Insecta Mundi
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Institute
- Extern (65)
0204
The American species belonging to the genera Alloblackburneus Bordat and Blackburneus Schmidt (Coleoptera: Scarabaeidae: Aphodiinae) are redescribed and figured. Seven new species are described: Alloblackburneus guadalajarae, A. ibanezbernali; Blackburneus amazonicus, B. sanfilippoi, B. surinamensis, B. teposcolulaensis, B. thomasi. The neotype of Scarabaeus rubeolus Palisot de Beauvois, 1809 is designated. The lectotype of Blackburneus argentinensis (Schmidt, 1909) and of Blackburneus laxepunctatus (Schmidt, 1910) are
designated. The following new combinations are proposed: Alloblackburneus aegrotus (Horn, 1870); Alloblackburneus cavidomus (Brown, 1927); Alloblackburneus cynomysi (Brown, 1927); Alloblackburneus fordi (Gordon, 1974); Alloblackburneus geomysi (Cartwright, 1939); Alloblackburneus lentus (Horn, 1870); Alloblackburneus rubeolus (Palisot de Beauvois, 1805); Alloblackburneus saylori (Hinton, 1934); Alloblackburneus tenuistriatus (Horn, 1887); Alloblackburneus troglodytes (Hubbard, 1894).
0086
0269
Systematic revision of the genus Orodaliscoides Schmidt, 1913 (Coleoptera: Scarabaeidae: Aphodiinae)
(2012)
The species belonging to the genus Orodaliscoides Schmidt (Coleoptera: Scarabaeidae: Aphodiinae) are redescribed and figured. The following new combinations are proposed: Orodaliscoides fimbripes (Brown, 1928) and Orodaliscoides giulianii (Gordon, 1977).
0383
Besides the two species at present known belonging to the genus Trichonotuloides Balthasar (T. glyptus (Bates) and T. latecrenatus (Bates)), two new Mexican species, T. alfonsinae and T. hansferyi, are herein described (Coleoptera: Scarabaeidae: Aphodiinae). The complete set of fi gures is supplied for all taxa herein dealt with.
590
A systematic redefinition of the species belonging to the genus Geomyphilus Gordon and Skelley, 2007 (Coleoptera: Scarabaeidae: Aphodiinae) of Mexico and neighboring countries is presented. The new species G. tuzincola of Mexico is described and figured. The new combination Coelotrachelus macgregori (Islas, 1955) is proposed.
791
The click beetle genus Carlota Arias-Bohart (Coleoptera: Elateridae: Agrypninae: Agrypnini) was considered as a junior synonym of Candanius Hayek recently. However, there are deep morphological differences between these genera which justify the validity of Carlota. The morphology of this genus was re-examined in detail and based on the short and shallow antennal grooves, strongly serrate antennae from antennomeres 3 through 10, subquadrate pronotum with four distinct subcircular depressions, and straight prosternal process not bent dorsally, I resurrect the genus Carlota from synonymy.
0310
The following genera of Leiodini (Coleoptera: Leiodidae: Leiodinae) of the continental United States and Canada are reviewed: Cyrtusa Erichson, with two species; Isoplastus Horn, with two species (one new); Liocyrtusa Daffner, with three species; Lionothus Brown, with five species (three new), and Zeadolopus Broun, with four species (all genera are in the “Cyrtusa genus group”) and Ecarinosphaerula Hatch, with one named species (in the “Leiodes genus group”). The new species are Isoplastus floridanus Peck and Cook of Florida; Lionothus bidentatus Peck and Cook of Texas and Oklahoma, Lionothus exiguus Peck and Cook of Florida and Texas, and Lionothus parvoculus Peck and Cook of Arizona and New Mexico. Bionomic data on the species are given, and complete known distributions are mapped.
0392
We clarify the taxonomy and nomenclature of several species-group names and the family-group name for the anticheirine scarabs (Scarabaeidae: Rutelinae: Rutelini), a group that includes nearly 550 species and subspecies in 44 genera and subgenera. These clarifications are necessary due to taxonomic and nomenclatural inaccuracies in recent revisionary works. We provide a list of genera included in the anticheirine scarabs sensu Soula. Numerous nomenclatural changes are necessary due to invalid type designations or misspellings: the valid type species of Aequatoria Arrow is Chlorota associata Waterhouse; the valid type species of Chlorota Burmeister is Rutela terminata Le Peletier de Saint-Fargeau and Audinet-Serville; the valid type species of Dorysthetus Blanchard is Macraspis chlorophana Burmeister; the valid type species of Anticheira Eschscholtz is Scarabaeus virens Drury. Changes to the type species of Anticheira result in new combinations: Anticheira virens virens (Drury), new combination, Anticheira virens bleuzeni (Soula), new combination and Anticheira virens jossi (Soula), new combination. Some anticheirine species names were not associated with species descriptions or type designations, making these names nomina nuda and unavailable: “Chlorota smithi”, “Macraspis cincta guatemalensis”, “Parachlorota equatoriana”, “Parachlorota pardoi”, “Pseudothyridium (Megathyridium) bousqueti”, “Theuremaripa rivae brasiliensis” and “Thyridium cupriventre blanchardi”. Dorysthaetus rufipennis Dejean is a nomen nudum and an unavailable name. We correct numerous misspellings from Soula’s five volumes on anticheirine scarabs and provide a list to avoid propagation of these errors by future researchers. Eighteen names are unavailable infrasubspecific names, and thirty-six of Soula’s names were misspelled. These names confuse and obscure ruteline systematics. Lastly, we provide a comprehensive list of infrasubspecific names in the anticheirine scarabs and we establish these as available or unavailable names.
921
Our expanded efforts in genomic sequencing to cover additional skipper butterfly (Lepidoptera: Hesperiidae) species and populations, including primary type specimens, call for taxonomic changes to restore monophyly and correct misidentifications by moving taxa between genera and proposing new names. Reconciliation between phenotypic characters and genomic trees suggests three new tribes, two new subtribes, 23 new genera, 17 new subgenera and 10 new species that are proposed here: Psolosini Grishin, new tribe (type genus Psolos Staudinger, 1889), Ismini Grishin, new tribe (type genus Isma Distant, 1886), Eetionini Grishin, new tribe (type genus Eetion de Nicéville, 1895), Orphina Grishin, new subtribe (type genus Orphe Godman, 1901), Carystoidina Grishin, new subtribe (type genus Carystoides Godman, 1901), Fulvatis Grishin, new genus (type species Telegonus fulvius Plötz, 1882), Adina Grishin, new genus (type species Nascus adrastor Mabille and Boullet, 1912), Ornilius Grishin, new genus (type species Ornilius rotundus Grishin, new species), Tolius Grishin, new genus (type species Antigonus tolimus Plötz, 1884), Lennia Grishin, new genus (type species Leona lena Evans, 1937), Trida Grishin, new genus (type species Cyclopides barberae Trimen, 1873), Noxys Grishin, new genus (type species Oxynthes viricuculla Hayward, 1951), Gracilata Grishin, new genus (type species Enosis quadrinotata Mabille, 1889), Hermio Grishin, new genus (type species Falga ? hermione Schaus, 1913), Eutus Grishin, new genus (type species Cobalus rastaca Schaus, 1902), Gufa Grishin, new genus (type species Phlebodes gulala Schaus, 1902), Godmia Grishin, new genus (type species Euroto chlorocephala Godman, 1900), Rhomba Grishin, new genus (type species Eutychide gertschi Bell, 1937), Rectava Grishin, new genus (type species Megistias ignarus Bell, 1932), Contrastia Grishin, new genus (type species Hesperia distigma Plötz, 1882), Mit Grishin, new genus (type species Mnasitheus badius Bell, 1930), Picova Grishin, new genus (type species Vorates steinbachi Bell, 1930), Lattus Grishin, new genus (type species Eutocus arabupuana Bell, 1932), Gubrus Grishin, new genus (type species Vehilius lugubris Lindsey, 1925), Koria Grishin, new genus (type species Hesperia kora Hewitson, 1877), Corta Grishin, new genus (type species Eutychide lycortas Godman, 1900), Calvetta Grishin, new genus (type species Hesperia calvina Hewitson, 1866), Oz Grishin, new genus (type species Astictopterus ozias Hewitson, 1878), Praxa Grishin, new subgenus (type species Nascus prax Evans, 1952), Bron Grishin, new subgenus (type species Papilio broteas Cramer, 1780), Turis Grishin, new subgenus (type species Pyrgus 1955, and Synale Mabille, 1904 of Carystus Hübner, [1819]. The following 20 genera are treated as junior subjective synonyms: Leucochitonea Wallengren, 1857 of Abantis Hopffer, 1855; Sapaea Plötz, 1879 and Netrobalane Mabille, 1903 of Caprona Wallengren, 1857; Parasovia Devyatkin, 1996 of Sebastonyma Watson, 1893; Pemara Eliot, 1978 of Oerane Elwes and Edwards, 1897; Ankola Evans, 1937 of Pardaleodes Butler, 1870; Arotis Mabille, 1904 of Mnaseas Godman, 1901; Chalcone Evans, 1955, Hansa Evans, 1955, and Propertius Evans, 1955 of Metrocles Godman, 1900; Jongiana O. Mielke and Casagrande, 2002 of Cobaloides Hayward, 1939; Pamba Evans, 1955 of Psoralis Mabille, 1904; Brownus Grishin, 2019 of Styriodes