Insecta Mundi
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0458
0457
In recent years, populations of the Asian citrus psyllid, Diaphorina citri Kuwayama (Hemiptera: Liviidae), have increased in rural citrus orchards and urban backyard gardens. In order to fi nd biological control options for D. citri, a search for natural enemies was conducted in the department of Valle del Cauca, Colombia. The collections were carried out in citrus orchards in rural areas and house gardens and street trees in urban areas.
Natural enemies were collected from D. citri found on Citrus spp. and Murraya paniculata (L.) Jack (Rutaceae). A total of 16 species of natural enemies of D. citri distributed in six families in fi ve orders are reported from Colombia.
Herein we provide an updated list of 95 species of arthropod (arachnids and insects) natural enemies of D. citri distributed in nine orders and 23 families recorded worldwide.
0456
0455
A new scarab species of the genus Lycomedes Brème, 1844 is described, totaling fi ve species of the genus reported for Colombia. Lycomedes salazari was collected in an oak forest at the high Andean region of Santurbán in the municipality of California, Santander Department, Colombia. Besides inhabiting the highest altitude known for the genus, L. salazari has affi nities with L. buckleyi Waterhouse, 1880, but differs on the cephalic, thoracic and leg morphologies. Biological notes and an updated key for the adults of the genus are included.
0454
Two new species of Corimbion Martins, 1970 are described from Bolivia: Corimbion kuckartzi and Corimbion ledezmae. A previous key to the South American species of Corimbion (Martins 2009) is herein modified to include the new species. Dorsal, ventral and lateral habitus illustrations, as well as variation in color and dorsal pattern for C. kuckartzi, are also presented.
0453
New species of anthophilous Cerambycinae (Coleoptera, Cerambycidae) are described from Bolivia:
Molorchini, Merionoedopsis zamalloae sp. nov.; Heteropsini, Chrysoprasis azurearegina sp. nov., and C. maryhowardae sp. nov.; Rhopalophorini, Dihammaphora densiserrata sp. nov., D. dilmanappae sp. nov., D. espinotibia sp. nov., D. fosterorum sp. nov., D. paraperforata sp. nov., D. pilcomayoensis sp. nov., and Rhopalophora santacruzensis sp. nov. All species are illustrated and host flower records are provided.
0452
Klugiatragus gen. nov. is described for Epimelitta laticornis (Klug, 1825) because this species has closed procoxal cavities, a crucial diagnostic incompatible with Epimelitta Bates, 1870, which has open procoxal cavities. Both sexes of this species are illustrated.
0451
Information about the range of distribution of Centris (Centris) testacea Lepeletier has not been historically accurate. Range of distribution and identifi cation of the species is clarifi ed. Images of both sexes as well as male terminalia are presented. Centris testacea is a good species, with no synonym names. According to the available information, C. testacea should be considered endemic to the Greater Antillean island of Hispaniola.
0450
The nine Nearctic species of Laemophloeus Dejean (Coleoptera: Laemophloeidae) are reviewed, keyed, and illustrated. One species, L. apache Thomas, n. sp., is described as new. Two previously described species are synonymized: L. californicus Casey (= L. biguttatus (Say), n. syn.) and L. woodruffi Thomas (= L. fervidus Casey, n. syn.). A neotype is designated for L. biguttatus (Say), and lectotypes are designated for L. terminalis Casey and L. fervidus Casey. A checklist of the described world species is provided.
0449
This paper updates the knowledge about the occurrence of armored scale insects on tropical fruit trees in Argentina, and Pseudaulacaspis cockerelli (Cooley) is recorded for the fi rst time from Argentina and South America. Herein we present three new associations for the world: Aonidiella aurantii (Maskell) / Musa paradisiaca L.; Hemiberlesia cyanophylli (Signoret) / M. paradisiaca, and Acutaspis paulista (Hempel) / Persea americana Miller, and 10 new associations of armored scale / tropical fruit for Argentina are presented. Acutaspis paulista, Hemiberlesia palmae (Cockerell), Mycetaspis personata (Comstock) and P. cockerelli are cited for the first time on tropical fruits in this country.
0448
A collaborative survey of the scale insects (Hemiptera: Coccoidea) found on dracaena and fi cus plants in Cambodia, Laos, Thailand and Vietnam conducted in 2015 identifi ed 49 species of scale insects belonging to 36 genera in six families (25 species on dracaena, 42 species on fi cus). Of the species of scale insects that have been reported on dracaena and fi cus plants in southeastern Asia in the ScaleNet database (Ben-Dov et al. 2015), 32% and 17% were species collected in this survey, respectively. Twenty-three species (47%) of scale insects have been intercepted at Korean ports of entry on imported dracaena and fi cus plants from southeastern Asia from 1996 to 2014 (PIS 2015). Additionally, this list of species collected on these plants from exporting countries could be utilized as a basis for possible preventive measures in quarantine and inspection of traded products.
0447
Ichneumonidae are incredibly diverse, yet there have been few Guatemalan ichneumonid studies. We studied the phenology of 215 specimens of Zagryphus zulaya Gauld (Hymenoptera: Ichneumonidae: Tryphoninae), most captured in a light trap and two Malaise traps in a montane (1850m) forest from 2002 to 2014. One of the Malaise traps caught over 95% of the specimens. The results suggest that Z. zulaya is most abundant in May, June, July, and August, the rainy season. Zagryphus vegai Gauld is newly recorded from Guatemala; it was previously known only from Costa Rica. It was found at an altitude of 1850m, though its previous known range was 700m to 1500m.
0446
A new protocol to prepare material of Coccoidea (Insecta: Hemiptera) for observation with scanning electron microscopy is presented. This technique consists of cleaning samples (stored in 96% alcohol) with acetone and xylene, taking them directly to the hydration step, and subsequently dehydrating them in alcohol. Fixation in glutaraldehyde and post-fi xation in osmium tetroxide are omitted. The cleaning of the samples and the correct critical point dryer are very important.
0445
Seven species of Drapetes Megerle (Coleoptera: Elateridae) are recognized from northern North America. Drapetes chiricahua new species is described from southeastern Arizona and north-central New Mexico. Drapetes parallelus Van Dyke is a senior homonym of Drapetes paralellus Cobos by variant spelling, and a new synonym of Drapetes cylindricus Fall. Drapetes paralellus Cobos is replaced with Drapetes cobosi new name. Drapetes clarki Bonvouloir is not a North American species and is given a redesignated type locality in Brazil. Drapetes plagiatus (Boheman) is not a California species and is given a redesignated type locality in Panama. New state records are reported for Drapetes exstriatus (Say) from Arkansas, Colorado, Nebraska, and Oklahoma. A key to the species of the region is provided and each species is illustrated.
0444
Pandirodesmus rutherfordi, n. sp., represented by 18 individuals including eight adult males, occurs in secondary forests near Charlotteville and Speyside, Tobago, Trinidad and Tobago. Along with the type and second species, P. disparipes Silvestri, from Guyana and known only from females, the segmental legs of P. rutherfordi alternate between long (anterior pairs) and short (posterior ones), spiracular openings are on straw-like tubules, and ozopores are located on paramedian metatergal spines. These features appear to be adaptations for biotopes of loose sand, detritus, or frass, and 17 specimens, including the six juveniles, exhibit coatings of “sand grains” that are loosely cemented together and to the smooth, translucent, grayish-white exoskeleton. The tubules and spines elevate the spiracles and ozopores above the coating, thereby ensuring that they remain open and functional.
The coating, which provides camoufl age and lends strength and rigidity to the poorly sclerotized exoskeleton, is a subuniform “pavement” that covers the entire animal except the labrum/clypeus, tarsal and antennal apices, prozonae, paraprocts, and the gonopods in males. Ramose/dendritic setae, particularly on narrowly rounded podo-/antennomeres, trap “sand grains,” and the ozopore secretions apparently constitute the “glue” that cements the coating, as evidenced circumstantially by layers of “sand” between the spines on the anterior metaterga, where they are physically closest. The alternating segmental leg lengths, in part due to differing ventrolateral and ventromedial origins, appear to be an adaptation for lateral/sideways motion in which the long (anterior) legs extend laterally and pull the body to the level of the short (posterior) ones, which continue the motion while the anterior legs extend to begin the next stroke. The opposing legs perform the complementary pushing motion a fraction after the long legs initiate the pulling stroke and hence are slightly and purposefully out of sync. An adult male paratype lacks the coating, probably because it had just molted and lacked time to amass it; the juvenile female paratype of P. disparipes also is “naked,” as was, according to Silvestri, the now lost adult female holotype. Until fresh material is collected, coatings cannot be confi rmed for P. disparipes even though it shares the anatomical modifi cations that seem adaptions for such. The minute, triramous gonotelopodites of P. rutherfordi are unlike any known for a chelodesmid, so the current generic placement, in a monotypic tribe in the nominate chelodesmid subfamily, is retained. With species in both South America and the southern Antilles, Pandirodesmus/ini had to exist on both the “proto-Antillean” terrane and the adjoining part of Pangaean Gondwana before the former rifted in the Cretaceous/Paleocene, ~66 million years ago, and P. rutherfordi is a remnant of the former population that became isolated on present-day Tobago when the terrane fragmented. Affi nity between Guyanan and southern Antillean platyrhacid millipeds (Polydesmida: Leptodesmidea) suggest that Pandirodesmus/ini may occur sporadically as far north in the island chain as St. Lucia.
