Six new species of false click beetle (Coleoptera: Eucnemidae) are described. These new species are: Fornax dixiensis sp. nov. (Florida, USA), Dromaeolus comayaguiensis sp. nov. (Honduras), Asiocnemis bicolor sp. nov. (Ecuador), Miruantennus chinensis sp. nov. (Yunnan prov., China), Miruantennus cuneiformis sp. nov. (Malaysia) and Nematodes africanus sp. nov. (Côte d’Ivoire). Identifi cation keys are provided for species of Fornax in the Nearctic ecozone and Miruantennus in the Palearctic and Indo-Malayan ecozones. A new key to include all the genera within the tribe Nematodini Leiler is also provided.
567
The taxonomic history of the rhinotragine genera Phygopoda Thomson, 1864 and Pseudophygopoda Tavakilian and Peñaherrera-Leiva, 2007 (Coleoptera: Cerambycidae: Cerambycinae) are discussed, and evidence is presented to suggest that some recent taxonomic changes made by Carelli and Monné (2015) were unjustified. Consequently, Phygopoda nigritarsis Gounelle, 1911 is moved to the genus Neophygopoda Melzer, 1933, creating the new combination Neophygopoda nigritarsis, the genera Panamapoda Clarke, 2014 and Paraphygopoda Clarke, 2014 are revalidated, and the species Paraphygopoda viridimicans (Fisher, 1952) and Paraphygopoda nappae Clarke, 2014 are also revalidated.
566
Dipropus tequesta Johnson new species (Coleoptera: Elateridae) is described from southern Florida. Dipropus fuscus (LeConte) is a new synonym of D. soleatus (Say), and D. granosus (Fall) is a new synonym of D. asper (LeConte). The fl ightless female of D. asper is described and provides the fi rst report of brachyptery and endogean habits in Dipropus. A key to the species of the eastern United States and a new checklist of species for the country are provided.
578
Tragosoma soror (Coleoptera: Cerambycidae: Prioninae), a new Nearctic species, is described and compared to its North American congeners. Tragosoma harrisii LeConte, revised status, and Tragosoma spiculum Casey, revised status are both resurrected from synonymy and considered valid species distinct from T. depsarium (Linnaeus). The Mexican T. nigripenne Bates is confi rmed as a valid species. Tragosoma chiricahuae Linsley, new synonym, is placed in synonymy under T. spiculum Casey, and T. sodale Casey, new synonym, T. parvicolle Casey, new synonym, and T. repens Casey new synonym are placed in synonymy under T. harrisii. Tragosoma repens is a junior objective synonym of T. parvicolle. A lectotype is designated for T. sodale. An updated synonymical and distributional checklist of Tragosoma of the world is provided.
575
Redescription of adult and observations of the weevil Revena plaumanni Bondar (Coleoptera: Curculionidae), preying on seeds of the palm tree Butia eriospatha (Mart. ex Drude) Becc. (Arecaceae), as well as oviposition and larval behavior, pupation site, and adult emergence are reported for the fi rst time in Brazil. Photos of host plant, immature forms (egg, larva and pupa) and adult are given. Illustrations of male and female genitalia are also provided.
576
579
A statewide survey of Wisconsin’s Nemonychidae and Anthribidae (Coleoptera: Curculionoidea) was conducted over one full (2012) and two partial (2011, 2013) fi eld seasons. Specimens were collected using a variety of techniques. Two species of the nemonychid genus Cimberis and 22 species of Anthribidae in 13 genera are now confi rmed. Generic and species-level keys for Wisconsin species are provided along with generic and species diagnoses. Species treatments also include a synonymy, a description, and information on natural history, phenology, distribution and collecting methods. Dorsal and lateral habitus images are also provided for most of the species.
580
602 [korr. Vers.]
