Insecta Mundi
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746
An abundance-based checklist of eastern Buprestidae (Coleoptera) was compiled from collection records from invasive insect surveys conducted during 2010-2018. Reported are 111 species in 17 genera based on 33,047 specimens examined from 10 states. Sixty-three new state records in nine states are reported. Collection date ranges by month for each state are provided.
757
The scale insects (Hemiptera: Coccomorpha) are one of the most successful groups of plant-feeding arthropods. Most species of shade trees, fruit trees and ornamental shrubs are subject to scale insect attacks. Based on the review of the literature and survey results, the host plant list of the scale insects in South Korea was developed and updated. Herein, an updated list of 253 species of host plants in 71 families associated with scale insects is provided and 162 species in 15 scale insect families are listed.
747
The adventive mole cricket species (Orthoptera: Gryllotalpidae) in Hawaii is apparently Gryllotalpa krishnani Arun Prasanna et al., 2012. The adventive was long thought to be Gryllotalpa africana Palisot de Beauvois, 1805. From 1983 until 2008 it seemed that Gryllotalpa orientalis Burmeister, 1839 might be the adventive’s correct name. However, male genitalia and tegmental characters of Hawaiian specimens match features of G. krishnani and not G. orientalis.
831
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784
Reinstatement of Carposina ottawana Kearfott, 1907 (Lepidoptera: Carposinidae) as a valid species
(2020)
Carposina ottawana Kearfott, 1907 (Lepidoptera: Carposinidae), revised status, formerly considered a synonym of C. sasakii Matsumura, 1900, is returned to species status. Morphological features that separate the Asian species C. sasakii and C. niponensis Walsingham, 1900 from the North American C. ottawana are described and illustrated. A heuristic maximum likelihood (ML) analysis based on the mitochondrial gene cyto-chrome oxidase I (DNA barcode) further supports C. ottawana and C. sasakii as distinct taxa.
751
Cyrtinus pygmaeus (Haldeman, 1847) (Coleoptera: Cerambycidae: Lamiinae) is redescribed and newly recorded from Mexico. The female of Decarthria stephensii Hope, 1834 is also redescribed, the number of specimens in the type series is corrected, as is the depository of the types, and the species is newly recorded from Dominica. Two new species of Cyrtinus LeConte, 1852 are described from Mexico: C. fisheri Wappes, Santos-Silva and Nascimento; and C. howdeni Wappes, Santos-Silva and Nascimento. A key to species of Decarthria Hope, 1834 (adapted from an earlier key to Cyrtinini) is provided.
753
Although well studied in the Afrotropical Region, the genus Cosmorrhyncha Meyrick, 1913 (Lepidoptera: Tortricidae: Olethreutinae), has received little attention in the New World, where it apparently is restricted to the Neotropics from Guatemala south to Paraguay. Seven species are recognized, five of which are described as new: C. tonsana (Walker, 1863) (Type locality: Brazil); C. ocelliferana (Walker, 1863) (TL: Brazil); C. landryi Brown and Razowski, sp. n. (TL: French Guiana); C. parintina Brown and Razowski, sp. n. (TL: Brazil); C. macrospina Brown and Razowski, sp. n. (TL: Brazil); C. albistrigulana Brown and Razowski, sp. n. (TL: Costa Rica); and C. osana Brown and Razowski, sp. n. (TL: Costa Rica). Our circumscription of C. ocelliferana is rather broad and most likely encompasses a species complex rather than a single entity. Larvae of C. tonsana have been reared from Picramnia latifolia Tul. (Picramniaceae), Dialium guianense (Aubl.) Sandwith (Fabaceae), and Machaerium seemannii Benth. ex Seem. (Fabaceae) in Costa Rica; and those of C. albistrigulana from Dialium guianense (Fabaceae).