0443
New host and geographic records are given from Mexico, Guatemala, and Costa Rica for Megaplatypus exaratus (Blandford), previously reported only from three localities in Guatemala. Like most Platypodinae it is polyphagous and breeding hosts are reported from fi ve genera in four unrelated families. A redescription and updated diagnosis is included.
0442
The fauna of Panama is species-rich due to its location and topography, and the resulting diversity of microclimates and habitats. The last summary of information about the caddisfly fauna (Insecta: Trichoptera) of Panama in 1992 noted 168 taxa in 13 families and 39 genera. Since then, and through 2014,
a large number of publications, based on work by a dedicated cadre of individuals, have recorded a net gain of 78 species and six genera. In 2015, 17 new species to science and 38 new country records were added, including one new family and two new genera for the country; and, two species were removed. Thus, 299 species of caddisflies are now known from the Republic of Panama and are distributed among 14 families
and 47 genera. Given the higher diversity in neighboring Costa Rica (>500 species), we feel certain that there are many more species yet to be discovered.
0441
This revision of the Nearctic biting midges in the Culicoides (Monoculicoides) nubeculosus-stigma complex recognizes four species: C. grandensis Grogan and Phillips, C. riethi Kieffer, C. stigma (Meigen) and C. shemanchuki, new species, from Alberta, Canada and North Dakota, USA. Culicoides stigma is recorded for the fi rst time in the Nearctic region from Alberta, Canada. Culicoides gigas Root and Hoffman is a junior synonym of Culicoides riethi Kieffer (new synonym). A key is provided for the recognition of both sexes of the four North
American species in the Culicoides (Monoculicoides) nubeculosus-stigma complex.
0440
Xyleborus is one of the most diverse and abundant genera of ambrosia beetles, several species of which are considered important tropical pests. Given the current importance of this group, we present taxonomic keys for the 18 Mexican species of Xyleborus, using distinct external morphological features. A brief diagnosis and illustrations are included for each species.
0439
The fi rst natural history account of Discomorpha (Discomorpha) biplagiata (Guérin) (Chrysomelidae: Cassidinae: Omocerini) from Ecuador is presented. Larvae and adults feed on the leaves and stems of Cordia hebeclada Johnst. (Boraginaceae), the fi rst documented host plant for the species. Oviposition and pupation also occur on this host. Young larvae suffered heavy predation, especially from Ectatomma sp. (Formicidae: Ponerinae) and Oplomus nr. marginalis (Pentatomidae: Asopinae).
0438
Identification of caddisfly specimens (Insecta: Trichoptera) from Vietnam collected by or for the Royal Ontario Museum, the American Museum of Natural History, and the Museum für Naturkunde der Humboldt-Universität zu Berlin revealed nine new species in a variety of families and genera. New species include:
Philopotamidae—Dolophilodes carpenteri and Wormaldia montuosa; Stenopsychidae—Stenopsyche siniaevi; Polycentropodidae—Nyctiophylax hatinh; Psychomyiidae Eoneureclipsis afonini; Hydropsychidae—Maesaipsyche lappa; Glossosomatidae—Agapetus darlingi and Agapetus grimaldi; and, Molannidae—Molannodes sapa. In addition, new records for 11 species and two genera (Georgium and Tagalopsyche) of caddisflies from Vietnam are herein noted.
0437
The Trichoptera of Panama. II. Ten new species of microcaddisflies (Trichoptera: Hydroptilidae)
(2015)
In the Republic of Panama, the family Hydroptilidae (Insecta: Trichoptera) is currently represented by 82 species distributed among 14 genera. In this publication we provide descriptions and illustrations for 10 new species of hydroptilids in the subfamily Hydroptilinae: Tribe Leucotrichiini—Zumatrichia teribe and Z. zegla; Tribe Neotrichiini—Neotrichia pamelae and N. parabullata; and, Tribe Ochrotrichiini—Metrichia nowaczyki, M. sencilla, Nothotrichia panama, Ochrotrichia abrelata, O. nimmoi, and O. pulgara. The genus Nothotrichia is recorded from Panama for the first time.
0436
Plesioclytus morrisi Wappes and Skelley new species (Coleoptera: Cerambycidae) from the Ohoopee dune system in central Georgia is described with comments on the biology of the new species. The taxonomic placement of Plesioclytus Giesbert in the tribe Clytini is questioned as key characters are found to differ from the current characters used to define the tribe in the New World, resulting in its transfer to the newly erected Plesioclytini Wappes and Skelley new tribe, defined herein. Habitat photos for the new species and habitus photos for it and P. relictus Giesbert are provided.
0435
The Republic of Panama currently has 245 recorded species of Trichoptera distributed among 13 families. Herein we add 32 new country records for Panama, including one new family (Limnephilidae) and one new genus (Limnephilus). We also provide the first collection information for Oxyethira maya Denning for Panama. These results are part of an ongoing effort to characterize the caddisfly fauna of Panama, and to evaluate that country’s major cuencas (water basins).
0434
0433
The fauna of Dynastinae (Scarabaeidae) on the island of Saba, Dutch Caribbean, was investigated through fi eldwork during 2006 to 2015. Three species, belonging to the three tribes Cyclocephalini, Pentodontini and Phileurini, are newly recorded from Saba and are discussed, with summaries of all relevant information from the West Indies. Detailed locality data, temporal distributions, and habitus photographs are presented for each species.
0432
The checklist revealed 40 species of scale insects (Hemiptera: Coccoidea) intercepted at Korean ports of entry on dracaena and fi cus plants from southern Asia from 1996 to 2014. Brief diagnostic criteria and related information of the intercepted species are given to assist in the identifi cation of specimens of scale insects intercepted from these plant hosts. Additionally, this preliminary checklist of scale insects could be utilized as a basis for preventive measures in quarantine.
0431
The Colombian fluted scale (CFS), Crypticerya multicicatrices Kondo & Unruh (Hemiptera: Monophlebidae) is a polyphagous scale insect which affects about 100 species of plants. Between 2010–2013, the species was reported as an invasive pest on the islands of San Andres and Old Providence, Colombian territory in the Caribbean sea.
Currently, populations of the CFS also have increased in the city of Cali, Valle del Cauca State, mainland Colombia, affecting different host plants, especially leguminous trees such as Caesalpinia peltophoroides Benth., Pithecellobium dulce (Roxb.) Benth., P. saman (Jacq.) Benth. (Fabaceae) and mango, Mangifera indica L. (Anacardiaceae). With the aim of searching for natural enemies to control the CFS, field studies were conducted in the city of Cali between February 2013 and October 2014. As a result of this study, two species of lady beetles were found feeding on CFS, i.e., Anovia punica Gordon and Rodolia cardinalis (Mulsant) (Coleoptera: Coccinellidae). This is the first record of R. cardinalis preying on CFS.
0430
0429
0428
Erythemis Hagen, 1861 (Odonata: Libellulidae: Sympetrinae) is a Neotropical genus with ten species in which morphological characters vary widely. The aim of this paper is to study the taxonomic diversity of the genus Erythemis and to test the diagnostic value of morphological characters used to discriminate species. The diagnostic value of the morphometric characters is tested using discriminant function analysis, principal component analysis, and graphical exploration of the data. A total of 134 characters were studied; of those, 53 are recoded and 81 are proposed in this work. Discrete characters such as color, genitalia, ventral teeth of male cercus, extension of dark basal area in hind wing, and morphometric characters of abdominal carinae and antenodal wing venation are the most useful for species determination. In contrast, abdomen length/HW length ratio, vulvar lamina length, and spines of femoral structure are highly variable. A lectotype is designated for Diplax credula Hagen, 1861. Taxonomic keys for males and females are included, and variation in several characters is presented.
0427
Lectotypes are designated for the following species-level taxa, based on specimens in the Museum of Comparative Zoology, Harvard University: Tetratomidae (Hallomenus scapularis Melsheimer, 1846; Hallomenus serricornis LeConte, 1878); Melandryidae (Hypulus bicinctus Horn, 1888; Dircaea liturata LeConte, 1866; Dircaea fusca LeConte, 1878; Dircaea prona LeConte, 1878; Dircaea riversi LeConte, 1884; Hypulus trifasciatus Melsheimer, 1846; Microtonus sericans LeConte, 1862; Scraptia fl avicollis Haldeman, 1848; Scraptia rugosa Haldeman, 1848; Melandrya maculata LeConte, 1850; Melandrya striata var. bicolor Melsheimer, 1846; Melandrya striata var.
thoracica Melsheimer, 1846; Hypulus fulminans LeConte, 1859; Microscapha arctica Horn, 1893; Microscapha clavicornis LeConte, 1866; Orchesia castanea Melsheimer, 1846; Orchesia gracilis Melsheimer, 1846; Orchesia ornata Horn, 1888; Amblyctis praeses LeConte, 1879; Dircaea sericea Haldeman, 1848; Serropalpus obsoletus Haldeman, 1848; Serropalpus substriatus Haldeman, 1848; Carebara longula LeConte, 1866; Hallomenus quadripustulata Melsheimer, 1846); Mycteridae (Mycterus canescens Horn, 1879; Mycterus quadricollis Horn, 1874); and Boridae (Crymodes discicollis LeConte, 1850).