Schaus, 1913; Mnasilus Godman, 1900 of Papias Godman, 1900; Sucova Evans, 1955 of Mnasitheus Godman, 1900; Pyrrhocalles Mabille, 1904 and Asbolis Mabille, 1904 of Choranthus Scudder, 1872; Miltomiges Mabille, 1903 of Methionopsis Godman, 1901; Sacrator Evans, 1955 of Thracides Hübner, [1819]; and Lychnuchoides Godman, 1901 of Perichares Scudder, 1872. Arunena Swinhoe, 1919 is a junior subjective synonym of Stimula de Nicéville, 1898 (not of Koruthaialos Watson, 1893). The following 27 names are species-level taxa (some in new combinations) reinstated from synonymy: Salantoia gildo (Mabille, 1888) (not Salatis cebrenus (Cramer, 1777)), Bungalotis corentinus (Plötz, 1882) (not Bungalotis midas (Cramer, 1775)), Telegonus cretellus (Herrich-Schäffer, 1869) (not Telegonus cassander (Fabricius, 1793)), Santa palica (Mabille, 1888) (not Chiothion asychis (Stoll, 1780)), Camptopleura cincta Mabille and Boullet, 1917 (not Camptopleura auxo (Möschler, 1879)), Camptopleura orsus (Mabille, 1889) (not Nisoniades mimas (Cramer, 1775)), Metron voranus (Mabille, 1891) and Metron fasciata (Möschler, 1877) (not Metron zimra (Hewitson, 1877)), Limochores catahorma (Dyar, 1916) (not Limochores pupillus (Plötz, 1882)), Pares viridiceps (Mabille, 1889) (not Thoon modius (Mabille, 1889)), Tigasis wellingi (Freeman, 1969) (not Tigasis arita (Schaus, 1902)), Rectava sobrinus (Schaus, 1902) (not Papias phainis Godman, 1900), Nastra subsordida (Mabille, 1891) (not Adlerodea asema (Mabille, 1891), previously in Eutychide Godman, 1900), Lerema pattenii Scudder, 1872 (not Lerema accius (J. E. Smith, 1797)), Lerema (Morys) ancus (Möschler, 1879) (not Cymaenes tripunctus theogenis (Capronnier, 1874)), Cobalopsis zetus (Bell, 1942) (not Cobalopsis nero (Herrich-Schäffer, 1869)), Lerema (Geia) etelka (Schaus, 1902) (not Lerema (Geia) geisa (Möschler, 1879), previously in Morys Godman, 1900), Cymaenes isus (Godman, 1900) (not Cymaenes trebius (Mabille, 1891)), Vehilius labdacus (Godman, 1900) (not Vehilius inca (Scudder, 1872)), Papias amyrna (Mabille, 1891) (not Papias allubita (Butler, 1877), previously in Mnasilus Godman, 1900), Papias integra (Mabille, 1891) (not Papias subcostulata (Herrich-Schäffer, 1870)), Metiscus atheas Godman, 1900 (not Hesperia achelous Plötz, 1882), Dion agassus (Mabille, 1891) (not Dion uza (Hewitson, 1877), previously in Enosis Mabille, 1889), Picova incompta (Hayward, 1942) (not Lerema (Morys) micythus (Godman, 1900), previously in Morys Godman, 1900), Lucida melitaea (Draudt, 1923) (not Lucida lucia (Capronnier, 1874)), Methionopsis modestus Godman, 1901 (not Methionopsis ina (Plötz, 1882)), and Thargella (Volus) volasus (Godman, 1901) (not Eutocus facilis (Plötz, 1884)). The following 57 taxa are elevated from subspecies to species, new status (some in new combinations): Dyscophellus doriscus (Hewitson, 1867) (not Dyscophellus porcius (C. Felder and R. Felder, 1862), Phocides vida (A. Butler, 1872) (not Phocides urania (Westwood, 1852)), Tagiades (Daimio) ceylonica Evans, 1932 (not Tagiades litigiosa Möschler, 1878), Tagiades (Daimio) tubulus Fruhstorfer, 1910 (not Tagiades sambavana Elwes and Edwards, 1897), Tagiades (Daimio) kina Evans, 1934, Tagiades (Daimio) sheba Evans, 1934, Tagiades (Daimio) martinus Plötz, 1884, Tagiades (Daimio) sem Mabille, 1883, and Tagiades (Daimio) neira Plötz, 1885 (not Tagiades trebellius (Hopffer, 1874)), Tagiades (Daimio) korela Mabille, 1891 and Tagiades (Daimio) presbyter Butler, 1882 (not Tagiades nestus (C. Felder, 1860)), Tagiades obscurus Mabille, 1876, Tagiades ravi (Moore, [1866]), Tagiades atticus (Fabricius, 1793), Tagiades titus Plötz, 1884, Tagiades janetta Butler, 1870, Tagiades inconspicua Rothschild, 1915, and Tagiades hovia Swinhoe, 1904 (not Tagiades japetus (Stoll, [1781])), Tagiades silvia Evans, 1934 and Tagiades elegans Mabille, 1877 (not Tagiades gana (Moore, [1866])), Tapena bornea Evans, 1941 and Tapena minuscula Elwes and Edwards, 1897 (not Tapena thwaitesi Moore, [1881]), Darpa dealbata (Distant, 1886) (not Darpa pteria (Hewitson, 1868)), Perus manx (Evans, 1953) (not Perus minor (Schaus, 1902)), Canesia pallida (Röber, 1925) (not Carrhenes canescens (R. Felder, 1869)), Carrhenes conia Evans, 1953 (not Carrhenes fuscescens (Mabille, 1891)), Anisochoria extincta Hayward, 1933 and Anisochoria polysticta Mabille, 1876 (not Anisochoria pedaliodina (Butler, 1870)), Anisochoria verda Evans, 1953 (not Anisochoria minorella Mabille, 1898), Bralus alco (Evans, 1953) (not Bralus albida (Mabille, 1888)), Ephyriades jamaicensis (Möschler, 1879) (not Ephyriades brunnea (Herrich-Schäffer, 1865)), Koruthaialos (Stimula) frena Evans, 1949 (not Koruthaialos focula (Plötz, 1882)), Euphyes kiowah (Reakirt, 1866) (not Euphyes vestris (Boisduval, 1852)), Mnaseas inca Bell, 1930 (not Mnaseas bicolor (Mabille, 1889)), Metron hypochlora (Draudt, 1923) (not Metrocles schrottkyi (Giacomelli, 1911), previously in Metron Godman, 1900), Decinea huasteca (H. Freeman, 1969), Decinea denta Evans, 1955, and Decinea antus (Mabille, 1895) (not Decinea decinea (Hewitson, 1876)), Xeniades pteras Godman, 1900 (not Xeniades chalestra (Hewitson, 1866)), Xeniades difficilis Draudt, 1923 (not Xeniades orchamus (Cramer, 1777)), Xeniades hermoda (Hewitson, 1870) (not Tisias quadrata (HerrichSchäffer, 1869)), Hermio vina (Evans, 1955) (not Hermio hermione (Schaus, 1913), previously in Lento Evans, 1955), Cymaenes loxa Evans, 1955, (not Cymaenes laureolus (Schaus, 1913)), Niconiades peri (Evans, 1955) (not Rhinthon bajula (Schaus, 1902), previously in Neoxeniades Hayward, 1938), Gallio danius (Bell, 1941) (not Vehilius seriatus (Mabille, 1891)), Gallio massarus (E. Bell, 1940) (not Gallio garima (Schaus, 1902) previously in Tigasis Godman, 1900), Cymaenes edata (Plötz, 1882), Cymaenes miqua (Dyar, 1913) and Cymaenes aequatoria (Hayward, 1940) (not Cymaenes odilia (Burmeister, 1878)), Lychnuchus (Enosis) demon (Evans, 1955) (not Lychnuchus (Enosis) immaculata (Hewitson, 1868), previously in Enosis Mabille, 1889), Naevolus naevus Evans, 1955 (not Naevolus orius (Mabille, 1883)), Lucida scopas (Mabille, 1891), Lucida oebasus (Godman, 1900), and Lucida leopardus (Weeks, 1901) (not Lucida lucia (Capronnier, 1874)), Corticea schwarzi (E. Bell, 1941) and Corticea sylva (Hayward, 1942) (not Corticea mendica (Mabille, 1898)), and Choranthus orientis (Skinner, 1920) (not Choranthus antiqua (Herrich-Schäffer, 1863), previously in Pyrrhocalles Mabille, 1904). Borbo impar bipunctata (Elwes and J. Edwards, 1897) is a valid subspecies, not a synonym of Borbo impar tetragraphus (Mabille, 1891), here placed in synonymy with Lotongus calathus (Hewitson, 1876), new synonym. We confirm the species status of Telegonus cassius (Evans, 1952) and Lerema (Morys) valda Evans, 1955. Euphyes chamuli Freeman, 1969 is placed as a subspecies of Euphyes kiowah (Reakirt, 1866), new status. The following 41 taxa are junior subjective synonyms, either newly proposed or transferred from synonymy with other species or subspecies: Telegonus mutius Plötz, 1882 of Euriphellus phraxanor (Hewitson, 1876), Telegonus erythras Mabille, 1888 of Dyscophellus damias (Plötz, 1882), Aethilla jaira Butler, 1870 of Telegonus cretellus (Herrich-Schäffer, 1869), Paches era Evans, 1953 of Santa palica (Mabille, 1888), Antigonus alburnea Plötz, 1884 of Tolius tolimus robigus (Plötz, 1884) (not of Echelatus sempiternus simplicior (Möschler, 1877)), Echelatus depenicillus Strand, 1921 of E. sempiternus simplicior (not of T. tolimus robigus), Antigonus aura Plötz, 1884 of Theagenes dichrous (Mabille, 1878) (not of Helias phalaenoides palpalis (Latreille, [1824])), Achlyodes impressus Mabille, 1889 of Camptopleura orsus (Mabille, 1889), Augiades tania Schaus, 1902 of Metron voranus (Mabille, 1891), Pamphila verdanta Weeks, 1906 of Metron fasciata (Möschler, 1877), Niconiades viridis vista Evans, 1955 of Niconiades derisor (Mabille, 1891), Pamphila binaria Mabille, 1891 of Conga chydaea (A. Butler, 1877) (not of Cynea cynea (Hewitson, 1876)), Psoralis concolor Nicolay, 1980 of Ralis immaculatus (Hayward, 1940), Hesperia dido Plötz, 1882 of Cynea (Quinta) cannae (Herrich-Schäffer, 1869) (not of Lerema lochius (Plötz, 1882)), Proteides osembo Möschler, 1883 of Cynea (Cynea) diluta (Herrich-Schäffer, 1869) (not of Cynea (Quinta) cannae (Herrich-Schäffer, 1869)), Cobalopsis brema E. Bell, 1959 of Eutus rastaca (Schaus, 1902), Psoralis panamensis Anderson and Nakamura, 2019 of Rhomba gertschi (Bell, 1937), Cobalus asella Herrich-Schäffer, 1869 of Amblyscirtes alternata (Grote and Robinson, 1867) (not of Amblyscirtes vialis (W. H. Edwards, 1862)), Papias trimacula Nicolay, 1973 of Nastra subsordida (Mabille, 1891), Pamphila bipunctata Mabille, 1889 and Sarega staurus Mabille, 1904 of Lerema pattenii Scudder, 1872 (not of Cymaenes lumina (Herrich-Schäffer, 1869), previously in Lerema Scudder, 1872), Hesperia aethra Plötz, 1886 of Lerema lineosa (Herrich-Schäffer, 1865) (not