793
The New World genus Axina Kirby (Coleoptera: Cleridae) is revised for the first time. Thirty-two new species are described: Axina acutipennis, A. adelosa, A. atmis, A. bahia, A. bella, A. brunnea, A. chiasta, A. furcula, A. heveli, A. ignota, A. klisis, A. latilinea, A. lobispinula, A. luzia, A. macilenta, A. megaspina, A.minas, A. ochra, A. oligocheia, A. ordinis, A. orcastomata, A. pallidioccabus, A. phallospina, A. piperata, A. pollex, A. polycaula, A. rio, A. schenklingi, A. spina, A. trinalis, A. villa, and A. vista. The 19 previously described species are: Axina analis Kirby, A. apicalis Pic, A. basalis Schenkling, A. bifasciata (Chevrolat), A. centrimaculata Schenkling, A. conspicua Schenkling, A. diversesignata Pic, A. equestris (Schenkling), A. fasciata Kirsch, A. fortipes Pic, A. lateralis Pic, A. longevittata Pic, A. munda Schenkling, A. nigrifrons Schenkling, A.parcepunctata Schenkling, A. picta Schenkling, A. plagiata Schenkling, A. proxima (Chevrolat), and A. sexmaculata Spinola. Two species Priocera equestris Schenkling and Priocera proxima Chevrolat, are transferred into the genus Axina becoming new combinations: Axina equestris (Schenkling) and Axina proxima (Chevrolat). Two names, Priocera podagrica Schenkling, 1900, and P. podagrica variety pygmaea Schenkling, 1902, are new synonymies of Priocera proxima Chevrolat, 1876. Lectotypes are here designated for nine species: Axina basalis Schenkling, 1900; Axina centrimaculata Schenkling, 1900; Axina conspicua Schenkling, 1900; Axina diversesignata Pic, 1946; Axina munda Schenkling, 1900; Axina nigrifrons Schenkling, 1906; Axina parcepunctata Schenkling, 1900; Axina picta Schenkling, 1907; and Axina plagiata Schenkling, 1900. It is proposed that Axina species are predators of lignicolous insects, particularly bark beetles. The species of Axina can be classified into eight species groups and a theory of their phylogenetic relationships is proposed via WINCLADA in conjunction with NONA. Of the 51 species that now comprise Axina, only one traversed the Panamanian portal before the Colombian Andes reached their modern altitudes. This work includes a generic-level morphological analysis, brief treatise of natural history, key to species, comments about Axinazoogeography, and hypotheses of species-group phylogeny.
788
Psyllids are an economically important group of insects. Several species are serious emerging pests with regulatory significance. About 20 adventive species have been discovered in Florida in the past 20 years, including several pests. Additionally, five species new to science have been found. We provide an annotated checklist of Florida species with taxonomic information and identification tools, including keys to Florida genera and known species. Seventy species of Psylloidea currently are reported from Florida. Forty-one are native to Florida, with 12 endemic to the state. Twenty are adventive, the majority being from the Neotropics. One was introduced deliberately for biological control, seven represent temporary populations (eradicated, reared in quarantine), and one is a dubious record. Craspedolepta euthamiae Burckhardt and Halbert, new species, Katacephala wineriterae Burckhardt and Halbert, new species, Pseudophacopteron gumbolimbo Burckhardt and Halbert, new species, Nothotrioza longipedis Burckhardt and Halbert, new species, and Trioza myresae Burckhardt and Halbert, new species are described from Florida and are native endemic species. Aphalara persicaria Caldwell is redescribed and separated from similar species. Aphalara persicaria var. cubana Caldwell is confirmed as a junior synonym of Aphalara persicaria. The Florida records of Craspedolepta spp. are revisited and revised, including Craspedolepta euthamiae Burckhardt and Halbert, new species. Bactericera nigrilla (Crawford), new combination, revived status is recognized from Florida, redescribed, and distinguished from similar species. Rhinopsylla caldwelli Tuthill is transferred to Kuwayama Crawford and becomes Kuwayama caldwelli (Tuthill), new combination.Trioza maritima Tuthill is transferred to Leuronota Crawford and becomes Leuronota maritima (Tuthill), new combination. Species of Bactericera Puton on Salix L. (Salicaceae) in North America are reviewed. Bactericera flori (Crawford), new combination, new status. is determined to be the correct name for Trioza assimilis Crawford nec Flor (= Trioza flori Crawford, replacement name, = Trioza pomonae Aulmann, replacement name), and Trioza dubia Patch, new synonym. Lectotypes are designated for Trioza marginata Crawford, Trioza minuta Crawford, Trioza minuta similis Crawford, and Trioza nigra Crawford.