0426
At Topes de Collantes Natural Park, Alturas de Trinidad, in the mountains of Guamuhaya province of Sancti Spiritus, Cuba, 30 species of scale insects belonging to 21 genera, four families and an endemic species were identifi ed. Specimens were deposited at the insect zoological collection of the Ecology and Systematics Institute (CZACC), Cuba. A literature review was carried out and the world data base on scale insects, ScaleNet was consulted. A cartographic map indicating the collecting sites at the scale of 1: 250,000 was created using MapInfo Professional Version 4.5 program. Twelve species are recorded for the fi rst time for the mountains of Guamuhaya and further 10 species are recorded for other locations. 82% of the species were polyphagous, 11% were oligophagous and 7% were monophagous. 57% were introduced species, 43% were native and 79% are cosmopolitan or widely distributed species. 30 species and 21 families of host plants were identifi ed, of which 23 plant species were new host plant records for 19 scale insect species, and 11 botanical families are for the fi rst time recorded as hosts for 10 scale insect species. There were signifi cant differences in the distribution of scale insects and their host plant species, botanical families and vegetation type.
0425
0424
A total of 70 species of Nitidulidae are recorded from the state of Indiana. Nine of these species are recorded as new state records. The myrmecophile Amphotis ulkei LeConte, and the adventive Carpophilus marginellus Motschulsky, originally from Southeast Asia are added to the state fauna. The methods with which these new taxa were found are given, along with a brief discussion on trapping techniques used for the family as a whole.
Targeted use of vinegar-based baits and yellow sticky cards were used in addition to the general implementation of fermenting brown sugar and malt beer bait fi lled jugs, UV lighting, and Lindgren funnels. An updated checklist of all sap beetle species known to occur in Indiana is presented within a modern taxonomic framework for the family.
0423
Tiphallus torreon n. sp., the fi rst rhachodesmid milliped from Coahuila, Mexico, displays an iridescent turquoise pigmentation with patterned white paranotal markings and a truncated, subapical projection from the broad, non-descript gonopodal acropodite. Four genera – Strongylodesmus Saussure, Mexidesmus Loomis, and Ceuthauxus and Tiphallus, both by Chamberlin – contain forms exhibiting this general condition, but the last is the only one whose type species does. Synthetic treatments are essential to advance familial knowledge beyond the descriptive stage, and revising these four taxa would constitute a meaningful initial study. Rhachodesmidae extend from northern Nuevo León, Mexico, ca. 77 km (48 mi) from the Rio Grande, to central Costa Rica; Glomeridae (Glomerida), Platydesmidae (Platydesmida), and Stemmiulidae (Stemmiulida) show similar distributions whereas Allopocockiidae (Spirobolida) and Rhysodesmus Cook (Polydesmida: Xystodesmidae) traverse the river and occupy southernmost Texas. Tridontomidae, the other component of Rhachodesmoidea, occupies a small enclave in Alta Verapaz, Guatemala. Rhachodesmidae/oidea do not occur in Panama and are initially recorded from Belize; localities are needed from Honduras.
0422
The white coconut scale, Parlagena bennetti Williams, 1969 (Hemiptera: Diaspididae) is reported for the fi rst time on New Zealand fl ax, Phormium tenax J.R. Forst. & G. Forst. (Xanthorrhoeaceae), coconut, Cocos nucifera L., Manila palm, Veitchia merrillii (Becc.) H.E. Moore, oil palm, Elaeis guineensis Jacq. and an oil palm hybrid OxG: E. oleifera x E. guineensis (Arecaceae) in continental Colombia, and on coconut from Venezuela. Previously, P. bennetti was known only on the Caribbean islands of Trinidad and San Andres, causing serious damage to coconut, its only known host until now. A brief characterization of P. bennetti and an updated list of 23 species of scale insects of the family Diaspididae reported worldwide on Elaeis spp. is provided.
0421
Fifth instars for eight of the nine species of Microrhagus Dejean (Coleoptera: Eucnemidae: Melasinae:
Dirhagini), all endemic in the Nearctic region, are described and illustrated. Biological information is provided, along with a brief discussion of the larval morphology among the eight Nearctic species of Microrhagus. Adult descriptions
are provided for four new species. These new species include Microrhagus breviangularis new species, Microrhagus carinicollis new species, Microrhagus lecontei new species and Microrhagus opacus new species.
A checklist and identifi cation keys for both adults and most known larvae are provided.
0420
0419
0418
Mimuloria Chamberlin 1928 is revived from synonymy under Nannaria Chamberlin 1918a for Nannariini (Polydesmida: Xystodesmidae) with simple but apically ornamented gonopodal acropodites that arch or lean mediad and cross body midlines and opposing acropodites in situ. It encompasses two assemblages based primarily on the nature of the ornamentations, the castanea and dilatata species groups. The former includes three established species [M. castanea (McNeill 1887) M. missouriensis Chamberlin 1928 and M. davidcauseyi (Causey 1950a)], and the latter contains two new ones (M. dilatata [M. d. dilatata, M. d. sigmoidea], and M.
rhysodesmoides). Castanaria Causey 1950b is returned to synonymy under Mimuloria, and C. depalmai Causey 1950b is placed under M. castanea, thereby constituting a new synonymy. The fi rst illustrations of the holotype gonopods of Fontaria oblonga C. L. Koch 1847 and N. minor Chamberlin 1918a unequivocally establish their identities, and the convoluted nomenclatural tangle involving Oenomaea Hoffman 1964 and O. pulchella (Bollman 1889a) is detailed. Whether in Oenomaea or a new genus, separate generic status seems appropriate for Nannariini with subterminal solenomeres; N. morrisoni Hoffman 1948 and its potential synonym N. shenandoa Hoffman 1949 may also belong here. Initial tribal localities are reported from Alabama, South Carolina, and coastal Virginia and Maryland, and “O. pulchella” occurs in northern Alabama north/west of the Tennessee River; M. castanea is newly recorded from Missouri and Tennessee. A horizontally subtriangular distribution in the eastern and midwestern states is projected for Nannariini, which even occur on South Bass Island, Ohio, in Lake Erie, and may thus inhabit
nearby Pelee Island, Ontario, Canada.
0417
A new species, Chrysobothris cerceripraeda Westcott and Thomas (Coleoptera: Buprestidae), is described from prey specimens found in nests of the ground-nesting wasp, Cerceris fumipennis Say (Hymenoptera: Crabronidae), near Jacksonville, Florida, USA. A listing for all species of Buprestidae taken as prey of the wasp in that state is provided, four of which, including the species described herein, are new. Notes are also given for
four new state records for Buprestidae.
0416
0415
The genus Bembidion Latreille (Carabidae: Bembidiini) is reviewed for New Zealand. Thirty-six species-group taxa are recognized. Seven species are described as new: Bembidion (Zecillenus) karikari new species, Bembidion (Zecillenus) puponga new species, Bembidion (Zecillenus) tepaki new species, Bembidion (Zecillenus) waimarama new species, Bembidion (Zemetallina) bullerense new species, Bembidion (Zemetallina) mangamuka new species, Bembidion (Zemetallina) waiho new species. The taxonomic status of two species-group taxa is changed (valid names listed after equal sign): Bembidion (Zeactedium) orbiferum giachinoi Toledano, 2005 = Bembidion (Zeactedium) giachinoi Toledano, 2005; Bembidion (Zeperyphodes) nesophilum Broun, 1886 (previously synonymized with Bembidion (Zeperyphodes) callipeplum Bates, 1878) is resurrected from synonymy. A new synonymy is established (valid name listed after equal sign): Bembidion (Ananotaphus) rotundicolle eustictum Bates, 1878 = Bembidion (Ananotaphus) rotundicolle Bates, 1874. A concise review of the taxonomy of all taxa is provided.
Descriptions, identifi cation keys, illustrations of male genitalia, habitus photos, as well a s distributional data and
maps are given. Extensive information on ecology, biology, dispersal power, and collecting techniques is included for each species.
0414
Originally described from Great Guana Cay, Exuma Islands, Bahamas, Choranthus richmondi L. Miller, was also recorded from North Andros Island. Three specimens were collected on South Andros Island, Bahamas, in March 2014, representing a new record for this island. Photographs of adults, a complete diagnosis, genitalia illustrations, and a description of the habitat, nectar sources, and potential hostplant are presented. The male genitalia are illustrated for the fi rst time.
0413
0412
Two new species of Aulonothroscus Horn are described from The Bahamas and a third species is newly reported. Aulonothroscus inawa new species is described from Great Inagua and Aulonothroscus sibateo new species is described from Eleuthera. Aulonothroscus convergens (Horn) is reported from Andros, providing an island and new country record. These are the fi rst Aulonothroscus identifi ed to species from the Lucayan Archipelago and from a West Indies locality other than Guadeloupe. A key to the species of Bahamian Aulonothroscus is provided.