of Lerema (Morys) compta Butler, 1877), Megistias miaba Schaus, 1902 of Cobalopsis valerius (Möschler, 1879), Phanis sylvia Kaye, 1914 of Lerema etelka (Schaus, 1902) (not of Lerema (Geia) geisa (Möschler, 1879), previously in Morys Godman, 1900), Carystus odilia Burmeister, 1878, Pamphila trebius Mabille, 1891 and Megistias corescene Schaus, 1902 of Cymaenes lumina (Herrich-Schäffer, 1869), Hesperia phocylides Plötz, 1882 of Cymaenes edata (Plötz, 1882) (not of Lerema accius (J. E. Smith, 1797)), Pamphila xenos Mabille, 1898 of Vehilius inca (Scudder, 1872), Mnasilus guianae Lindsey, 1925 of Papias amyrna (Mabille, 1891), Pamphila nubila Mabille, 1891 of Papias integra (Mabille, 1891) (not of Cynea corisana (Plötz, 1882)), Enosis matheri H. Freeman, 1969 of Metiscus atheas Godman, 1900 (previously in Enosis Mabille, 1889), Hesperia infuscata Plötz, 1882 of Mnaseas derasa derasa (Herrich-Schäffer, 1870) (previously Arotis Mabille, 1904), (not of Papias subcostulata (Herrich-Schäffer, 1870)), Pamphila astur Mabille, 1891 of Metiscus angularis (Möschler, 1877) (not of Cymaenes tripunctus theogenis (Capronnier, 1874)), Anthoptus macalpinei H. Freeman, 1969 of Anthoptus inculta (Dyar, 1918), Methionopsis typhon Godman, 1901 of Methionopsis ina (Plötz, 1882), Methionopsis dolor Evans, 1955 of Thargella volasus (Godman, 1901), Hesperia cinica Plötz, 1882 of Dubiella dubius (Stoll, 1781), Cobalus disjuncta Herrich-Schäffer, 1869 of Dubiella dubius (Stoll, 1781) (not of Vettius lafrenaye (Latreille, [1824])), and Saliana vixen Evans, 1955 of Neoxeniades parna (Evans, 1955). The following are new and revised genusspecies combinations: Euriphellus cebrenus (Cramer, 1777) (not Salatis Evans, 1952), Gorgopas extensa (Mabille, 1891) (not Polyctor Evans, 1953), Clytius shola (Evans, 1953) (not Staphylus Godman and Salvin, 1896), Perus narycus (Mabille, 1889) (not Ouleus Lindsey, 1925), Perus parvus (Steinhauser and Austin, 1993) (not Staphylus Godman and Salvin, 1896), Pholisora litus (Dyar, 1912) (not Bolla Mabille, 1903), Carrhenes decens (A. Butler, 1874) (not Antigonus Hübner, [1819]), Santa palica (Mabille, 1888) (not Chiothion Grishin, 2019), Bralus nadia (Nicolay, 1980) (not Anisochoria Mabille, 1876), Acerbas sarala (de Nicéville, 1889) (not Lotongus Distant, 1886), Caenides sophia (Evans, 1937) (not Hypoleucis Mabille, 1891), Hypoleucis dacena (Hewitson, 1876) (not Caenides Holland, 1896), Dotta tura (Evans, 1951) (not Astictopterus C. Felder and R. Felder, 1860), Nervia wallengrenii (Trimen, 1883) (not Kedestes Watson, 1893), Testia mammaea (Hewitson, 1876) (not Decinea Evans, 1955), Oxynthes trinka (Evans, 1955) (not Orthos Evans, 1955), Metrocles argentea (Weeks, 1901) (not Paratrytone Godman, 1900), Metrocles scitula (Hayward, 1951) (not Mucia Godman, 1900), Metrocles schrottkyi (Giacomelli, 1911) (not Metron Godman, 1900), Niconiades derisor (Mabille, 1891) (not Decinea Evans, 1955), Paratrytone samenta (Dyar, 1914) (not Ochlodes Scudder, 1872), Oligoria (Cobaloides) locutia (Hewitson, 1876) (not Quinta Evans, 1955), Psoralis (Saniba) laska (Evans, 1955) (not Vidius Evans, 1955), Psoralis (Saniba) arva (Evans, 1955) and Psoralis (Saniba) umbrata (Erschoff, 1876) (not Vettius Godman, 1901), Psoralis (Saniba) calcarea (Schaus, 1902) and Psoralis (Saniba) visendus (E. Bell, 1942) (not Molo Godman, 1900), Alychna gota (Evans, 1955) (not Psoralis Mabille, 1904), Adlerodea asema (Mabille, 1891) and Adlerodea subpunctata (Hayward, 1940) (not Eutychide Godman, 1900), Ralis immaculatus (Hayward, 1940) (not Mucia Godman, 1900), Rhinthon braesia (Hewitson, 1867) and Rhinthon bajula (Schaus, 1902) (not Neoxeniades Hayward, 1938), Cymaenes lochius Plötz, 1882 (not Lerema Scudder, 1872), Paracarystus ranka (Evans, 1955) (not Thoon Godman, 1900), Tricrista aethus (Hayward, 1951), Tricrista canta (Evans, 1955), Tricrista slopa (Evans, 1955), Tricrista circellata (Plötz, 1882), and Tricrista taxes (Godman, 1900) (not Thoon Godman, 1900), Gallio madius (E. Bell, 1941) and Gallio seriatus (Mabille, 1891) (not Vehilius Godman, 1900), Gallio garima (Schaus, 1902) (not Tigasis Godman, 1900), Tigasis corope (HerrichSchäffer, 1869) (not Cynea Evans, 1955), Tigasis perloides (Plötz, 1882) (not Cymaenes Scudder, 1872), Amblyscirtes (Flor) florus (Godman, 1900) (not Repens Evans, 1955), Vidius fraus (Godman, 1900) (not Cymaenes Scudder, 1872), Nastra celeus (Mabille, 1891) (not Vehilius Godman, 1900), Nastra nappa (Evans, 1955) (not Vidius Evans, 1955), Vehilius warreni (Weeks, 1901) and Vehilius limae (Lindsey, 1925) (not Cymaenes Scudder, 1872), Cymaenes lumina (Herrich-Schäffer, 1869) (not Lerema Scudder, 1872), Cobalopsis valerius (Möschler, 1879) (not Cobalopsis Godman, 1900), Cobalopsis dictys (Godman, 1900) (not Papias Godman, 1900), Lerema (Morys) venias (Bell, 1942) (not Cobalopsis Godman, 1900), Papias latonia (Schaus, 1913) (not Cobalopsis Godman, 1900), Dion iccius (Evans, 1955) and Dion uza (Hewitson, 1877) (not Enosis Mabille, 1889), Vistigma (Vistigma) opus (Steinhauser, 2008) (not Thoon Godman, 1900), Saturnus fartuga (Schaus, 1902) (not Parphorus Godman, 1900), Phlebodes fuldai (E. Bell, 1930) (not Vettius Godman, 1901), Mnasitheus padus (Evans, 1955) (not Moeris Godman, 1900), Naevolus brunnescens (Hayward, 1939) (not Psoralis Mabille, 1904), Lamponia ploetzii (Capronnier, 1874) (not Vettius Godman, 1901), Mnestheus silvaticus Hayward, 1940 (not Ludens Evans, 1955), Rigga spangla (Evans, 1955) (not Sodalia Evans, 1955), Corticea vicinus (Plötz, 1884) (not Lento Evans, 1955), Mnasalcas thymoetes (Hayward, 1942) (not Mnasicles Godman, 1901), Mnasalcas boyaca (Nicolay, 1973) (not Pamba Evans, 1955), Vertica brasta (Evans, 1955) (not Lychnuchus Hübner, [1831]), Carystina discors Plötz, 1882 (not Cobalus Hübner, [1819]), Zetka irena (Evans, 1955) (not Neoxeniades Hayward, 1938), and Neoxeniades parna (Evans, 1955) (not Niconiades Hübner, [1821]). The following are new or revised species-subspecies combinations: Tagiades neira moti Evans, 1934, Tagiades neira canonicus Fruhstorfer, 1910, Tagiades sheba vella Evans, 1934, Tagiades sheba lola Evans, 1945, Tagiades korela biakana Evans, 1934, Tagiades korela mefora Evans, 1934, Tagiades korela suffusus Rothschild, 1915, Tagiades korela brunta Evans, 1949, Tagiades ravi ravina Fruhstorfer, 1910, Tagiades atticus carnica Evans, 1934, Tagiades atticus nankowra Evans, 1934, Tagiades atticus helferi C. Felder, 1862, Tagiades atticus balana Fruhstorfer, 1910, Tagiades inconspicua mathias Evans, 1934, Tagiades hovia kazana Evans, 1934, Tagiades elegans fuscata de Jong and Treadaway, 2007, Tagiades elegans semperi Fruhstorfer, 1910, Metron hypochlora tomba Evans, 1955, Decinea denta pruda Evans, 1955, and Choranthus orientis eleutherae (Bates, 1934) (previously in Pyrrhocalles Mabille, 1904). In addition to the abovementioned changes, the following new combinations involve newly proposed genus group names: Fulvatis fulvius (Plötz, 1882) and Fulvatis scyrus (E. Bell, 1934) (not Salatis Evans, 1952); Adina adrastor (Mabille and Boullet, 1912) (not Bungalotis Watson, 1893); Nascus (Praxa) prax Evans, 1952, Nascus (Bron) broteas (Cramer, 1780), and Nascus (Bron) solon (Plötz, 1882) (not Pseudonascus Austin, 2008); Chirgus (Turis) veturius (Plötz, 1884); Paches (Tiges) liborius (Plötz, 1884), and Paches (Tiges) mutilatus (Hopffer, 1874) (not Antigonus Hübner, [1819]); Paches (Tiges) exosa (A. Butler, 1877); Tolius tolimus (Plötz, 1884) and Tolius luctuosus (Godman & Salvin, 1894) (not Echelatus Godman and Salvin, 1894); Ancistroides (Ocrypta) caerulea (Evans, 1928), Ancistroides (Ocrypta) renardi (Oberthür, 1878), Ancistroides (Ocrypta) waigensis (Plötz, 1882), Ancistroides (Ocrypta) aluensis (Swinhoe, 1907), Ancistroides (Ocrypta) flavipes (Janson, 1886), and Ancistroides (Ocrypta) maria (Evans, 1949) (not Notocrypta de Nicéville, 1889); Lennia lena (Evans, 1937), Lennia binoevatus (Mabille, 1891), Lennia maracanda (Hewitson, 1876), and Lennia lota (Evans, 1937) (not Leona Evans, 1937); Trida barberae (Trimen, 1873) and Trida sarahae (Henning and Henning, 1998) (not Kedestes Watson, 1893); Noxys viricuculla (Hayward, 1951) (not Oxynthes Godman, 1900); Xeniades (Tixe) quadrata (Herrich-Schäffer, 1869), Xeniades (Tixe) rinda (Evans, 1955), Xeniades (Tixe) putumayo (Constantino and Salazar, 2013) (not Tisias Godman, 1901); Gracilata quadrinotata (Mabille, 1889) (not Styriodes Schaus, 1913); Hermio hermione (Schaus, 1913) (not Lento Evans, 1955); Cynea (Nycea) hycsos (Mabille, 1891), Cynea (Nycea) corisana (Plötz, 1882), Cynea (Nycea) popla Evans, 1955, Cynea (Nycea) iquita (E. Bell, 1941), Cynea (Nycea) robba Evans, 1955, Cynea (Nycea) melius (Geyer, 1832), and Cynea (Nycea) irma (Möschler, 1879); Eutus rastaca (Schaus, 1902) (not Eutychide Godman, 1900); Eutus yesta (Evans, 1955) (not Thoon Godman, 1900); Eutus mubevensis (E. Bell, 1932) (not Tigasis Godman, 1900); Gufa gulala (Schaus, 1902) (not