752
767
Several taxonomic and nomenclatural issues are reviewed, clarified, and resolved for multiple genera of the Erotylinae (Coleoptera: Erotylidae). Generic-group names discussed: Brachymerus Dejean, 1836, Cypherotylus Crotch, 1873, Cytorea Laporte, 1840, Erotylus Fabricius, 1775, Eudaemonius Lewis, 1887, Eutriplax Lewis,1887, Gibbifer Voet, 1806, Neobarytopus Alvarenga, 1965, Neomorphoides Alvarenga, 1977, Ogcotriplax Heller,1920, Paratritoma Gorham, 1888, Platichna Thomson, 1863, Pseudochrysomela Voet, 1806, Pseudotriplax Heller,1920, Triplax Herbst, 1793, Tritomapara Alvarenga, 1970, Typocephalus Hope,1841, and Xestus Wollaston, 1864. Reviewing these issues resulted in a several nomenclatural actions. Eutriplax Lewis,1887,was found to be an unnecessary replacement name for Eudaemonius Lewis,1887.The genus name is reverted to Eudaemonius,
resulting in one new combination: Eudaemonius quinquepustulatus (Li and Ren, 2006).
The Neotropical Tritomapara Alvarenga,1970,was found to be a new objective synonym of Paratritoma Gorham,1888, which is a synonym of Triplax Herbst,1793,leading to the following eight new combinations:
Triplax atricaudata (Kuhnt,1910),Triplax brasiliensis (Guérin,1946),Triplax bruchi (Kuhnt,1910),Triplax caduca (Gorham,1888),Triplax dimidiata (Gorham,1888),Triplax melanoderes (Kuhnt,1910),Triplax triplacoides
(Crotch,1876), and Triplax vivida (Gorham,1888). Erotylus tibialis Duponchel, 1825, is recognized as the valid type species for Brachymerus Dejean 1836, which moves the name Brachymerus to a different genus-group taxon and renders Neomorphoides Alvarenga, 1977, a new synonym. This revalidates Neobarytopus Alvarenga, 1965, as originally proposed. These genus-group names are presently subgenera in Iphiclus Dejean, 1836, and the move creates 23 new combinations in Iphiclus(Brachymerus) Dejean, 1836: I. (B.) amazonus (Crotch, 1876), I. (B.) atriventris (Mader,1943), I. (B.) bicolor(Lacordaire,1842), I. (B.) clavicornis (Olivier,1792), I. (B.) columbiae (Crotch,1876), I. (B.) costaricensis (Mader,1943), I. (B.) disconigrum (Mader,1942), I. (B.) dorsonotatus (Lacordaire, 1842), I. (B.) fulviventris (Gorham,1888), I. (B.) humeropictus (Mader,1943), I. (B.) lateripunctatus (Crotch,1876), I. (B.) melanopus (Gorham,1888), I. (B.) neglectus (Guérin,1956), I. (B.) nigritarsis (Mader,1942), I. (B.) nigriventris (Crotch, 1876), I. (B.) nigropectus (Mader,1942), I. (B.) posticenigrum (Mader,1942), I. (B.) pyrrhocephalus (Erichson,1847), I. (B.) rubripennis (Lacordaire,1842), I. (B.) signaticollis (Kuhnt,1910), I. (B.) simplex (Lacordaire,1842), I. (B.) spilotus (Gorham,1888), I. (B.) tibialis (Duponchel,1825); and, 75 new combinations in Iphiclus (Neobarytopus) Alvarenga, 1965: I. (N.) adustus (Duponchel,1825), I. (N.) alboniger (Guérin,1956), I. (N.) amictus (Erichson,1847), I. (N.) andicola (Kirsch,1867), I. (N.) assequens (Mader,1942), I. (N.) bajulus (Lacordaire,1842), I. (N.) batesi (Gorham, 1889),I. (N.) bellulus (Lacordaire,1842), I. (N.) bicinctus (Olivier,1807), I. (N.) bistrifoliatus (Gorham,1889), I. (N.) bizonatus (Crotch,1876), I. (N.) bremei (Guérin-Méneville,1841), I. (N.) brongniarti (Lacordaire,1842), I. (N.) brunneostriolatus (Kuhnt, 1910), I. (N.) cerasinus (Lacordaire,1842), I. (N.) conformis (Lacordaire,1842), I. (N.)distinctus (Duponchel, 1825), I. (N.) divisus (Guérin,1956), I. (N.) dorsalis (Olivier,1792), I. (N.) eburneus (Crotch,1876), I. (N.) elegans (Mader,1942), I. (N.) epipleuralis (Crotch,1876), I. (N.) erichsoni (Lacordaire,1842), I. (N.) flavofasciatus (Duponchel,1825), I. (N.) flavosignatus (Duponchel,1825), I. (N.) fragmentatus (Gorham,1888), I.