0411
Twelve new country records are presented for Diplocladus compactus Gerstmeier and Weiss, 2009, D. louvelii (Spinola, 1844), D. oculicollis (Fairmaire, 1885), D. rufus (Pic, 1953) Strotocera brevefasciata Gerstmeier and Weiss, 2009, and S. emerita (Péringuey, 1899) and the distribution of S. chicoi Gerstmeier and Weiss, 2009, is corrected.
0410
Within the tribe Coelidiini, subfamily Coelidiinae (Cicadellidae: Hemiptera), fragmentation of the genera Calodia Nielson, Olidiana McKamey and Taharana Nielson established the following 13 new genera: Cladolidia, type-species, Lodiana cladopenis Zhang; Creberulidia, type-species, Calodia paucita Nielson; Glaberana, type-species, Glaberana spadix, sp. nov.; Hamusolidia, type-species, Hamusolidia introrsa, sp. nov.; Hiatusorus, typespecies, Taharana schonhorsti Nielson; Laosolidia, type-species, Laosolidia complexa, sp. nov.; Orbisolidia, typespecies, Calodia spinocava Nielson; Singillatus, type-species, Lodiana furcata Nielson; Trinoridia, type-species, Trinoridia calcaris, sp. nov.; Tripesidia, type-species, Calodia warei Nielson; Tumidorus, type-species, Lodiana nielsoni Zhang; Webbolidia, type-species, Taharana webbi Nielson and Zhangolidia, type-species, Lodiana polyspinata Zhang. Nineteen genera in the tribe are treated.
The following 62 new species in 12 genera are described, illustrated and photographed: Calodia bicompressa (India); C. birama (Philippines); C. propennata (India); C. sichuanensis (China); C. sinuata (Laos); C. vincula (China, Vietnam); Creberulidia corniger (Laos); C. inflata (Thailand); C. multipenicula (Cambodia); C. ordospinosa (Thailand); C. penicula (Thailand); Glaberana ampla (Thailand); G. dentilamina (Thailand); G. longilamina (Thailand); G. penita (Laos); G. spadix (Laos); G. stylafurcata (Indonesia); Hamusolidia introrsa (Laos); Hiatusorus aviformus (Laos); H. robustus (China); H. supraspinosus (Thailand); Laosolidia complexa (Laos); L. tuberis (Laos); L. longiserrata (Laos); Olidiana tuberis (Vietnam); O. bispiculata (Laos); O. filiata (Thailand); O. implicata (Thailand); O. inaequabilia (Thailand); O. lata (Laos); O. parafringa (Laos); O. pennata (Laos); O. tonkinensis (Vietnam); O. vincula (Vietnam); Singillatus gracilius (Indonesia); S. ventrospinatus (India); Taharana abstrusa (Thailand); T. angusta (Vietnam); T. biavicula (Thailand); T. biunca (Thailand); T. brevicutata (Thailand); T. caverna (Malaysia); T. exiquitas (Thailand); T. forcipia (Thailand); T. gracilata (Thailand); T. incisura (Thailand); T. intimacalcara (Thailand); T. lacertosa (Thailand); T. mediolata (Thailand); T. minutura (Thailand); T. oblongiserrata (Laos); T. subspinata (Thailand); T. sublamina (Thailand); T. phetchahabunesis (Thailand); T. protriangulata (Thailand); T. subtumida (Thailand); T. truncata (Thailand); Trinoridia calcaris (India); T. trifida (Malaysia); Tripesidia kubani (Laos); Webbolidia kristenseni (Thailand); W. magna (Laos).
Taxonomy of all the genera is elucidated with a revised key to genera and species. The following formerly suppressed species are herein reinstated: Olidiana (Lodiana) flavofasciana Li, 1989, Olidiana (Lodiana) nigritibiana Li, 1987, Olidiana rufofasciana Li and Wang, 1989 and Webbolidia (Taharana) uniaristata Zhang, 1990. The following 3 species are new junior synonyms: Calodia flavinota Cai and Kuoh, 1993: 220 [= Calodia patricia (Jacobi), 1944:49], Olidiana yangi McKamey 2006: 502 [= Lodiana (Olidiana) hamularis Xu, 2000: 220] and Taharana yinggenensis Zhang and Zhang, 1994: 96 [= Taharana (Coelidia) sparsa (Stål) 1854: 254]. Taharana hainana Zhang 1994: 132 is a nomen nudum based on the same name in Zhang’s thesis (1988) which name was cited later by Li and Wang 1991: 275. Lodiana hainana Cai and He 2002: 139 is also a nomen nudum. A replacement name proposed herein is caii, nom. nov. in the genus Olidiana; Zhang’s 1994: 71 illustrations of subgenital plate (I) and aedeagus (M) of “fasciana Li” does not appear to represent the respective illustrations in Li 1991: 357 and may represent a new species in the genus Glaberana. The name of Olidiana nigridorsum (Cai and Shen) is changed to Olidiana nigridorsa (Cai and Shen) to agree with gender. Among 12 genera, 102 species are proposed in new combinations. Lodiana reductusi Xu and Kuoh, 1997 and Lodiana spicata Xu and Kuoh, 1997 are declared incertae sedis after attempts failed to locate the original descriptions and type specimens. Both species are provisionally assigned to the genus Olidiana. Two syntype specimens of Jassus egregius Schumacher, previously thought to be lost, were located in the Senckenberg Deutsches Entomologisches Institut, Müncheberg, Germany. The male specimen is designated lectotype herein. Six species in the Olidiana brevis (Walker) interspecific variation complex are elucidated with illustrations of male genitalia features.
New records, an updated checklist and a synoptic catalogue are also provided. All taxa including 264 valid species and 304 names are indexed.
0409
0408
0407
0406
We report on collections of seven species of pleasing lacewings (Neuroptera: Dilaridae) from the Americas. New country distributional records are reported for Nallachius pulchellus (Banks) from Honduras and Trinidad, and Nallachius phantomellus Adams from Ecuador. Nallachius ovalis Adams and Nallachius prestoni (McLachlan) are reported from Brazil representing the second reported collections of those species. Additional state records and in-country distributional information are presented for the other species. Two species could not be identified with certainty.
0405
0404
A new genus and four new species of false click beetles
(Coleoptera: Eucnemidae) from Southeast Asia
(2015)
A new genus and four new species of false click beetle (Coleoptera: Eucnemidae) are described. Bermilloides new genus is described from Bornean Malaysia. New species are: Calyptocerus iridis new species (Philippines), Bermilloides lumawigi new species (Malaysia), Spinifornax elongatus new species (Malaysia) and Ceratus antennatus new species (Thailand). Dorsal and ventral habitus, as possible, for each species are illustrated.
Male aedeagi are illustrated for Calyptocerus iridis and Ceratus antennatus. A new key is provided for Calyptocerus species in Southeast Asia.
0403
Pseudaulacaspis kiushiuensis (Kuwana), Pseudaulacaspis latiloba (Takagi and Kawai) and Pseudaulacaspis
ulmicola Tang, occurring on Quercus, Carpinus and Zelkova plants are newly documented in the Korean fauna of armored scales (Hemiptera: Diaspididae). Characteristics of these species are briefl y redescribed and illustrative photographs and information on their distribution and hosts are provided. A dichotomous key to species of Pseudaulacaspis from Korea is given for correct species identifi cation. In addition, the current status of Pseudaulacaspis forsythiae (Kanda) and Pseudaulacaspis nishikigi (Kanda) in Korea is discussed based on the results of a survey of the armored scales of Korea and related papers.
0402
Described, illustrated and discussed is a strikingly distinct new species of the scarabaeine genus Anomiopus Westwood, A. apuskispay Figueroa and Edmonds, from the Chanchamayo montane forest region of eastern Peru. It is a close relative of the recently described A. pishtaco, also from eastern Peru. Both species are noteworthy because of the bizarre form of the male clypeus, which bears extravagantly developed marginal teeth.