Mucia Godman, 1900); Gufa fusca (Hayward, 1940) (not Tigasis Godman, 1900); Godmia chlorocephala (Godman, 1900) (not Onophas Godman, 1900); Rhomba gertschi (E. Bell, 1937) (not Justinia Evans, 1955); Mnasicles (Nausia) nausiphanes (Schaus, 1913) (not Tigasis Godman, 1900); Amblyscirtes (Flor) florus (Godman, 1900) (not Repens Evans, 1955); Rectava ignarus (E. Bell, 1932) (not Papias Godman, 1900); Rectava vorgia (Schaus, 1902) (not Cobalopsis Godman, 1900); Rectava nostra (Evans, 1955) (not not Vidius Evans, 1955); Lerema (Geia) geisa (Möschler, 1879) and Lerema (Geia) lyde (Godman, 1900) (not Morys Godman, 1900); Contrastia distigma (Plötz, 1882) (not Cymaenes Scudder, 1872); Mit (Mit) badius (E. Bell, 1930) (not Styriodes Schaus, 1913); Mit (Mit) gemignanii (Hayward, 1940), (not Mnasitheus Godman, 1900); Mit (Rotundia) schausi (Mielke and Casagrande, 2002), (not Enosis Mabille, 1889); Picova steinbachi (E. Bell, 1930) (not Saturnus Evans, 1955); Lattus arabupuana (E. Bell, 1932) (not Eutocus Godman, 1901); Gubrus lugubris (Lindsey, 1925) (not Vehilius Godman, 1900); Thargella (Pseudopapias) tristissimus (Schaus, 1902) (not Papias Godman, 1900); Koria kora (Hewitson, 1877) (not Justinia Evans, 1955); Justinia (Septia) septa Evans, 1955; Corta lycortas (Godman, 1900) (not Orthos Evans, 1955); Vertica (Brasta) brasta (Evans, 1955) (not Lychnuchus Hübner, [1831]); Calvetta calvina (Hewitson, 1866) (not Cobalus Hübner, [1819]); Neoxeniades (Bina) gabina (Godman, 1900) (not Orthos Evans, 1955); Oz ozias (Hewitson, 1878) and Oz sebastiani Salazar and Constantino, 2013 (not Lychnuchoides Godman, 1901); and Carystoides (Balma) balza Evans, 1955 and Carystoides (Balma) maroma (Möschler, 1877). Finally, unless stated otherwise, all subgenera, species, subspecies and synonyms of mentioned genera and species are transferred together with their parent taxa, and taxa not mentioned in this work remain as previously classified.
769
Five new species and one new genus in Neotropical Lamiinae (Coleoptera: Cerambycidae) are de-scribed: Cotycicuiara lingafelteri Wappes and Santos-Silva, from Panama (Acanthoderini); Nesozineus morrisi Wappes and Santos-Silva, from Bolivia (Acanthoderini); Trichoanoreina panamensis Wappes and Santos-Silva, from Panama (Acanthoderini); Callisema skillmani Wappes and Santos-Silva, from Bolivia (Calliini); Rileyellus panamensis Wappes and Santos-Silva, new genus and species, from Panama (Desmiphorini). Callia pulchra Melzer, 1930 is placed in synonymy with Callia axillaris (Dalman, 1823).
0217
The taxonomic position of Onthophagus (Palaeonthophagus) lemuroides d’Orbigny, 1898 and Onthophagus
(Palaeonthophagus) fortigibber Reitter, 1909 is discussed (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini).
A key to the species is given. Photos of type specimens of the two taxa and significant chromatic varieties, and
drawings of aedeagi are presented.
1032
We propose several nomenclatural changes for taxa in the subfamily Cladodinae and the lampyrid tribes Cratomorphini, Lamprocerini, Lampyrini, Photinini, and Pleotomini in the subfamily Lampyrinae (Coleoptera: Lampyridae). We present evidence for the correct year of description of Calotrachelum flavolineatum Pic, 1941, Diaphanes lateapicalis breveapicalis Pic, 1954, Photinus conradti Pic, 1940, and all species described by Charles Émile Blanchard. Spellings are corrected for Diaphanes lateapicalis breveapicalis Pic, 1954, Diaphanes latipennis dartevillei Pic, 1952, Diaphanes longecarinatus Pic, 1955, Diaphanes moultoni latemarginatus Pic, 1938, Lampyris olivieriana von Heyden, 1890, and Photinus reductemarginalis Pic, 1941. We propose Lucidota puertoestrellaensis Keller and Martin nomen novum to replace Lucidota boliviana Pic, 1927, and Photinus laticollis brasiliensis Keller and Martin nomen novum for Photinus laticollis latior Pic, 1941. Seventy taxa described as variations or aberrations in the tribes Cratomorphini, Lamprocerini, Lampyrini, Photinini, and Pleotomini in the subfamily Lampyrinae by McDermott (1966), are evaluated to subspecies with their availability determined based on ICZN (1999: Article 45.6). Photinus motschulskyi Zaragoza-Caballero, Zurita-Garcia, and Ramírez-Ponce, 2023 is synonymized under Photinus intercalatus Gemminger, 1870. Lastly, we correct the type species of the genus Lucidina Gorham, 1833, and address the dates of publication for fireflies described by Blanchard.
ZooBank registration. urn:lsid:zoobank.org:pub:DCD31B02-DF17-4115-A9A8-A6CBB02D8684
965
Several nomenclatural changes for taxa in the firefly subfamily Luciolinae (Coleoptera: Lampyridae) are proposed. Evidence is presented to correct the year of description for Luciola dregei Motschulsky 1853 to Motschulsky 1854 when the name was ratified. We correct the authorities and years of descriptions for type species designations for the genera Delopleurus Motschulsky, 1853, and Delopyrus Motschulsky, 1853 to Motschulsky 1854 for both. All remaining taxa described as variations in the subfamily Luciolinae by McDermott (1966), are evaluated to subspecies with their availability determined based on ICZN (1999) Article 45.6.
ZooBank registration. urn:lsid:zoobank.org:pub:618A5442-2644-4E9A-BE70-07C413810DB9
987
We propose several nomenclatural changes for taxa in the lampyrid subfamilies Ototretinae, Photurinae, and Psilocladinae (Coleoptera: Lampyridae). Evidence for the correct year of description of Drilaster albicornis lateobscura (Pic, 1921) is presented. Spellings are corrected for Drilaster debilis holzi (Pic, 1914), Drilaster impustulata fukienensis (Pic, 1955), Drilaster moutoni (Pic, 1911), Drilaster pendleburyi (Pic, 1943) and Pyrogaster lunifera (Eschscholtz, 1822). We also explain the validity of the name Photuris flavicollis Fall, 1927. Fifty-eight taxa described as variations or aberrations in the subfamilies Ototretinae, Photurinae, and Psilocladinae by Delkeskamp (1977), McDermott (1966), Wittmer (1944), and Pic (1924c), are evaluated, with their availability determined based on ICZN (1999: Article 45.6).
ZooBank registration. urn:lsid:zoobank.org:pub:65697128-997F-4A52-AC9E-6860B0BF997D
1055
The exclusively New World velvet ant genus Timulla Ashmead, 1899 (Hymenoptera: Mutillidae: Mutillinae: Trogaspidiini) contains 180 species and ten subspecies. Most of these species are known from a single sex, and the validity of the subspecies has not been evaluated since their original description in 1937 and 1938. The Timulla fauna of the United States of America includes thirty species and eight subspecies, and the fauna of Canada includes three species. The faunas of these two countries were critically studied with the following results. Out of the eight total subspecies, seven were found to be structurally identical to and sympatric with the nominate subspecies. The subspecific differences were limited to cuticle and/or setal coloration in males and intergrades between them were found in several cases. With the senior synonym listed first, seven subspecies-level synonymies are proposed, which include: Timulla barbigera (Bradley, 1916) = T. barbigera rohweri Mickel, 1937, new synonym; T. dubitata (Smith, 1855) = T. dubitata fugitiva Mickel, 1937, new synonym; T. hollensis (Melander, 1903) = T. hollensis melanderi Mickel, 1937, new synonym; T. ocellaria Mickel, 1937 = T. ocellaria rufidorsa Mickel, 1937, new synonym; T. suspensa (Gerstaecker, 1874) = T. suspensa jonesi Mickel, 1937, new synonym, = T. suspensa sonora Mickel, 1937, new synonym; T. vagans (Fabricius, 1798) = T. vagans rufinota Mickel, 1937, new synonym. The final remaining subspecies, Timulla navasota coahuila Krombein, 1951, is raised to a full species, Timulla coahuila Krombein, 1951, new status, based on its unique female morphology. Also, four new sex associations are proposed, which include: Timulla barbata (Fox, 1899) = T. wileyae Mickel, 1937, new synonym; T. euterpe (Blake, 1879) = T. compressicornis Mickel, 1937, new synonym; T. neobule Mickel, 1937 = T. nicholi Mickel, 1937, new synonym; T. subhyalina Mickel, 1937 = T. dubitatiformis Mickel, 1937, new synonym. The former species, T. dubitatiformis, which was previously known only from females, is here recognized as being a morphologically-conservative complex of species; its synonymy with T. subhyalina effectively associates the remaining male-based members of the Timulla ocellaria species-group with it as well, which includes Timulla hollensis (Melander, 1903), T. kansana Mickel, 1937, T. ocellaria Mickel, 1937, T. rufosignata (Bradley, 1916), T. sayi (Blake, 1871), T. subhyalina Mickel, 1937, and T. tolerata Mickel, 1937. Further, two species-level synonymies are proposed, which include: Timulla dubitata (Smith, 1855) = T. murcia Mickel, 1938, new synonym; T. vagans (Fabricius, 1798) = T. huntleyensis Mickel, 1937, new synonym. Finally, Timulla cyllene (Cameron, 1894) is newly recorded in the United States of America from the state of Arizona.