(N.) friedei (Mader,1938), I. (N.) geometra (Lacordaire,1842), I. (N.) hebriacus (Lacordaire,1842), I. (N.) hexastictus (Crotch,1876), I. (N.) incas (Gorham,1889), I. (N.) iris (Guérin,1956), I. (N.) jacinthoi (Alvarenga,1977), I.(N.) laceratus (Mader,1938), I. (N.) lugens (Lacordaire,1842), I. (N.) lunaris (Guérin,1956), I. (N.) luteozonatus (Crotch, 1876), I. (N.) miles (Mader,1942), I. (N.) mirus (Mader,1942), I. (N.) musicalis (Lacordaire,1842), I. (N.) neophyta (Lacordaire, 1842), I. (N.) nigripennis (Demay,1838), I. (N.) nigropictus (Lacordaire,1842), I. (N.) nigrofasciatus (Mader,1942), I. (N.) nitidulus (Oliver,1807), I. (N.) obsoletesignatus (Crotch, 1876), I. (N.) octoguttatus (Olivier,1807), I. (N.) octopustulatus (Guérin,1956), I. (N.) odyneroides (Crotch, 1876), I. (N.) ornatus (Kuhnt,1909), I. (N.) pantherinus (Kuhnt, 1909), I. (N.) pauper (Guérin,1956), I. (N.) peraffinis (Crotch,1876), I. (N.) perplexus (Mader,1942), I. (N.) peruvianus (Mader,1942), I. (N.) planipennis (Kuhnt,1909), I. (N.) puncticollis (Kirsch,1876), I. (N.) quadrifasciatus (Kirsch,1865), I. (N.) quinquefasciatus (Lacordaire,1842), I. (N.) ramosus (Olivier,1807), I. (N.) regularis (Erichson, 1848), I. (N.) rhomboidalis (Guérin,1956), I. (N.) rufipennis (Panzer,1798), I. (N.) salamandra (Erichson,1847), I. (N.) spectabilis (Lacordaire,1842), I. (N.) stramineus (Lacordaire,1842), I. (N.) subsanguineus (Crotch,1876), I. (N.) superbus (Mader,1942), I. (N.) tigrinatus (Guérin,1956), I. (N.) tricinctus (Duponchel,1825), I. (N.) trifasciatus (Olivier,1807), I. (N.) tripartitus (Lacordaire,1842), I. (N.) ucayalensis (Gorham,1889), I. (N.) venezuelae (Crotch, 1876), I. (N.) westwoodi (Guérin-Méneville,1841).
The works of Voet (1766–1778,1806) do not follow binominal nomenclature and are therefore unavailable by the International Code of Zoological Nomenclature, Article 11.4.Thus, Voet’s (1806) generic names “Pseudochrysomela” and “Gibbifer”, and the species names proposed in each, are unavailable. Removing them from nomenclatural considerations resulted in the following nomenclatural acts: the resurrection of Erotylus rufipennis Panzer,1798, now Iphiclus (Neobarytopus) rufipennis (Panzer) new combination; the proposal of a new name, Iphiclus (Brachymerus) fabricii Skelley for Erotylus rufipennis Fabricius, 1801, not Erotylus rufipennis Panzer, 1798; new combinations for the two species, Cypherotylus adrianae (Alvarenga, 1976) and Cypherotylus borgmeieri (Alvarenga, 1976); and revalidated status for the five species names, Erotylus variegatus Fabricius,
1781, Barytopus gronovii (Herbst, 1783), Prepopharus notatus (Olivier, 1792), Iphiclus (Iphiclus) sedecimguttatus (Olivier, 1792), and Cypherotylus duponcheli Arrow, 1937.
839
Glaresis franki Keller and Skelley new species and Glaresis thomasi Keller and Skelley new species (Coleoptera: Scarabaeoidea: Glaresidae) are described and illustrated. They represent the first record of the family for the West Indies. Both species are placed into the phoenicis species group. A key to the two West Indies species is presented.