0401
A revision of the genus Acyphoderes Audinet-Serville, 1833 (Coleoptera: Cerambycidae), with a brief synopsis of the genus Bromiades Thomson, 1864, is presented. Acyphoderes to comprise eight South American species, Acyphoderes aurulenta (Kirby, 1818), A. abdominalis (Olivier, 1795), A. auricapilla Fisher, 1947, A. crinita (Klug, 1825), A. hirtipes (Klug, 1825), A. rubrohirsutotibialis Tippmann, 1953, A. carinicollis Bates, 1873 and A. amboroensis Clarke 2013; and Acyphoderes is divided into three informal species groups. A new genus, Ameriphoderes, is proposed for eleven Mexican and Central American species, Ameriphoderes acutipennis (Thomson, 1860), A. amoena (Chemsak and Linsley, 1979), A. ayalai (Chemsak and Linsley, 1988), A. bayanicus (Giesbert, 1991), A. cribricollis (Bates, 1892), A. magna (Giesbert, 1991), A. parva (Chemsak and Linsley, 1979), A. prolixa (Chemsak and Linsley, 1979), A. suavis (Bates, 1885), A. velutina (Bates, 1885) and A. yucateca (Bates, 1892); and Ameriphoderes is divided into two informal species groups. Acyphoderes delicata Horn, 1894 is placed in its own genus, Amerispheca. Other new genera are proposed for the following; Acyderophes for Acyphoderes fulgida Chemsak and Linsley, 1979; Brachyphoderes for Acyphoderes dehiscens Chemsak, 1997 and A. longicollis Chemsak and Noguera, 1993; Acutiphoderes for Acyphoderes odyneroides White, 1855; and Anomaloderes for Acyphoderes itaiuba Martins and Galileo, 2004. Acyphoderes sexualis Bates, 1885 and Sphecomorpha forficulifera (Gounelle, 1913) are moved to the new genus Forficuladeres. Acyphoderes synoecae Chemsak and Noguera, 1997 from Mexico is treated as a junior synonym of Sphecomorpha vespiventris
(Bates, 1880) from Guatemala. Acyphoderes cracentis Chemsak and Noguera, 1997 from Mexico, and Acyphoderes violaceus Bezark, Santos-Silva and Martins, 2012 from Costa Rica are placed in the new genus Odontogracilis, together with two species, Odontocera exilis Fisher, 1947 from Mesoamerica and Odontocera gracilis (Klug, 1825) from Brazil. Identification keys are provided for the genera, species groups (and the species in each one). All species are illustrated (both male and female when available). Plates illustrating the following are also provided; types of abdomen, hind leg, and aedeagus (tegmen and median lobe). Host-flower records are provided for many species.
0400
Distributional data are presented for Neotropical spongillafl ies (Sisyridae). New country records from Uruguay are presented for Climacia carpenteri Parfi n and Gurney, C. insolita Flint, C. versicolor Flint. Climacia desordentata Monserrat is synonymized with Climacia basalis Banks, NEW SYNONOMY. For the fi rst time, Sisyra apicalis Banks is reported from Guatemala, Suriname, and Uruguay, and S. panama Parfi n and Gurney is reported from Peru. Additional distributional data are presented for other species.
0399
Characterized by small body size, apically rounded/lobed anterior gonopod telopodites, long slender posterior gonopod telopodites, and torsion in the cyphopod receptacles, Floridobolus fl oydi, n. sp., is described from the southern sector of the Brooksville Ridge in northwestern peninsular Florida. It inhabits sandy “Big Scrub” environments like F. penneri Causey, 1957, and F. orini Shelley, 2014, and is documented from the sector’s center and northern periphery, in Hernando and Citrus Counties, respectively, with a sight record from the eastern periphery. Its discovery supports the thesis that each sand ridge in peninsular Florida may harbor a unique species of this endemic genus.
0398
The ladybeetle Anovia sp. (Coleoptera: Coccinellidae: Noviini), known as an important natural enemy of the Colombian fluted scale, Crypticeria muticicatrices Kondo and Gullan (Hemiptera: Coccoidea: Monophlebidae) is here identified as A. punica Gordon, 1972, and reported for the first time from Peru. Here we provide information and illustrations of the great variation in color and dorsal patterns of A. punica found in Colombia and Peru.
0397
The fauna of the small carrion beetles and round fungus beetles (Leiodidae) of the oceanic islands of the West Indies is reviewed with 11 genera and 81 species recorded. Keys to adults of all genera and species, descriptions, and figures are provided to aid in identification. All species are endemic to the islands of the West Indies. Most species are endemic to a single island, but some species in the Lesser Antilles occur on more than one island. It is certain that more species remain to be discovered, especially on larger and less explored islands. Two new genera are described: Parvocyrtusa (type species Parvocyrtusa hispaniolensis), and Pseudolionothus (type species Pseudolionothus insularis). The genus Pseudoagathidium Angelini is reported from the New World for the first time with one species. The higher taxa and 61 new species and their island distributions are as follows: Cholevinae, Eucatopini, Eucatops Portevin (first West Indian record): E. annulus, Hispaniola. Ptomaphagini, Proptomaphaginus Szymczakowski (four species, no new taxa), Greater Antilles and Bahamas (new genus record). Anemadini, Dissochaetus Reitter (five species, one new species): D. santalucia, St. Lucia. Leiodinae, Agathidiini, Agathidium Panzer (first West Indian record): A. minutum, Hispaniola. Pseudoagathidium Angelini (first New World record): P. ignotum, St. Vincent. Leiodini, Isoplastus Horn (first West Indian record): I. hispaniolensis, Hispaniola. Zeadolopus Broun (five known species, 28 new species): Z. acinaces, Hispaniola; Z. angulatus, St. Vincent; Z. antiguensis, Antigua, Saba, Montserrat; Z. atratus, Cuba; Z. bahamensis, Bahamas Islands (Andros Island); Z. caborojo, Hispaniola; Z. carinatus, Jamaica; Z caymanensis, Cayman Islands (Grand Cayman); Z. cubensis, Cuba; Z. dominica, Dominica; Z. exiguus, Hispaniola; Z. flavidus, Cuba; Z. hatomayor, Hispaniola; Z. hispaniolensis, Hispaniola; Z. iviei, Hispaniola; Z. jarabacoa, Hispaniola; Z. lavega, Hispaniola; Z. longipes, Hispaniola; Z. lucidus, Cuba; Z. miniusculus, Hispaniola; Z. nanus, Hispaniola; Z. nesiotes, St. Lucia and Martinique; Z. oviedoensis, Hispaniola; Z. parvantilliensis, Montserrat, Guadeloupe, Dominica, Martinique, St. Lucia, St. Vincent, Grenada; Z. paulus, Hispaniola; Z. pedernales, Hispaniola; Z. pusillus, Cuba. Parvocyrtusa (new genus, one new species): P. hispaniolensis, Hispaniola. Pseudolionothus (new genus, two new species): P. andersoni, Cuba; P. insularis, Hispaniola. Scotocryptini, Aglyptinus Cockerell (five known species, 19 new species): A. angulatus, Hispaniola; A. bahamensis, Bahamas Islands (Andros Island); A. biserriatus, Cuba; A. capitaneus, Cuba; A. dominica, Dominica; A. fortipunctatus, Cuba; A. grandis, Hispaniola; A. grenadensis, Grenada; A. hemipterus, Jamaica; A. hispaniolensis, Hispaniola; A. longipalpus, Hispaniola; A. luciae, St. Lucia; A. maculatus, Jamaica; A. martiniquensis, Martinique; A. minutus, Cuba; A. parvoculus, Jamaica; A. parvus, St. Lucia; A. sinuatus, Cuba; A. vincentii, St. Vincent. Creagrophorus Matthews (one known species, seven new species): C. bicolor, Martinique; C. cubensis, Cuba; C. dominica, Dominica; C. hispaniolensis, Hispaniola; C. microdentatus, Hispaniola; C. santalucia, St. Lucia; C. unidentatus, St. Vincent and Grenada.
0396
A new species of soft scale from Mexico, Toumeyella martinezi Kondo and Gonzalez sp. nov. (Hemiptera: Coccoidea: Coccidae) collected on Myrtillocactus geometrizans (Mart. ex Pfeiff.) Console (Cactaceae) is described and illustrated. An updated taxonomic key to the soft scale insects of the genus Toumeyella Cockerell known from Mexico is provided. A list of all currently known species of Toumeyella worldwide is given.
0395
The genus Paragymnopleurus Shipp, 1897 (Coleoptera: Scarabaeidae: Scarabaeinae: Gymnopleurini) is characterized and its constituent taxa are keyed and illustrated. Twelve species and five subspecies are deemed valid, and five species groups are recognized. Three new synonymies include: Paragymnopleurus stipes japonicus Balthasar is synonymized with P. ambiguus Janssens, and P. maurus malayanus Ochi and Kon and P. maurus pauliani Janssens are synonymized with the nominotypical subspecies. First country and provincial records are reported for P. brahminus (Waterhouse), P. maurus (Sharp) and P. sinuatus szechouanicus Balthasar. A lectotype is here designated for Gymnopleurus singularis Waterhouse, validating an unpublished designation. A checklist of valid species and synonyms is provided.
0394
We provide new state and county records of biting midges in the genus Culicoides Latreille (Diptera: Ceratopogonidae) from the southeastern United States collected with CDC miniature light traps during 2007–2012 in Florida, Georgia, Alabama, Mississippi, Louisiana, Arkansas, and Texas. The primary goals of the surveys were to identify the presence of exotic Culicoides, and determine the ranges of known and possible vectors of bluetongue virus (BTV) and epizootic hemorrhagic disease virus (EHDV). Included are the fi rst records of: Culicoides (Amossovia) beckae Wirth and Blanton from Louisiana and Mississippi, C. (A.) oklahomensis Khalaf from Alabama and Arkansas, C. (Avaritia) alachua Jamnback and Wirth from Alabama, C. (Culicoides) neopulicaris Wirth from Alabama, C. (Drymodesmyia) butleri Wirth and Hubert from Texas, C. (Hoffmania) insignis Lutz from Mississippi, C. (Oecacta) barbosai Wirth and Blanton from Georgia, C. (Silvaticulicoides) loisae Jamnback from Alabama, and C. kirbyi Glick and Mullen from Mississippi. We also provide new Florida county records for C. alachua, C. barbosai, C. (Beltranmyia) hollensis (Melander and Brues), C. insignis, and C. (Monoculicoides) sonorensis Wirth and Jones; a new Georgia county record for C. alachua; and new Alabama county records for C. insignis, and C. sonorensis.