ZooBank registration. urn:lsid:zoobank.org:pub:891E0C92-B8BF-4487-84D4-42EB2254AF4A
751
Cyrtinus pygmaeus (Haldeman, 1847) (Coleoptera: Cerambycidae: Lamiinae) is redescribed and newly recorded from Mexico. The female of Decarthria stephensii Hope, 1834 is also redescribed, the number of specimens in the type series is corrected, as is the depository of the types, and the species is newly recorded from Dominica. Two new species of Cyrtinus LeConte, 1852 are described from Mexico: C. fisheri Wappes, Santos-Silva and Nascimento; and C. howdeni Wappes, Santos-Silva and Nascimento. A key to species of Decarthria Hope, 1834 (adapted from an earlier key to Cyrtinini) is provided.
0090
A taxonomic overview for the fritillary genus Speyeria Scudder (= Argynnis Fabricius) and its placement within the Heliconiinae (Nymphalidae) is presented. Taxonomic accounts, type images, and relevant literature for the 25 subspecies within the Speyeria atlantis (Edwards) and Speyeria hesperis (Edwards) complexes and the 16 nominate Speyeria species are included. Errors in nomenclature are identified, taxonomic, life history, and distributional information are updated, and type locality information is discussed. Images of primary type specimens for all 16 Speyeria species and the 25 subspecies in the atlantis - hesperis complexes appear together in color here for the first time. One new combination of a species-subspecies is created: Speyeria hesperis hanseni Emmel, Emmel, and Matoon, 1998, new combination, which was previously Speyeria atlantis hanseni Emmel, Emmel, and Matoon.
983
Autarcontes lopezi Fisher, 1925 (Coleoptera: Buprestidae), is transferred to the genus Agrilus Curtis, 1825 (new combination). Colobogaster bella Kirsch, 1873, is transferred to the genus Chrysobothris Eschscholtz, 1829 (new combination). Ectinogonia isamarae Moore, 1994, is resurrected as the valid name for the species previously called E. obscuripennis Cobos, 1954, as the latter is unavailable as infrasubspecific. Conognatha jakobsoni Obenberger, 1928, is resurrected over C. germaini Théry in Hoscheck, 1934, as the former name has priority. Callimicra lucida Waterhouse, 1889, is resurrected as the valid name over C. hoscheki Obenberger, 1922, which has been used due to several historical errors and misinterpretations.
ZooBank registration. urn:lsid:zoobank.org:pub:651D001D-1C91-4A1A-B8BE-335BC7E7DD0F
0066
The New World euparine scarab genera Parataenius Balthasar, 1961 and Pseudataenius Brown, 1927 are revised. Ataenius brunneus Schmidt is transferred to the genus Parataenius becoming Parataenius brunneus (Schmidt), new combination. The monospecific genus Ataenioides Petrovitz, 1973, is synonymized with Pseudataenius Brown, 1927, (new synonymy) and the type species, Ataenioides gracilitarsis Petrovitz, is given the new combination Pseudataenius gracilitarsis (Petrovitz). New species of Parataenius are described from southern South America: Parataenius selvae, P. estero, and P. martinezi. Keys for species of both genera are presented and pertinent morphological details are illustrated.
0050
The Tetragonoderus (Peronoscelis) quadriguttatus assemblage is a postulated monophyletic part of the intersectus complex. This assemblage is characterized against a background that includes review of a classification of the genus and comparison, in the form of a key, to other unrelated Western Hemisphere genera that share elongate tibial spurs with Tetragonoderus Dejean. The principal, easily observed feature that unites the three members (T. laevigatus Chaudoir, 1876; T. deuvei, new species (type locality: Cuijaba, Mato Grosso, Brazil), and T. quadriguttatus Dejean) of the quadriguttatus assemblage is the four-spotted elytra (two spots per elytron). These species are treated in detail, including key, synonymy (as required), comparisons, description (external and male genitalic features), habitat (if known), locality data, and geographical range map. Also, T. subfasciatus Putzeys, 1846, the elytral color pattern of whose members may be confused with the spotted pattern of the quadriguttatus assemblage, is treated similarly. The following new synonymies were established: T. quadriguttatus Dejean 1829 = T. columbicus Steinheil 1875 = T. lacordairei Chaudoir 1876 = T. tetragrammus Chaudoir 1876; and T. laevigatus Chaudoir 1876 = T. chaudoiri Liebke 1928 (replacement name for the junior homonym, T. unicolor Chaudoir, 1876). Lectotypes are designated for T. unicolor Chaudoir, T. lacordairei Chaudoir, and T. subfasciatus Putzeys. A neotype is designated for T. quadriguttatus Dejean. New distribution records in the West Indies for T. quadriguttatus are recorded for the islands of Grand Cayman, Jamaica and Hispaniola. These records may be the result of recent natural overseas dispersal from northern South America, or they may be the result of human-mediated accidental introduction, or a combination of both. The southern Floridian (Nearctic) records for T. laevigatus probably represent a recent accidental introduction through commerce, followed by dispersal through flights of adults.
952
This paper presents a taxonomic review of the genus Boreocanthon Halffter, restored generic status, a group of ball-rolling (telocoprid) dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) endemic to North America. The genus, heretofore treated as a subgenus of Canthon, comprises 13 species, each keyed, diagnosed, illustrated and presented with information on distribution, relationships, and biology (*signifies restored generic combination): *Boreocanthon ateuchiceps (Bates, 1887); B. coahuilensis (Howden, 1966); *B. depressipennis (LeConte, 1859c); *B. ebenus (Say, 1823); *B. forreri (Bates, 1887); *B. integricollis (Schaeffer, 1915); *B. lecontei (Harold, 1868); *B. melanus (Robinson, 1948); *B. praticola (LeConte, 1868); *B. probus (Germar, 1823); *B. puncticollis (LeConte, 1866); *B. simplex (LeConte, 1857); as well as Boreocanthon halffteri Edmonds here described as a new species. Other actions taken are a) Canthon mixtus Robinson declared junior subjective new synonym of Boreocanthon puncticollis; b) Canthon bisignatus Balthasar, 1939, declared junior subjective new synonym of Boreocanthon simplex; c) Boreocanthon coahuilensis (Howden) new generic combination; and d) neotype designated for the type species of the genus Boreocanthon, Ateuchus ebenus Say, 1823, here Boreocanthon ebenus (Say).
ZooBank registration. urn:lsid:zoobank.org:pub:74CC7BA2-4E8D-4780-BB1B-E47370CBB19D
1014
t. This paper presents a taxonomic review of the genus Melanocanthon Halffter, a group of ball-rolling (telocoprid) dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) endemic to eastern North America. The genus comprises five species, each keyed, diagnosed, illustrated and presented with information on distribution, relationships, and biology: Melanocanthon punctaticollis (Schaeffer), M. granulifer (Schmidt), M. nigricornis (Say), M. bispinatus (Robinson) and Melanocanthon vulturnus Edmonds, new species.
ZooBank registration. urn:lsid:zoobank.org:pub:FA7D5D5E-CEB8-48ED-A442-74C315FCF5E4
879
Two subspecies of Cosmodela duponti (Dejean) (Coleoptera: Cicindelidae) are elevated to species rank: Cosmodela barmanica (Gestro) and C. indica (Fleutiaux). The lectotypes of all the above-mentioned species are designated as well. Short redescriptions of the three species are provided together with a key, and illustrations of their habitus.
848
Within Nearctic Cicindelidae, Cicindelini, Eunota togata (LaFerté-Sénectère, 1841) (Coleoptera: Cicindelidae) exists as several distinct phenotypes, some described as subspecies whose taxonomic status has been misinterpreted. Eunota togata togata applies to adults marked with broadly sinuate inner margins on shiny brown elytra common across northern coastal Gulf of Mexico. Eunota togata fascinans (Casey,1914), new synonymy, incorrectly applied to populationsin western Texas and New Mexico, United States, is considered a junior synonym of E. togata globicollis (Casey,1913), the predominant subspecies with broadly sinuate inner margins on coppery elytra found from eastern New Mexico and western Texas into the Southern Great Plains of North America. Eunota togata leucophasma, new subspecies, is comprised of adults with mostly white elytra that are isolated in Hudspeth County, Texas, and Doña Ana, Otero, and Torrance Counties, New Mexico. Intergrades between E. togata globicollis and E. togata leucophasma, new subspecies, are found sporadically in eastern New Mexico and western Texas; however, suggested intergrades between E. togata globicollis and E. togata togata in western Texas are reinterpreted as more typical of intergrades farther north in New Mexico. Adults marked with acutely sinuate inner margins on darker elytra found only in northcentral Kansas to southeast Nebraska should be referred to as E. togata latilabris (Willis,1967), new status. All subspecies are collectively known as the white-cloaked tiger beetle. Geological conditions influencing development of saline habitats and subsequent evolution of divergent white E. togata subspecies in Texas and New Mexico, and darker E. togata subspecies in Kansas and Nebraska, are discussed and compared to more uniformly marked E. togata subspecies found elsewhere in North America.
0379
Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae)
(2014)
The North American species of the genus Eusphalerum Kraatz (Coleoptera, Staphylinidae, Omaliinae) are revised. The taxonomic history, natural history, geographical distribution of the genus, characters, species groups, diversity, and biogeography of North American species are presented. Two main phylogenetic lineages and 13 species groups are provisionally recognized. The following new synonymies are proposed: Eusphalerum farrarae (Hatch, 1944) = E. lawrencei Hatch, 1957; Eusphalerum californicum (Fauvel, 1878) = E. atriventre (Casey, 1894), = E. nigerrimum (Casey, 1894), = E. gilvipenne (Casey, 1894), = E. dichroum (Fall, 1922), = E. bonnelli (Hatch, 1944), = E. lunae Hatch, 1957; Eusphalerum fraternum (Casey, 1894) = E. minskae (Hatch, 1944); Eusphalerum rugulosum (Mäklin, 1853) = E. grayae (Hatch, 1944); Eusphalerum orientale (Bernhauer, 1912) = E. frosti (Bernhauer, 1928). The following lectotypes are designated: E. subangulatum (Casey), E. californicum (Fauvel), E. gilvipenne (Casey), E. diversicolle (Casey), E. convexum (Fauvel), E. fraternum (Casey), E. horni (Fauvel), E. orientale (Bernhauer), E. pothos (Mannerheim), and E. punctatum (Casey). The following new species are described: Eusphalerum pilosum (California); E. klimaszewskii (British Columbia); E. chatzimanolisi (California); E. carolinensis (Kentucky, Missouri, North Carolina, Ohio, Pennsylvania, Tennessee, Virginia); E. caterinoi (California); E. luteipes (California); E. thayeranum (Alberta, British Columbia, Idaho, Indiana (doubtful record), Oregon, Washington); E. margaretae (Tennessee); E. newtoni (British Columbia, Oregon, California); E. parvispiculum (California, Oregon); E. uncinatum (British Columbia, California, Oregon, Washington). Eusphalerum lapponicum (Mannerheim, 1830) is excluded from the North American fauna. The following new combination is proposed: Xylodromus segmentarius (Mäklin, 1852: 322) (ex Omalium), wrongly attributed to Eusphalerum in the literature. Omalium marginatum Say, 1832 is considered a doubtful species, probably not Eusphalerum. A key to the 27 recognized North American Eusphalerum species and a catalog of the species are provided.