768
Arcola malloi (Pastrana, 1961) is a junior subjective synonym of Macrorrhinia endonephele (Hampson, 1918) syn. nov. (Lepidoptera: Pyralidae). The species is a biological control agent introduced in United States and Australia to control alligatorweed, Alternanthera philoxeroides (Mart.) Griseb. (Amaranthaceae). The synonymy is recognized by comparison of type specimens, genitalic dissections, and DNA COI barcoding. Vogtia Pastrana, 1961 syn. nov. and Arcola Shaffer, 1995 syn. nov. are synonymized with Macrorrhinia Ragonot, 1887. Macror-rhinia megajuxta (Neunzig and Goodson, 1992) comb. nov. is transferred from Ocala Hulst, 1892. Lectotypes are designated for Divitiaca ochrella Barnes and McDunnough, 1913, and Divitiaca simulella Barnes and Mc-Dunnough, 1913.
783
791
The click beetle genus Carlota Arias-Bohart (Coleoptera: Elateridae: Agrypninae: Agrypnini) was considered as a junior synonym of Candanius Hayek recently. However, there are deep morphological differences between these genera which justify the validity of Carlota. The morphology of this genus was re-examined in detail and based on the short and shallow antennal grooves, strongly serrate antennae from antennomeres 3 through 10, subquadrate pronotum with four distinct subcircular depressions, and straight prosternal process not bent dorsally, I resurrect the genus Carlota from synonymy.
829
We honor the life and accomplishments of Michael C. Thomas with a short narrative of his professional life along with appendices listing his scientific artwork, bibliography and patronyms. This paper is the first of a Festschrift with contributed remembrances and separate papers honoring him with additional patronyms.
763
In 2017, a new project was begun to assess the biodiversity of national parks and forest reserves in the Republic of Panama. Designated “Proyecto Sistema de Producción Sostenible Conservación de la Biodiversidad (PSPSCB)”, this project is managed by Panama’s Ministerio de Ambiente. The first park sampled in 2017 was Omar Torrijos Herrera National Park (OTHNP). Trichoptera (Insecta) were collected at four locations using both Malaise traps and UV light traps. The rugged terrain and lack of access in this remote park limited the sampled area. Sampling included streams in both the Caribbean and Pacific drainages. Seven new species of microcaddisflies (Hydroptilidae: Alistotrichia coclensis Armitage and Harris, Cerasmatrichia akanthos Armitage and Harris, Metrichia corazones Armitage and Harris, Neotrichia espinosa Armitage and Harris, Neotrichia michaeli Armitage and Harris, Neotrichia pierpointorum Armitage and Harris, and Neotrichia yayas Armitage and Harris) and one new country record, Metrichia macrophallata Flint, were identified from this preliminary survey and are reported herein. Other recently described species are reported here for the first time outside of their type localities. Based on other areas more extensively sampled compared to this modest survey, many more new species and new country records await discovery in OTHNP as are reported herein. There are now 439 species distributed among 15 families and 55 genera known from Panama.
807
Cypherotylus dromedarius (Lacordaire, 1842) (Coleoptera: Erotylidae) was described from French Guiana. Information about its current distribution is scattered throughout the literature and the internet. Specimens of this species were recently observed and photographed in the Boquete area, Chiriqui Province, Panama, and constitute a new record for the country of this species. Subsequently, we located two specimens deposited in the National Reference Collection of the G. B. Fairchild Museum of Invertebrates, University of Panama, from the same locality in Chiriqui Province. We summarize the distribution of this species from scattered sources and provide five new records.
764
After publication of Armitage and Harris (2020), it was noticed that the wrong illustrations were used for the species Neotrichia espinosa Armitage and Harris (Fig. 9) and Neotrichia michaeli Armit-age and Harris (Fig. 10), resulting in images that did not correspond to the captions. The authors and the Insecta Mundi editorial staff apologize for this error. The intended versions of Figures 9 and 10 are reproduced here. Insecta Mundi has also released a revised version of the Armitage and Harris (2020) manuscript, with this error corrected. However, the revised version is not an official peer-reviewed article, and anyone wishing to reference the findings of Armitage and Harris (2020) should cite the original manuscript or this erratum.