0393
A revised checklist for the butterfl y families Hedylidae and Hesperiidae (Lepidoptera) of Trinidad (Trinidad and Tobago) is presented, bringing nomenclature in line with modern usage, and indicating synonyms from earlier lists and papers. The following are new records for Trinidad: Thessia athesis (Hewitson), Bungalotis clusia Evans, Sarmientoia eriopis (Hewitson), Pellicia tyana toza Evans, Quadrus contubernalis contubernalis (Mabille) and Decinea decinea derisor (Mabille). The checklist includes two species of Hedylidae and 314 species of 157 genera of Hesperiidae, comprising 87 species of 37 genera of Eudaminae, 73 species of 41 genera of Pyrginae, and 154 species of 80 genera of Hesperiinae.
0392
We clarify the taxonomy and nomenclature of several species-group names and the family-group name for the anticheirine scarabs (Scarabaeidae: Rutelinae: Rutelini), a group that includes nearly 550 species and subspecies in 44 genera and subgenera. These clarifications are necessary due to taxonomic and nomenclatural inaccuracies in recent revisionary works. We provide a list of genera included in the anticheirine scarabs sensu Soula. Numerous nomenclatural changes are necessary due to invalid type designations or misspellings: the valid type species of Aequatoria Arrow is Chlorota associata Waterhouse; the valid type species of Chlorota Burmeister is Rutela terminata Le Peletier de Saint-Fargeau and Audinet-Serville; the valid type species of Dorysthetus Blanchard is Macraspis chlorophana Burmeister; the valid type species of Anticheira Eschscholtz is Scarabaeus virens Drury. Changes to the type species of Anticheira result in new combinations: Anticheira virens virens (Drury), new combination, Anticheira virens bleuzeni (Soula), new combination and Anticheira virens jossi (Soula), new combination. Some anticheirine species names were not associated with species descriptions or type designations, making these names nomina nuda and unavailable: “Chlorota smithi”, “Macraspis cincta guatemalensis”, “Parachlorota equatoriana”, “Parachlorota pardoi”, “Pseudothyridium (Megathyridium) bousqueti”, “Theuremaripa rivae brasiliensis” and “Thyridium cupriventre blanchardi”. Dorysthaetus rufipennis Dejean is a nomen nudum and an unavailable name. We correct numerous misspellings from Soula’s five volumes on anticheirine scarabs and provide a list to avoid propagation of these errors by future researchers. Eighteen names are unavailable infrasubspecific names, and thirty-six of Soula’s names were misspelled. These names confuse and obscure ruteline systematics. Lastly, we provide a comprehensive list of infrasubspecific names in the anticheirine scarabs and we establish these as available or unavailable names.
0391
0390
Brachiacantha Dejean (Coccinellinae: Hyperaspidini), containing 49 species, is discussed, species described, illustrations provided, and a key to all recognized taxa included. Coccinella octopustulata F., Cleothera groendali Mulsant, Hyperaspis argentinica Weise, Hyperaspis blandula Weise, and Hyperaspis egae Crotch are transferred to Brachiacantha. Cleothera billoti parva Mulsant is recognized as a valid species and transferred to Brachiacantha. Brachiacantha arrowi Brèthes, Brachiacantha australe Leng, Brachiacantha manni Nunenmacher, and Brachiacantha propria Kirsch are recognized as synonyms of Brachiacantha groendali (Mulsant). A total of 29 new species of Brachiacantha are described; B. amber, B. anita, B. april, B. cathy, B. clara, B. charlotte, B. danielle, B. darlene, B. debbie, B. eleanor, B. emma, B. esther, B. eva, B. gail, B. hazel, B. jamie, B. joanne, B. jill, B. juanita, B. lauren, B. leslie, B. lynn, B. monica, B. pauline, B. pseudoarrowi, B. regina, B. sally, B. valerie, and B. veronica. Lectotypes are designated for B. argentinica (Weise), B. armandi (Mulsant), B. arrowi Brèthes, B. bahiensis Brethès, B. bilineata Weise, B. blandula Weise, B. bruchi Weise, B. loricata (Mulsant), B. octopustulata (F.), B. propria Kirsch.
0389
Pocket gopher burrows were sampled from 22 counties within Arkansas to determine the associated faunal composition of three major families of Coleoptera (Histeridae, Leiodidae and Scarabaeidae) commonly associated with pocket gopher burrows. We collected eight species of Histeridae, four species of Leiodidae and eight species of Scarabaeidae from the burrows of Geomys breviceps Baird. Three of the Histeridae were new state records, Geomysaprinus goffi Ross, G. rugosifrons (Fall) and Margarinotus felipae (Lewis). All of the Leiodidae were new state records and one Scarabaeidae was a new state record, Dellacasiellus concavus (Say). The most commonly collected scarab beetles were Cryptoscatomaseter haldemani (Horn) and Geomyphilus insolitus (Brown). The most commonly collected hister beetle was Onthophilus kirni Ross. The Leiodidae were infrequently captured.
0388
Trixagus steineri (Coleoptera: Throscidae), a new species and first genus record from The Bahamas
(2014)
Trixagus Kugelann 1794 is the second largest genus of Throscidae with 80 valid species presently assigned. Horn (1885, 1890) reviewed the species for the United States and Mesoamerica, and then Blanchard (1917) revised part of the family for Canada and the United States in a posthumous article edited by H.C. Fall. Schenkling (1928) provided the only published worldwide catalog. Yensen (1975) provided a modern revision for the species for Canada and the United States, and then (Yensen 1980) described T. cobosi from Panama and provided a new key to all of the described American species. Among these studies only T. chevrolati (Bonvouloir 1859) was recorded from southern Florida, thus being a species potentially shared with The Bahamas. Aulonothroscus bicarinatus Fleutiaux (1911, 1947) (Blackwelder 1944), from Guadeloupe, is the only other throscid species previously described from the West Indies. Prior to the work of Lawrence and Newton (1995) the subfamily Lissominae received divergent treatment and was usually treated as a subfamily of Throscidae (e.g., Schenkling 1928, Blackwelder 1944, Yensen 1975). This subfamily is represented in The Bahamas by two species of Drapetes Megerle 1821 (Turnbow and Thomas 2008). Representation of the subsequently restricted family in The Bahamas was provided by two undetermined species of Aulonothroscus Horn listed by Turnbow and Thomas (2008) from Andros, Eleuthera, and Great Inagua islands; these will be treated separately from here. The reporting of a new species of Trixagus from New Providence Island provides a second genus from the country and the entire Lucayan Archipelago, and suggests that related species of both genera from Hispaniola and Cuba are probable and await discovery (Peck 2005; Perez-Gelabert 2008).
0387
0384
Twenty six species of the genus Purenleon Stange (Neuroptera: Myrmeleontidae: Nemoleontini) are recognized from the New World, of which ten are described as new species: P. oaxacae, P. fernandezi, P. tibialis, P. adamsi, P. andinus, P. apache, P. aztecus, P. cavei, P. farri and P. toltecus. Neotypes are designated for Psammoleon banksi Esben-Petersen 1933 and for Formicaleo inaequalis Navás 1913. The larvae of sixteen species have been reared and are described with data on their biologies. The genus is subdivided into three species groups based mostly on leg structure. The inscriptus group consists of fi ve species, the tibialis group consists of two species and the rest of the species are in the bistictus group. Keys to the species based both on adults and larvae are provided and descriptions given.
0383
Besides the two species at present known belonging to the genus Trichonotuloides Balthasar (T. glyptus (Bates) and T. latecrenatus (Bates)), two new Mexican species, T. alfonsinae and T. hansferyi, are herein described (Coleoptera: Scarabaeidae: Aphodiinae). The complete set of fi gures is supplied for all taxa herein dealt with.
0382
The known range of Oxybleptes meridionalis Smetana (Coleoptera: Staphylinidae) is expanded in Florida, USA, from Indian River and Manatee counties to now include Brevard, Highlands, Orange, Seminole and Volusia. Oxybleptes davisi (Notman) is confi rmed to exist in Florida, with records from Leon, Liberty and Wakulla counties in the Panhandle, and Orange County in central Florida. Lissohypnus texanus Casey is newly reported from Florida. A new species, Lissohypnus fullertoni, is described from Florida. Diochus schaumii Kraatz reverts to this original spelling; its widespread form in Florida is identical to that in the northeastern USA.
0381
Aleurolobus confusus David and Subramaniam and Bemisia lespedezae (Danzig) occurring on Stephanandra incisa (Thunb.) and Lespedeza bicolor Turcz. are newly documented in the Korean fauna of whiteflies (Hemiptera: Aleyrodidae). Characteristics of these species are briefly redescribed and illustrative photographs and information on their distribution and hosts are provided. A dichotomous key to whiteflies of Korea is given for correct species identification. In addition, the current status of Bemisia takahashii (Danzig) in Korea is discussed based on the results of a survey of the whiteflies of Korea and related papers.