793
The New World genus Axina Kirby (Coleoptera: Cleridae) is revised for the first time. Thirty-two new species are described: Axina acutipennis, A. adelosa, A. atmis, A. bahia, A. bella, A. brunnea, A. chiasta, A. furcula, A. heveli, A. ignota, A. klisis, A. latilinea, A. lobispinula, A. luzia, A. macilenta, A. megaspina, A.minas, A. ochra, A. oligocheia, A. ordinis, A. orcastomata, A. pallidioccabus, A. phallospina, A. piperata, A. pollex, A. polycaula, A. rio, A. schenklingi, A. spina, A. trinalis, A. villa, and A. vista. The 19 previously described species are: Axina analis Kirby, A. apicalis Pic, A. basalis Schenkling, A. bifasciata (Chevrolat), A. centrimaculata Schenkling, A. conspicua Schenkling, A. diversesignata Pic, A. equestris (Schenkling), A. fasciata Kirsch, A. fortipes Pic, A. lateralis Pic, A. longevittata Pic, A. munda Schenkling, A. nigrifrons Schenkling, A.parcepunctata Schenkling, A. picta Schenkling, A. plagiata Schenkling, A. proxima (Chevrolat), and A. sexmaculata Spinola. Two species Priocera equestris Schenkling and Priocera proxima Chevrolat, are transferred into the genus Axina becoming new combinations: Axina equestris (Schenkling) and Axina proxima (Chevrolat). Two names, Priocera podagrica Schenkling, 1900, and P. podagrica variety pygmaea Schenkling, 1902, are new synonymies of Priocera proxima Chevrolat, 1876. Lectotypes are here designated for nine species: Axina basalis Schenkling, 1900; Axina centrimaculata Schenkling, 1900; Axina conspicua Schenkling, 1900; Axina diversesignata Pic, 1946; Axina munda Schenkling, 1900; Axina nigrifrons Schenkling, 1906; Axina parcepunctata Schenkling, 1900; Axina picta Schenkling, 1907; and Axina plagiata Schenkling, 1900. It is proposed that Axina species are predators of lignicolous insects, particularly bark beetles. The species of Axina can be classified into eight species groups and a theory of their phylogenetic relationships is proposed via WINCLADA in conjunction with NONA. Of the 51 species that now comprise Axina, only one traversed the Panamanian portal before the Colombian Andes reached their modern altitudes. This work includes a generic-level morphological analysis, brief treatise of natural history, key to species, comments about Axinazoogeography, and hypotheses of species-group phylogeny.
669
This study evaluated the population variation of Hemiberlesia lataniae (Signoret) (Hemiptera: Diaspididae) and associated parasitoids in Olea europaea L. (Oleaceae) groves located in the municipality of Barra do Ribeiro (30°30'54.95?S; 51°30'20.84?W), Rio Grande do Sul, Brazil. Eight samples were collected during the four seasons from November 2015 until October 2017; four branches with approximately 20 leaves were taken from 12 olive trees (four each from Arbequina, Arbosana and Koroneiki varieties). The collected material was placed in plastic bags and taken to the laboratory for further study. Scale insects were slide-mounted and were identified according to morphological characteristics of the adult female. Parts of two branches and 40 leaves of each variety infested with scale insects were sectioned and placed inside glass tubes, labeled, capped with cotton, kept in the laboratory at room temperature and examined daily to verify the emergence of parasitoids. Parasitoid specimens studied were critical point dried from ethanol and point-mounted. Some specimens were slide mounted in Canada balsam for more detailed study. In all samples, the presence of H. lataniae, as well as six associated parasitoid species, were identified. Metaphycus flavus (Howard) is recorded for the first time as a parasitoid of H. lataniae. Two other scale insects were also identified: Pseudaulacaspis pentagona (Targioni Tozzetti) (Diaspididae) and Saissetia coffeae (Walker) (Coccidae), but the population data of these did not permit statistical analysis. A key to identify the scale insect species recorded on olive trees in Brazil, based on adult females, is provided.
502
Taxonomic supplement (2001 to 2015) to the catalogue of New Zealand Carabidae (Insecta: Coleoptera)
(2016)
A supplement to the “Carabidae (Insecta: Coleoptera): catalogue” published by Larochelle and Larivière (2001; Fauna of New Zealand 43), is provided. A total of 99 genera and 547 species and subspecies belonging to 21 tribes and 8 subfamilies of Carabidae (including Cicindelini and Rhysodini), is recorded for New Zealand. Endemism is high, with 93% of the species and subspecies as well as 62% of the genera recorded from this country only. Changes to the 2001 catalogue are documented. The synonymy and type locality of taxa described between 2001 and 2015 are also provided.
0274
The purpose of this paper is to reassess the taxonomy of Phanaeus MacLeay (Coleoptera: Scarabaeidae) to accommodate new taxa and changes in taxonomic opinion since the publication of Edmonds’ 1994 revision of the genus. The two subgenera and 13 species groups established by Edmonds (1994) remain unchanged. A revised set of keys with accompanying comments and illustrations separates 54 recognized valid species. Seven recently described valid species are incorporated into the revised classification: Phanaeus blackalleri Delgado-Castillo, 1991; P. bordoni Arnaud, 1996; P. changdiazi Kohlmann and Solís, 2001; P. lecourti Arnaud, 2000; P. martinezorum Arnaud, 2000; P. yecoraensis Edmonds, 2004; and P. zapotecus Edmonds, 2006. The new name Phanaeus sororibispinus Edmonds and Zidek replaces Phanaeus alvarengai Arnaud, 1984, a primary junior homonym of P. alvarengai Pereira and d’Andretta, 1955. Three subspecies recognized in 1994 are elevated to species rank, new status: Phanaeus texensis Edmonds, 1994; P. pilatei Harold, 1863; and P. guatemalensis Harold, 1871. Phanaeus obliquans Bates, 1887 is removed from synonymy and given new status as a valid species. Twelve new junior subjective synonyms (bold) are recognized: P. tridens balthasari Arnaud, 2002 (of P. tridens Castelnau, 1840); P. dzidoi Arnaud, 2000 (of P. palaeno Blanchard, 1843); P. genieri Arnaud, 2002 (of P. amethystinus Harold, 1863); P. prasinus jolyi Arnaud, 2001 (of P. prasinus Harold, 1868); P. kirbyi ledezmai Arnaud, 2002 (of P. kirbyi Vigors, 1825); P. achilles lydiae Arnaud, 2000 (of P. achilles Boheman, 1858); P. chalcomelas grossii Arnaud, 2001 (of P. chalcomelas [Perty, 1830]); P pyrois malyi Arnaud, 2002 (of P. pyrois Bates, 1887); P. tridens moroni Arnaud, 2001 (of P. tridens Castenau, 1840); P. lecourti peruanus Arnaud, 2000 (of P. lecourti Arnaud, 2000); P. endymion porioni Arnaud, 2001 (of P. endymion Harold, 1863); P. pseudofurcosus Balthasar, 1939 (of P. tridens Castelnau, 1840); and P. prasinus trinidadensis Arnaud, 2001 (of P. prasinus Harold, 1868). “Phanaeus viridicollis” Olsoufieff, 1924 (sensu Arnaud 2002) is an unavailable name here considered a color variant of P. pyrois Bates, 1887.
0191
The taxonomy of the African and Madagascan species of the tiger beetle genus Chaetodera Jeannel 1946 (Coleoptera: Cicindelidae) is reviewed based on studies of primary types and additional museum specimens. Six species are recognized: C. andriana (Alluaud 1900), C. antatsima (Alluaud 1902), C. blanchardi (Fairmaire 1882), C. maheva (Künckel d’Herculais 1887), C. perrieri (Fairmaire 1897), and C. regalis (Dejean 1831). All species are illustrated, including color variants, and a key to species and maps of species distributions are provided. A hypothesis of phylogenetic relationships is proposed for the nine worldwide species of the genus Chaetodera based on computerized parsimony analysis of a matrix containing data on 16 adult morphological characteristics.
649
The linyphiid spiders Tenuiphantes zelatus (Zorsch, 1937), T. zibus (Zorsch, 1937), and Centromerus mariannae sp. nov. are diagnosed, described or redescribed and illustrated, including the previously undescribed female of T. zibus. All are common species of the Pacific Northwest of North America but were previously inadequately described (T. zelatus and T. zibus) or illustrated only but not described or named (C. mariannae).
762
A new species of the genus Termitodiellus Nakane, 1961—T. mindanaoensis Minkina and Kakizoe, new species from Mindanao Island in the Philippines is described and illustrated. Photographs of the epipharynx of Termitodiellus species are provided for the first time. A short discussion of the phylogeny of Rhyparini Schmidt, 1910 is presented.
0119
The Anthonomus juniperinus group, with descriptions of two new species (Coleoptera: Curculionidae)
(2010)
The Anthonomus juniperinus (Sanborn) species group is defined and two new species, Anthonomus sanborni, new species, and A. rileyi new species, from the United States are described, keyed and illustrated. The three species of the group are associated with the plant genus Juniperus and the larvae of A. juniperinus are known to develop in fungal galls of Gymnosporangium spp. as well as fruits of the Eastern redcedar, Juniperus virginiana L. The biology of the group and its taxonomic relationships to other species of Anthonomus Germar are also discussed.
0206
Among the four oriental genera of the tribe Helluonini, Omphra Dejean (Coleoptera: Carabidae), is unique for its endemism to the Indian subcontinent and aptery. High intraspecies variability in morphological characters and limited diagnostic information makes species differentiation of the genus Omphra a complicated task. The present study provides a description of a new species, Omphra drumonti n. sp. from the Western Ghats, redescriptions and a key to the species of Omphra, details of intraspecies variation, discussion of relationships between taxa and distributional patterns of the genus. Based on the distributional patterns in the Indian subcontinent and flightlessness of the genus, inability to cross the physical barrier of the Ganges–Brahmaputra delta between north and peninsular India is indicated as the reason for its absence in the northeastern Indian subcontinent and endemism to the lower Indian subcontinent.