Species of Mortoniella are revised for the northern and Andean part of the South American continent, including the countries of Bolivia, Peru, Ecuador, Colombia, Venezuela, and Guyana. All previously described species from the region are reillustrated and redescribed, except for Mortoniella santiaga Sykora, 1999 and M. quinuas Harper and Turcotte, 1985, whose types could not be located, and M. tranquilla Martynov, 1912, whose type is based on a female specimen and thus is currently unidentifiable. Included in the revision are 35 described species and 59 new species. Mortoniella similis Sykora, 1999 is considered a junior synonym of M. roldani Flint, 1991, and M. macuta (Botosaneanu, 1998) is considered a junior synonym of M. limona (Flint, 1981). A new subgenus, Nanotrichia, is recognized to accommodate species previously referred to as members of the ormina and velasquezi groups. Mexitrichia pacuara Flint, 1974 is designated the type species for the subgenus. Species previously referred to as members of the bilineata and leroda species groups are retained in the nominate subgenus, along with additional taxa not previously placed to species group, and treated within a number of subgroups. Previously described species of M. (Mortoniella) which are redescribed and reillustrated include: M. angulata Flint, 1963; M. apiculata Flint, 1963; M. atenuata (Flint, 1963); M. bifurcata Sykora, 1999; M. bilineata Ulmer, 1906; M. bolivica (Schmid, 1958); M. chicana Sykora, 1999; M. denticulata Sykora, 1999; M. elongata (Flint, 1963); M. enchrysa Flint, 1991; M. flinti Sykora, 1999; M. foersteri (Schmid, 1964); M. hodgesi Flint, 1963; M. iridescens Flint, 1991; M. leei (Flint, 1974); M. limona (Flint, 1981); M. marini (Rueda Martín and Gibon, 2008); M. paralineata Sykora, 1999; M. paraenchrysa Sykora, 1999; M. pocita (Flint, 1983); M. punensis (Flint, 1983); M. roldani Flint, 1991; M. simla (Flint, 1974); M. spinulata (Flint, 1991); M. squamata Sykora, 1999; M. unilineata Sykora, 1999; and M. wygodzinskii (Schmid, 1958). New species described in M. (Mortoniella), followed by their respective areas of distribution, include: M. acutiterga (Ecuador); M. adamsae (Peru); M. akrogeneios (Ecuador); M. applanata (Peru); M. auricularis (Colombia); M. barinasi (Venezuela); M. biramosa (Venezuela); M. bothrops (Peru); M. brevis (Ecuador, Venezuela); M. bulbosa (Peru); M. catherinae (Peru); M. chalalan (Peru); M. cornuta (Peru); M. cressae (Venezuela); M. croca (Peru); M. curtispina (Venezuela); M. curvistylus (Ecuador); M. dentiterga (Ecuador); M. dinotes (Peru); M. draconis (Ecuador); M. emarginata (Ecuador, Colombia); M. esrossi (Colombia); M. flexuosa (Colombia); M. furcula (Ecuador); M. gilli (Ecuador); M. gracilis (Venezuela); M. grandiloba (Venezuela); M. guyanensis (Guyana); M. hamata (Colombia); M. langleyae (Ecuador); M. longiterga (Ecuador); M. membranacea (Bolivia); M. monopodis (Colombia, Ecuador); M. parameralda (Ecuador); M. pica (Ecuador); M. proakantha (Ecuador); M. prolata (Peru); M. quadrispina (Ecuador); M. rectiflexa (Ecuador); M. ruedae (Bolivia); M. schlingeri (Colombia); M. silacea (Colombia, Ecuador); M. sinuosa (Bolivia, Peru); M. spatulata (Venezuela); M. tanyrhabdos (Venezuela); M. tridens (Peru); M. triramosa (Bolivia); M. tusci (Venezuela); and M. variabilis (Venezuela, Colombia). Species assigned to the subgenus M. (Nanotrichia) which are redescribed and reillustrated include: M. aequalis (Flint, 1963); M. aries (Flint, 1963); M. collegarum (Rueda Martín and Gibon, 2008); M. eduardoi (Rueda Martín and Gibon, 2008); M. macarenica (Flint, 1974); M. pacuara (Flint, 1974); M. usseglioi (Rueda Martín and Gibon, 2008); and M. velasquezi (Flint, 1991). Previously described species of Mortoniella, outside the area of coverage, that are transferred to the subgenus M. (Nanotrichia) include: M. alicula Blahnik and Holzenthal, 2011; M. bocaina Blahnik and Holzenthal, 2011; M. catarinensis (Flint, 1974); M. froehlichi Blahnik and Holzenthal, 2011; M. ormina (Mosely, 1939); M. rodmani Blahnik and Holzenthal, 2008; and M. tripuiensis Blahnik and Holzenthal, 2011. New species in the subgenus M. (Nanotrichia), followed by their respective areas of distribution, include: Mortoniella cognata (Ecuador, Venezuela); M. coheni (Ecuador); M. licina (Ecuador); M. paucispina (Peru); M. quadridactyla (Venezuela); M. simplicis (Venezuela); M. spangleri (Ecuador); M. triangularis (Ecuador); M. venezuelensis (Venezuela); and M. zamora (Ecuador). A key to the males of species from the region is also provided, as well as a key to females for the major subgroups and a species key to females of the velasquezi group. Finally, a partially resolved phylogeny of the species is presented, along with a discussion of evolutionary trends within the genus.