0380
Two new species of pselaphine staphylinids in the genus Batrisodes are described: B. (Declivodes) dorothae Ferro and Carlton from Feliciana Preserve, Louisiana; and B. (Babnormodes) spretoides Ferro and Carlton from Great Smoky Mountains National Park, Tennessee. They differ from all other described Batrisodes species in secondary male characters, especially details of the frontal region of the head. These two species bring the total diversity of the genus in North America to 88 species. Specimens were imaged using light microscopy, scanning electron microscopy, and X-ray microtomography (micro-CT) techniques. Utility of multiple imaging techniques, especially micro-CT, is discussed.
0379
Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae)
(2014)
The North American species of the genus Eusphalerum Kraatz (Coleoptera, Staphylinidae, Omaliinae) are revised. The taxonomic history, natural history, geographical distribution of the genus, characters, species groups, diversity, and biogeography of North American species are presented. Two main phylogenetic lineages and 13 species groups are provisionally recognized. The following new synonymies are proposed: Eusphalerum farrarae (Hatch, 1944) = E. lawrencei Hatch, 1957; Eusphalerum californicum (Fauvel, 1878) = E. atriventre (Casey, 1894), = E. nigerrimum (Casey, 1894), = E. gilvipenne (Casey, 1894), = E. dichroum (Fall, 1922), = E. bonnelli (Hatch, 1944), = E. lunae Hatch, 1957; Eusphalerum fraternum (Casey, 1894) = E. minskae (Hatch, 1944); Eusphalerum rugulosum (Mäklin, 1853) = E. grayae (Hatch, 1944); Eusphalerum orientale (Bernhauer, 1912) = E. frosti (Bernhauer, 1928). The following lectotypes are designated: E. subangulatum (Casey), E. californicum (Fauvel), E. gilvipenne (Casey), E. diversicolle (Casey), E. convexum (Fauvel), E. fraternum (Casey), E. horni (Fauvel), E. orientale (Bernhauer), E. pothos (Mannerheim), and E. punctatum (Casey). The following new species are described: Eusphalerum pilosum (California); E. klimaszewskii (British Columbia); E. chatzimanolisi (California); E. carolinensis (Kentucky, Missouri, North Carolina, Ohio, Pennsylvania, Tennessee, Virginia); E. caterinoi (California); E. luteipes (California); E. thayeranum (Alberta, British Columbia, Idaho, Indiana (doubtful record), Oregon, Washington); E. margaretae (Tennessee); E. newtoni (British Columbia, Oregon, California); E. parvispiculum (California, Oregon); E. uncinatum (British Columbia, California, Oregon, Washington). Eusphalerum lapponicum (Mannerheim, 1830) is excluded from the North American fauna. The following new combination is proposed: Xylodromus segmentarius (Mäklin, 1852: 322) (ex Omalium), wrongly attributed to Eusphalerum in the literature. Omalium marginatum Say, 1832 is considered a doubtful species, probably not Eusphalerum. A key to the 27 recognized North American Eusphalerum species and a catalog of the species are provided.
0378
Past concepts and synonymies of Anadenobolus monilicornis (Porat, 1876) (Spirobolida: Rhinocricidae), including the implied synonymy of Rhinocricus ectus Chamberlin, 1920, are consolidated into a formal account with the fi rst illustrations of the holotype. Prior to 1492, A. monilicornis was probably indigenous to an unknown number of southern Antillean islands, but through modern commerce, man has introduced it to Florida, Bermuda, Barbados, the Cayman Islands, and Jamaica, and probably repeatedly (re)introduced conspecifi c material to all the Lesser Antilles, resulting in subcontinuous gene pool mixing and reticulate evolution. A broad species concept is necessary to encompass the multitudinous variants, some of which have been recognized as species; only one true Caribbean species of Anadenobolus Silvestri, 1897, may exist, for which arboreus (Saussure, 1859) is the oldest name. The distribution of A. monilicornis presently extends from Bermuda and southern coastal Florida through the Greater and Lesser Antilles (excepting Cuba) to eastern coastal Venezuela and central Suriname, with outlier populations in Jamaica, the Cayman Islands, and Tampa Bay and the eastern Floridian panhandle; excepting Barbados, the indigenous range may have extended from Hispaniola through the same area. Introductions into Manitoba, Canada, and North Carolina, USA, have not yielded viable populations. Localities are newly recorded from St. Thomas, US Virgin Islands.
0377
Distributional records of the 71 tiger beetle species and subspecies known for Argentina are given together with a key and habitus photos (Coleoptera: Cicindelidae). Brasiella (Brasiella) cuyabaensis Mandl, 1970 and Brasiella (Gaymara) rotundatodilatata (Horn, 1925) are reported as new for the fauna of Argentina. Brasiella (Brasiella) stamatovi (Sumlin, 1979) is transferred to Cylindera (Plectographa), becoming C. (P.) stamatovi (Sumlin), new combination. Cylindera (Plectographa) siccalacicola (Sumlin, 1979) is placed into synonymy under Cylindera (Plectographa) hassenteufeli (Mandl, 1960), new synonymy. A lectotype is designated for Cylindera (Plectographa) patagonica bergiana (Horn, 1895).
0376
Five new species of Cerambycidae (Coleoptera) from Peru and Bolivia, and two new records for Peru
(2014)
The following four new species of Cerambycidae are described from Bolivia: Chrysoprasis imitatrix (Heteropsini); Carneades vigneaulti (Colobotheini); Colobothea larriveei (Colobotheini); Colobothea boliviana (Colobotheini). Esthlogena (Pseudotaxia) bella (Pteropliini) is described from Peru. A key to species of Carneades Bates, 1869 is provided. The other new species are included in previously published keys. Additionally, two new country records are reported for the fauna of Peru.
0375
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0372
A new species of Nephus Mulsant (Coleoptera: Coccinellidae) from southern Florida is described: Nephus (Nephus) alyssae. This represents the fi rst member of the subgenus Nephus reported in the southeastern United States. Florida species previously placed in the genus Nephus, now placed in Scymnobius Casey, are compared with the new species.
0371
A newly discovered population of Xystocheir brachymacris Shelley, 1996 (Polydesmida: Xystodesmidae: Xystocheirini), in Placer County (Co.), California, exhibits an unusual grayish-black color dorsally with mottled, ovoid patches at paranotal bases; it cons titutes northern generic and specifi c range extensions of ~28.4 km (17.6 mi). The gonopods differ from those in the El Dorado Co. population in having shorter/acuminate prefemoral processes and blade-like, rather than spatulate, processes “B” that angle away from the solenomere instead of overhanging it. Additionally, a strong distomedial prefemoral lobe, absent from the El Dorado population, arises from the stem in Placer Co. males. Authorship of Xystocheirini is properly attributed to Hoffman, 1980.
0370
The rediscovery of an older available name threatens the stability of the long accepted name of Strategus oblongus (Palisot de Beauvois, 1807) (Coleoptera: Scarabaeidae) from Hispaniola. Using Article 23.9 of the International Code of Zoological Nomenclature, Scarabaeus monoceros Nicolson, 1776 is designated a nomen oblitum to maintain nomenclatural stability while its junior synonym, Scarabaeus oblongus Palisot de Beauvois, 1807, is designated a nomen protectum.
0369
Multiple sources of confusion surround the identity of Ochodaeus grandiceps Fairmaire, 1897, from Sichuan, China (Coleoptera: Ochodaeidae). Herein the type specimen is illustrated to solve these taxonomic issues. Examination of the holotype indicates that the species must be transferred to the genus Nothochodaeus Nikolajev, 2005, resulting in a new combination. The genus Mimochodaeus Nikolajev, 2009, based on a misidentifi cation of O. grandiceps as its purported type species, is discussed. Finally, Cuban specimens had been treated erroneously as belonging to O. grandiceps, and so the species had remained undescribed. A new species, Parochodaeus perdidus, is described to accommodate them.
0368
A summary of the milliped faunas of Pakistan, Bangladesh, and Kashmir (Arthropoda: Diplopoda)
(2014)
Three female callipodidan samples from northern Pakistan are assigned to Bollmania kohalana (Attems, 1936) (Caspiopetalidae), the only ordinal representative documented from the country; a new record of Kaschmiriosoma loebli Jeekel, 2003 (Polydesmida: Paradoxosomatidae), is also provided. Localities are summarized for the 14 Pakistani, 6 Kashmirian, and 5 Bangladeshi diplopods. The last include one unidentifi able female of Zephronia Gray, 1832 (Sphaerotheriida: Zephroniidae), and two adventive species, Trachyjulus calvus (Pocock, 1893) (Spirostreptida: Cambalopsidae) and Asiomorpha coarctata (Saussure, 1860) (Polydesmida: Paradoxosomatidae); all constitute new country records. Two obscurely documented Bangladeshi diplopods are Gonoplectus cautus (Attems, 1936) (Spirostreptida: Harpagophoridae), and Trichopeltis watsoni Pocock, 1895 (Polydesmida: Cryptodesmidae). The Pakistani polydesmidan, Quasidesmus puschtun Golovatch, 1991, is transferred from Pyrgodesmidae to Cryptodesmidae.