0040
The bee fauna of the Greater Puerto Rico area was studied. A review of the previous relevant studies is presented. An annotated catalog and information about the origin and distributional patterns are also provided. Thirty-nine species of bees occur in Puerto Rico and adjacent islands. This fauna is composed of four elements: exclusive Puerto Rican endemics (26.5%); Antillean endemics occurring on multiple islands (76.5%); continental species that have also colonized the Antilles (23.5%); and species introduced through human activity (12.8%). The bee fauna was both low in its diversity and showed the highest level of disharmony in relation to other faunas of the Greater Antilles. A lectotype is here designated for Agapostemon krugii Wolcott, 1936.
0073
The beetle fauna of the island of Barbados is summarized. It is now known to contain 202 genera, and 254 species (in 40 families), of which 232 are named at the species level. Undoubtedly, the actual numbers of species on Barbados are much higher than now known. Nine species are possibly endemic to the island, 15 have been intentionally introduced, and 51 have probably been accidentally introduced through human activity. The remaining 157 named species may occur naturally as a result of natural over-water dispersal processes. These species mostly have a wide distribution in the Antilles and Latin America. The total named fauna is thus about 72% naturally occurring, and 28% the result of human-aided dispersal.
0178
The beetles of Martinique, Lesser Antilles (Insecta: Coleoptera); diversity and distributions
(2011)
This paper summarizes the published information on the beetle fauna of the island of Martinique, in the Lesser Antilles. The fauna is known to contain 42 families, with 201 genera, and 270 species. The families with the largest numbers of species are Cerambycidae (57), Curculionidae (43), Scarabaeidae (42), Tenebrionidae (18), and Staphylinidae (17). At least 15species (5.5%) were probably accidentally introduced to the island by human activities. Forty six species (17.0%) are endemic (restricted) to the island and likely speciated on the island. Sixty seven species (24.8%) are shared only with other islands of the Lesser Antilles (Lesser Antillean endemics), and 26 species (9.6%) are more widespread Antilles endemics. The remaining 116 species (42.8%) in the fauna are otherwise mostly widely distributed in the Antilles and the Neotropical Region. The Martinique beetle fauna has mostly originated elsewhere than on Martinique and is largely an immigrant fauna from other islands of the West Indies or the continental Neotropics. The numbers of Martinique species shared with other larger islands of the Lesser Antilles are (north to south) Montserrat (73), Guadeloupe (175), Dominica (98), St. Lucia (68), St. Vincent (93), and Grenada (88). Undoubtedly, the real number of species on Martinique is much higher than now reported and may actually be around 1600 or more species.
0106
The published beetle fauna of the island of St. Lucia is summarized. It contains 135 genera, and 175 species in 25 families. Four species are accidentally introduced by human activities. Twenty three species are endemic (restricted) to the island. Twenty seven species on St. Lucia are shared only with other islands of the Lesser Antilles, and 22 species are widespread Antilles endemics. The remaining 56.6% of the fauna is otherwise mostly one which is widely distributed in the Antilles and the Neotropics. This suggests that it is mostly an immigrant fauna originating in the continental Neotropics. Undoubtedly, the actual numbers of species on St. Lucia are many times higher than now reported and may originally have been as high as around 1400 species. Of the St. Lucia species known to occur on other islands, the largest numbers are shared with Guadeloupe (102), St. Vincent (79), and Martinique (55).
0144
This paper summarizes the information published on the beetle fauna of the island of St. Vincent (excluding the Grenadine islands). The fauna contains 62 families, with 371 genera, and 536 species. The families with the largest number of species are Staphylinidae (128), Curculionidae (54), Chrysomelidae (47), Scarabaeidae (31), Tenebrionidae (30), and Cerambycidae (29). At least 17 species (3.17%) were probably accidentally introduced to the island by human activities. One hundred four species (19.40%) are endemic (restricted) to the island and likely speciated on the island. One hundred twenty species (22.39%) are shared only with other islands of the Lesser Antilles (Lesser Antillean endemics), and 41 species (7.65%) are more widespread Antilles endemics. The remaining 254 species (47.38%) in the fauna are otherwise mostly widely distributed in the Antilles and the Neotropical Region. The St. Vincent beetle fauna has thus mostly originated elsewhere than on St. Vincent and is largely an immigrant fauna from other islands of the West Indies or the continental Neotropics. Of the St. Vincent species known to occur on other islands, the largest numbers are shared with (north to south) Guadeloupe (206), Dominica (115), Martinique (76), St. Lucia (87) and Grenada (298). Undoubtedly, the real number of species on St. Vincent is higher than now reported and may actually be around 1200 or more species.
0460
The island arc of the Lesser Antilles lies at the eastern margin of the Caribbean Sea in the Western Hemisphere, and stretches from the eastern end of the islands of the Greater Antilles (at the Virgin Islands), south to a position near the continental islands of Trinidad and Tobago at the north eastern corner of South America. The islands are a part of the West Indian Islands biodiversity “hotspot” and have been available for terrestrial colonization for about the past 15 million years. This is a status report on present knowledge of the beetle faunas of these islands, which is composed of 90 families, 1210 genera, and 2612 recognized species. Many additional species are not yet identified, or are unnamed, or remain to be discovered. Reported for the first time from the Lesser Antilles are four families, 49 genera, 105 species, and 1253 new island records. The largest families are Curculionidae (588 species), Staphylinidae (389 species), Chrysomelidae (181 species), Tenebrionidae (142 species), Cerambycidae (138 species), Scarabaeidae (127 species), and Carabidae (126 species). There are differing patterns of species distributions: 154 species are probably introduced by human activities; 985 are endemic species (limited to a single island); 465 are species endemic to more than one island of the Lesser Antilles; 212 are species limited to just islands of the West Indies; and 800 are native (naturally occurring) species which also have part of their distributional range in North, Central, or South America. Most of the widely distributed beetle fauna has probably come from South America by over-water dispersal. There is no compelling evidence for a vicariance origin of any part of the beetle fauna. Earlier colonists have had more time to form endemic genera (18) and endemic species. The more widely distributed species probably represent distributions achieved in and since the Pleistocene.
0324
We provide new records of biting and predaceous midges (Diptera: Ceratopogonidae) from Guadeloupe in the subfamily Ceratopogoninae, including descriptions and illustrations of three new predaceous species in the genera, Parabezzia Malloch, Stilobezzia Kieffer and Palpomyia Meigen, respectively, and the first records of the New World predaceous genus, Amerohelea Grogan and Wirth, from the Caribbean region. We also provide the first Guadeloupe records of the biting midges, Culicoides (Anilomyia) decor (Williston), C. (Avaritia) pusillus Lutz, C. (Drymodesmyia) bredini Wirth and Blanton, C. (D.) poikilonotus Macfie, C. (Haematomyidium) hoffmani Fox, C. (Hoffmania) insignis Lutz, C. rangeli Ortiz and Mirsa and C. trilineatus Fox, and the predaceous midges, Brachypogon (Brachypogon) bifidus Spinelli and Grogan, B. (B.) telesfordi Spinelli and Grogan, B. (B.) woodruffi Spinelli and Grogan, Monohelea maya Felippe-Bauer, Huerta and Ibáñez-Bernal, Stilobezzia (Stilobezzia) diminuta Lane and Forattini, S. (S.) thomsenae Wirth, Amerohelea galindoi Grogan and Wirth, Bezzia (Bezzia) flinti Spinelli and Wirth, B. (Homobezzia) venustula (Williston) and Palpomyia insularis Spinelli and Grogan.
0166
From 1995 to 2004 collections for Bruchidae (Coleoptera) were made in La Reserva de la Biósfera Sierra de Huautla, Morelos, Mexico. Specimens were reared from mature seedpods, but also collected by net, malaise trap, and light trap. In total 72 species in 13 genera of Bruchidae were recovered. Of those two new species are here described: Amblycerus montalvoi Romero and Acanthoscelides camerinoi Romero. We record 27 host plants for the bruchids found in the study area.
539
Six annotated lists are presented: A, a checklist of the butterflies (Lepidoptera, Papilionoidea) of Tobago (150 species); B, species for which there are no records in the last 80 years (49 species); C, species needing confi rmation from Tobago (5 species); D, species not accepted from Tobago (12 species); E, species which are likely to occur in Tobago, but have not been recorded (6 species); and F, species and subspecies recorded from Tobago, but not from Trinidad (2 species and 2 subspecies). Remarkably, 33% of the 150 recorded species have not been reported
in the last 80 years. While it is possible that some of these are not resident or have become extinct, it seems more likely that most have simply not been found in the last 80 years. The butterfly fauna of Tobago merits further study; year-round collecting in different habitats and areas, using a variety of techniques, will surely fi ll in many of the apparent gaps in our knowledge. Ouleus fridericus sheldoni ssp. nov. (Hesperiidae, Pyrginae) is described from Tobago, with illustrations of adults and male genitalia, and is compared to O. fridericus sinepunctis (Kaye) from Trinidad. Danaus plexippus tobagi A.H. Clark, 1941 is a syn. nov. of D. plexippus nigrippus (Haensch, 1909) (Nymphalidae, Danainae).
0250
Larvae of thirty one species of antlions (Neuroptera: Myrmeleontidae) belonging to eleven genera live in the protection of cave mouths or large rock overhangs in Australia. New taxa proposed here include the following three new genera: Australeon, Newleon, Speleon. The following twelve new species are described: Froggattisca kakadu; Froggattisca rennerensis; Newleon fragilis; Speleon cavernicolus; Speleon pilliga; Speleon yallingup; Stenoleon xanthopsis; Xantholeon cavernicolus; Xantholeon kakadu; Xantholeon newi; Xantholeon pallens; Xantholeon pentlandensis. Two cave species are transferred into a new genus becoming new combinations: Australeon illustris (Gerstaecker), Australeon manselli (New and Matsura). The small non-cave species previously placed in Stenoleon Tillyard are transferred to Bandidus Navás, becoming B. gradostriatus (New), B. copleyensis (New), B. grandithecus (New), and B. navasi (New), new combinations. All known species of six genera (Stenoleon Tillyard, Xantholeon Tillyard, Eophanes Banks and three new genera) are known only in cave mouths. Also, species of Heoclisis Navás, Froggattisca Esben Petersen, Glenoleon Banks, Heoclisis Navás and Myrmeleon Linnaeus contain species living in cave mouths. Two main types of caves are found in Australia; those with loose organic material and those with loose inorganic material. The cave habitat is divided into four zones and several subzones. Many species are restricted to one zone or another but species of Stenoleon may overlap zones. Discussions of the species and some of their biological requirements are provided. One new parasite record is given, an undetermined species of Echthrobacella Girault (Hymenoptera: Encyrtidae) reared from the larvae of Speleon yallingup Miller and Stange. Diagnoses or descriptions are given for all of these cave species including distributional data. Keys to the cave mouth inhabiting antlions of Australia (adults and larvae) are provided including a revised key to the Xantholeon Tillyard.