527
The melon fly, Bactrocera cucurbitae (Coquillett) (Diptera: Tephritidae), is a widespread, economically important tephritid fruit fly species. Bactrocera cucurbitae infests fruits and vegetables of a number of different plant species, with many host plants in the plant family Cucurbitaceae, but with additional hosts scattered across many other plant families. Although thought to be native to India, its distribution has spread throughout many countries in Oriental Asia, into a number of Pacifi c Island nations, and into Africa. The documented introductions into countries outside its native distribution show that this species could establish in other countries where it does not presently occur, particularly through the movement of infested fruit. As with other tephritid fruit fly species, establishment of B. cucurbitae can have signifi cant economic consequences, including damage and loss of food production, as well as requirements for implementation of costly quarantine treatments to permit export of commodities susceptible to infestation by B. cucurbitae and inspection of susceptible imported commodities. In order to avoid these adverse economic consequences, one needs to prevent the entry, establishment and spread of B. cucurbitae into a new habitat. To successfully achieve this, an accurate knowledge of the fly’s host plants is essential. Cognizant of this need, we prepared, and present here, a worldwide list of host plants for B. cucurbitae, with annotations on reported laboratory and fi eld infestation data. Overall, 136 plant taxa from 62 plant genera and 30 plant families are identifi ed as hosts of B. cucurbitae, based on reported fi eld infestation data. The predominant family, as expected, is Cucurbitaceae, with 56 plant taxa (41.2% of all host plant taxa) in which fi eld infestation by B. cucurbitae has been documented. The family with the 2nd highest number of documented infested plant taxa is Solanaceae, for which there are published fi eld infestation data for 20 plant taxa (14.7% of plant taxa for which there is documented fi eld infestation). Papers that list plants as hosts of B. cucurbitae based only on laboratory data, those that list plants as a host but do not report any fi eld infestation data, and those that report interception data add an additional 137 host plant taxa, representing a total of 80 genera and 39 plant families, 20 of which are additional plant families beyond those for which there is fi eld infestation data. These additional species must be considered “undetermined” hosts for which additional data are needed to document actual host status. This paper is a comprehensive documentation of host plants of the melon fly based on recorded infestations in laboratory and/or fi eld, interceptions at ports of entry, or “listing only” associations. Host records presented here will be used in vetting and developing the offi cial USDA list of host plants of the melon fly, which will be published by APHIS as a federal order.
523
The species of Dipropus Germar (Coleoptera: Elateridae) of the west-central region of North America are taxonomically reviewed. Historical records of D. approximatus (Candèze), D. ferreus (LeConte), D. simplex (LeConte) and D. soleatus (Say) in the region are based on misidentifi cations. Dipropus pericu new species is described from Baja California Sur. Dipropus reinae new species, D. sonora new species and D. yaqui new species are described from southern Arizona, New Mexico and Sonora. Dipropus warneri new species is described from Arizona, New Mexico and Texas. Dipropus sus (Candèze) is reported from Morelos, Sonora and Sinaloa as new state records. Keys to the species of the region are provided. The historical value of a specimen of Elater soleatus Say in the Dejean collection is discussed and its lectotype designated. Ischiodontus oblitus Candèze is treated as an objective synonym of Dipropus soleatus.
528
This revision concerns a small group of Western Palaearctic Copris species (Coleoptera: Scarabaeoidea:
Scarabaeidae) distinguished by having three lateral teeth on the foretibae. According to the literature, this group consists of four taxa: Copris armeniacus Faldermann, 1835, C. felschei Reitter, 1892, C. pueli Mollandin de Boissy, 1905 and C. umbilicatus Abeille de Perrin, 1901. Copris armeniacus is herein deemed a species inquirenda, and C. felschei, for which a neotype is designated, is deemed valid. Copris umbilicatus is recorded for the fi rst time from Turkey. A key to all species of the genus Copris known from the Western Palaearctic is provided. Variability of the cephalic and pronotal armature, and morphology of the parameres are illustrated.
526
524
A new species of Autostichidae, Eraina beechei nov. sp., is described from specimens collected in the Andean mountains of Ñuble, Biobío. Diagnostic characters for the species are presented; photographs of adults, illustrations of male and female genitalia, and a key for identifi cation of Eraina species are provided.
521
520