0367
A gomphid male from west-central Wisconsin (Eau Claire County, North Fork Eau Claire River, 11 June 1994, K. J. Tennessen leg) with characters that are intermediate between Ophiogomphus carolus Needham, 1897 and Ophiogomphus rupinsulensis (Walsh), 1862 is described and illustrated. The specimen appears to be a hybrid based on intermediate character states of 1) color pattern (slightly closer to O. carolus), 2) hamule morphology (shaped slightly more like those of O. carolus), and 3) anal appendage morphology (slightly more like those of O. rupinsulensis).
0366
An adventive female Julidae (Julida), discovered in a moist, grassy depression in the Peninsula de Brunswick south of Punta Arenas, Chile, and assigned to Cylindroiulus Verhoeff, 1894, is the fi rst vouchered milliped from southern Patagonia. The southernmost milliped ever collected in Chile, South America, and the Western Hemisphere, it may also constitute the southernmost in the world as the site is only ~1,176 km (735 mi) northwest of the Antarctic Peninsula. Records are consolidated of the two families, three genera, and fi ve species of this Holarctic order that are known from South America. They are documented from Argentina, Chile, and southern Peru and Brazil; three species are known from the Juan Fernandez Islands.
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Three new species of Pselnophorus are described from the Nearctic region. Pselnophorus chihuahuaensis Matthews, Gielis, and Watkins, Pselnophorus hodgesi Matthews, Gielis, and Watkins, and Pselnophorus kutisi Matthews, Gielis, and Watkins, are described and distinguished from the only previously named Nearctic congener Pselnophorus belfragei (Fish). Illustrations of the adults and male and female genitalia are provided along with a key to males.
0363
Eleven Neotropical species of Laemophloeus Dejean with antennal clubs composed of three antennomeres are reviewed, diagnosed, and illustrated. Six of the species are described as new: L. capitesculptus Thomas, n. sp., L. corporeflavus Thomas, n. sp., L. dozieri Thomas, n. sp., L. insulatestudinorum Thomas, n. sp., L. planaclavatus Thomas, n. sp., and L. taurus Thomas, n. sp. Four new synonymies are proposed: L. catharinensis Kessel (=L. incisus Sharp), new synonym; L. similans Kessel (=L. incisus Sharp), new synonym; L. distinguendus Sharp (=L. megacephalus Grouvelle), new synonym, and L. chevrolati Grouvelle (=L. lecontei Grouvelle), new synonym. A key to the species is provided.
0362
The crape myrtle aphid Sarucallis kahawaluokalani (Kirkaldy) (Hemiptera: Aphididae) collected on Lagerstroemia indica (Lythraceae) is herein reported as a new invasive species in the city of Palmira, State of Valle del Cauca, and on San Andres island, in the State of San Andres, Old Providence and Santa Catalina, Colombia. The species is illustrated and diagnosed. A brief review of recent invasive species in Colombia, i.e., Ceroplastes rubens Maskell (Hemiptera: Coccidae), Crypticerya multicicatrices Kondo and Unruh (Hemiptera: Monophlebidae) [invasive on the island of San Andres], Diaphorina citri Kuwayama (Hemiptera: Liviidae), Harmonia axyridis (Pallas) (Coleoptera: Coccinellidae), Maconellicoccus hirsutus (Green) (Hemiptera: Pseudococcidae) and Singhiella simplex (Singh) (Hemiptera: Aleyrodidae); and other adventive (but non-invasive) species in Colombia, i.e., Anagyrus kamali Moursi (Hymenoptera: Encyrtidae), Gyranusoidea indica Shafee, Alam and Agarwal (Hymenoptera: Encyrtidae), and Tamarixia radiata (Waterston) (Hymenoptera: Eulophidae) is provided.
0361
New North American records of Pyraloidea (Lepidoptera: Crambidae, Pyralidae) from southern Florida
(2014)
We report six new North American records, one new state record, and one rare record of pyraloid moths from southern Florida, together with diagnostic characters for all taxa. We transfer Ennomosia Amsel from Spilomelinae to Glaphyriinae, Cangetta micralis (Hampson) n. comb. from Deuterophysa Warren, and Microthyris lelex (Cramer) n. comb. from Cyclocena Möschler. We revise Pseudocabotia Blanchard and Knudson rev. stat. to a subgenus of Ancylosis Zeller, with its type species A. (P.) balconiensis (Blanchard and Knudson) n. comb., and discuss the classifi cation of Cabotia Ragonot as a subgenus of Ancylosis.
0360
Two new species and a new genus of Cerambycidae are described from South America: Cotyclytus arriagadai sp. nov., from Bolivia; and Lembu dieguezi, gen. nov., sp. nov., from Paraguay. Orthomegas irroratus (Lameere, 1915) is redescribed, based on the second and third known specimens, and its distribution is expanded to include Ecuador. The male of Jamesia fuscofasciata Dillon and Dillon, 1952 is described and illustrated for the fi rst time, and the distribution of the species is expanded to Peru. Thirty-two new country records (twelve for Paraguay, fi fteen for Peru, two for Ecuador, three for Bolivia) and one new province record (Argentina) are presented.
0359
Four new species and one new genus of Cerambycinae are described from French Guiana: Sphagoeme premarginata sp. nov. and Atenizus apicalis sp. nov. (Oemini); Paraniophis signatipes gen. nov., sp. nov., and Niophis brusteli sp. nov. (Ectenessini). Three new country records for French Guiana are provided: Sphagoeme paraensis Martins, 1977, Atenizus simplex Bates, 1884, and Macroeme vittipennis (Melzer, 1934). All taxa are illustrated.
0358
0357
The endemic Floridian milliped genus, Floridobolus Causey, 1957, more closely related to tylobolinines in the western United States (US), Mexico, and Guatemala than syntopic spirobolines, is incorporated into Spirobolidae (Spirobolida: Spirobolidea). With taxonomic priority by one year, its monotypic family is reduced to Floridobolinae, n. stat., comprising Floridobolini and Tylobolini, n. stats., the counterpart to Spirobolinae, comprising Spirobolini and Aztecolini, n. tribe; relationships are Floridobolini + (Tylobolini + (Aztecolini + Spirobolini)). Like F. penneri Causey, 1957, 208 km (130 mi) to the south in the Lake Wales Ridge, Polk and Highlands counties (cos.), F. orini n. sp., inhabits “Big Scrub” environments in the Ocala National Forest, Marion Co. Biogeographic reconstructions, compatible with broader hypotheses on the class’ evolutionary history, indicate that, from a presumptive source area in northern Mexico where the subfamilies overlap, spirobolid stock penetrated the “proto-US” four times, once per tribe, before the Western Interior Seaway developed in the Cretaceous Period, Mesozoic Era. Three expansions headed northeastward into future “Appalachia,” from which taxa spread southward as the Seaway receded. Floridobolini, the fi rst invader, had to be in “proto-Georgia” and positioned to penetrate Florida when the sand dunes that comprise the “Central Highlands” emerged from the sea in the Oligocene (Cenozoic), ~25 mya. As sea levels rose and fell, the dunes fragmented into islands and the subcontinuous Floridobolus population was partitioned. The southernmost became F. penneri; F. orini inhabited a northern island; and a graduate student is investigating other insular remnants for additional species. Shortly after Floridobolini began spreading, Hiltonius/ Tylobolini arose and expanded both southward to Guatemala and northwestward to California; Tylobolus Cook, 1904, diverged in the latter area and dispersed northward to Washington and eastward to Utah/Arizona. The third invader, and the second to disperse northeastward, was Aztecolini, which probably eradicated Floridobolini from some of its established range and was partitioned into Mexican (Aztecolus Chamberlin, 1943) and US (Chicobolus Chamberlin, 1947) taxa by the Seaway. The fi nal invader, Spirobolini, dispersed northwestward and northeastward to both the Pacifi c and Atlantic coasts; instead of Trans-Beringia, we prefer penetration of the Asian part of “Asiamerica,” when it temporarily formed during the Cretaceous, to explain the Mongolian fossil genus, Gobiulus Dzik, 1975, herein assigned to Tylobolini, and the occurrence of Spirobolus Brandt, 1833, in China and Taiwan today. In the east, Narceus Rafi nesque, 1820, spread across Appalachia, eradicated most remaining populations of Floridobolus and Chicobolus, and expanded to Maine and Québec after retreat of the Wisconsin glaciation. Chicobolus and Narceus also penetrated earliest Florida; the former established itself in the Central Highlands, spread through the widening peninsula as sea levels fell, and remained on insular refugia when waters rose. Apparently fueled by the different Floridian environments, Narceus underwent time-consuming speciation; consequently, Floridobolus and Chicobolus still survive on the peninsula, and an allopatric population of the latter inhabits coastal South Carolina. However, N. gordanus (Chamberlin, 1943) occurs syntopically with both in peninsular Florida and may be actively eradicating them from their last stronghold. Trigoniulus niger, takahasii, and segmentatus, all by Takakuwa, 1940, are removed from Spirobolidae and returned toTrigoniulidae (Trigoniulidea). New records in the Appendix include the fi rst of Aztecolus from Durango and Jalisco, Mexico.