0338
Scolopendra morsitans L., 1758, is documented from Honolulu, Oahu, Hawaiian Islands, the fi rst record of this anthropochoric chilopod from both the archipelago and state. Hawaii thus becomes the second American state to harbor the species, the other being Florida, where an individual has been taken in Jacksonville, Duval County. Meristic and morphological data are presented for three Hawaiian specimens. At least two other species of Scolopendra, both introduced, occur on these islands: S. polymorpha Wood, 1861, known only from one specimen from Oahu, and one or more representatives of the “S. subspinipes Leach, 1815, complex,” which is widespread and even inhabits Midway Atoll.
0315
A key is provided for the twelve taxonomic groups within the genus Therates Latreille (Coleoptera: Cicindelidae). The chennelli group is reviewed here and a key to the 53 species known for this group is provided. Each species is illustrated, and a detailed description of its morphology and distributional records are provided. Therates khaoyaii, Th. pearsoni, Th. safraneki, Th. schuelei and Th. sigridgeissleri are described as new to science. Therates differens Sawada and Wiesner, 1999 is placed into synonymy under Th. concinnus Gestro, 1888. Therates tonkinensis kubani Wiesner, 1988 is placed into synonymy under Th. tonkinensis Horn, 1902 and Th. belokobylskiyi Matalin and Wiesner, 2006 is placed into synonymy under Th. haucki Moravec and Wiesner, 2001. Lectotypes are designated for Th. waagenorum Horn, 1900, Th. clavicornis Horn, 1902, Th. tonkinensis Horn, 1902 and Th. annandalei Horn, 1908. Therates baolocensis Wiesner, 1996 is elevated to species rank.
490
Recent shifts in US policies towards Cuba suggest a relaxation or lifting of the embargo may occur in the near future. With the prospects of open travel and trade with Cuba come concerns over the introduction of agricultural pests. In an effort to assess these concerns the distribution-based introduction risk of pests listed in the 2015 Cooperative Agricultural Pest Survey’s (CAPS) list of priority pests of economic and environmental importance is reviewed. Of the 59 pests on the CAPS priority pest list, 20 have been recorded in the literature as being present in the Caribbean Basin, South America and Central America. For these 20 New World pests a commodity and distribution-based risk rating was assigned to describe their potential for introduction through the Cuba-Florida pest pathway. The highest rating was given to the six listed pests currently reported as being present in Cuba, and potential for introduction and subsequent impact of these six pests on Florida agriculture is discussed. In addition to the pests found on the 2015 CAPS priority pest list, information regarding pests of concern in the family Tephritidae and the Old World bollworm Helicoverpa armigera (Hübner), is also included, as is a description of the Cuban plant health and regulatory structure.
The significance of plant-pest introductions between Cuba and Florida is discussed, with an emphasis on proactive engagement in research and collaboration to address these issues.
948 [rev. ed.]
A key, an annotated checklist with detailed distribution, biological and host information, and color photographic plates are provided for the 91 species of dacine fruit flies (Diptera: Tephritidae: Dacini) known to occur in Oceania. Dacus virgatus Coquillett, previously a synonym of Bactrocera psidii (Froggatt), is instead considered a junior synonym of B. facialis (Coquillett). The species originally described in 1971 as Dacus (Asiadacus) perpusillus Drew, later reassigned as Bactrocera (Sinodacus) perpusilla (Drew) and in recent years as Zeugodacus (Sinodacus) perpusillus (Drew) actually belongs to genus Dacus, and is transferred back to Dacus, but to the subgenus Neodacus, restored combination. The presence of B. redunca (Drew) is recorded for the first time in New Caledonia. New male lure records include isoeugenol and dihydroeugenol for both B. neoxanthodes Drew and Romig and B. quadrisetosa (Bezzi) and zingerone for Dacus taui (Drew and Romig), all in Vanuatu.
ZooBank registration. urn:lsid:zoobank.org:pub:A105F057-F2A4-4C14-B82E-14912B319D57
948
A key, an annotated checklist with detailed distribution, biological and host information, and color photographic plates are provided for the 91 species of dacine fruit flies (Diptera: Tephritidae: Dacini) known to occur in Oceania. Dacus virgatus Coquillett, previously a synonym of Bactrocera psidii (Froggatt), is instead considered a junior synonym of B. facialis (Coquillett). The species originally described in 1971 as Dacus (Asiadacus) perpusillus Drew, later reassigned as Bactrocera (Sinodacus) perpusilla (Drew) and in recent years as Zeugodacus (Sinodacus) perpusillus (Drew) actually belongs to genus Dacus, and is transferred back to Dacus, but to the subgenus Neodacus, restored combination. The presence of B. redunca (Drew) is recorded for the first time in New Caledonia. New male lure records include isoeugenol and dihydroeugenol for both B. neoxanthodes Drew and Romig and B. quadrisetosa (Bezzi) and zingerone for Dacus taui (Drew and Romig), all in Vanuatu.
ZooBank registration. urn:lsid:zoobank.org:pub:A105F057-F2A4-4C14-B82E-14912B319D57
712
There are 214 species of the suborder Tipulomorpha (Diptera) known from Panama. Of these, 162 species were described by the noted American entomologist, Charles Paul Alexander, from Panama during the period from 1912 through 1979. Panama hosts two of the four families found within the Tipulomorpha: Limoniidae (197 species) and Tipulidae (17 species). We have records of only three species from multiple provinces (n = 2) within Panama and 211 species from just one province. The vast majority of the latter are known from a single location within their respective provinces. There are 142 endemic species (66.4%) included in Panama’s Tipulomorpha fauna, with the remainder being found elsewhere in South America (48 species), Central America (44), Mexico (23 species), the Caribbean (14 species), and the United States (4 species). Whereas, this taxonomic group has benefited from collecting in the Canal Zone and the popular western highlands in Chiriqui Province, we know little or nothing of the distribution of species within the country, their natural history, or their relationship to water quality.
0352
The Guadeloupe Archipelago, the French overseas Département de Guadeloupe, is a geographically associated group of islands and a natural biogeographic unit. The islands have been available for terrestrial colonization since the late Tertiary. From the viewpoint of beetle systematics and biodiversity, this is the most important set of islands of the Lesser Antilles because more species have been described or recorded from Guadeloupe than any other island or group in the Lesser Antilles. We present a summary of the 1338 beetle species recorded in the literature from the archipelago, in 60 families, and 719 genera. The families with the largest numbers of species are Curculionidae (420), Staphylinidae (153), Chrysomelidae (75), Cerambycidae (69), Scarabaeidae (64), and Tenebrionidae (59). Four hundred eighty two species are known only from one or more islands of the Guadeloupe group and likely speciated there. Guadeloupe is the type locality for an additional 59 species. At least 61 species have been accidentally introduced by human activities. A total of 261 species are known only from the Lesser Antilles including Guadeloupe. The remaining species are naturally more widespread in the Lesser Antilles, or the West Indies, and elsewhere in the New World. The actual number of species on the Guadeloupe Archipelago is estimated to be around 1850 or more species.
0159
This paper summarizes the published information on the beetle fauna of the northern Leeward Islands (Anguilla, Antigua, Barbuda, Nevis, Saba, St. Barthélemy, St. Eustatius, St. Kitts, St. Martin-St. Maarten, and smaller associated islands, excluding Montserrat). These islands are generally smaller, lower, and drier than the remaining Leeward and Windward islands of the Lesser Antilles island arc. The fauna contains 26 families, with 155 genera, and 218 species. The families with the largest number of recorded species are Staphylinidae (36), Cerambycidae (28), Scarabaeidae (25), Tenebrionidae (23), Curculionidae (18), and Carabidae (15). At least 7 species (3.2% of the fauna) were probably introduced to the island by human activities. Sixteen species (7.3%) are endemic (restricted) to a single paleo-island bank and likely speciated there. Twenty nine species (13.3%) are shared only with other islands of the Lesser Antilles (Lesser Antillean endemics), and 43 species (19.7%) are more widespread Antilles endemics. The remaining 123 species (56.4%) in the fauna are otherwise mostly widely distributed in the Antilles and the Neotropical Region. The local beetle fauna is largely an immigrant fauna and has mostly originated elsewhere than on the islands of the northern Leewards. Summary data on total species endemicity of the entire Lesser Antilles indicate the presence of at least 1278 endemic beetle species, which is a density of about 20.7 species per 100 km2. This is now equivalent to that of the endemic vascular plants of the Caribbean islands. This truly makes the Caribbean islands a biodiversity hotspot for beetles. For the northern Leewards, it is evident that the beetle diversity is markedly understudied, and that the actual number of species is many times higher than now known.
642
The dung beetle fauna of the Big Bend region of Texas (Coleoptera: Scarabaeidae: Scarabaeinae)
(2018)
This paper reports the results of a 2001-2009 field study of the scarabaeine dung beetle fauna (Coleoptera: Scarabaeidae) of the Big Bend region of Texas, a three-county area of the Trans-Pecos portion of the Chihuahuan Desert. The observed fauna comprises 10 native species, Canthon blumei Halffter and Halffter, C. imitator Brown, C. praticola LeConte, and C. mixtus Robinson; Onthophagus browni Howden and Cartwright, O. knausi Brown, O. velutinus Horn and O. brevifrons Horn; Copris arizonensis Schaeffer and Phanaeus texensis Edmonds; as well as two exotic species introduced in the 1970s, Digitonthophagus gazella (Fabricius) and Euoniticellus intermedius (Reiche). The existing native fauna antedates the completion of desertification approximately 9,000 yrs BP and is similar ecologically and taxonomically to those in southeastern Arizona and northern Mexico. Ecological distribution follows three broad, overlapping habitat zones: desert montane forest, desert grassland and desert scrub. Species accounts include diagnoses, geographic distribution data, and information on collection method, habitat distribution and daily activity.
0024
The seven genera and 13 species of dynastine scarabs recorded from the Bahamas are reviewed. Two of those species are endemic, including Cyclocephala dolichotarsa Ratcliffe and Cave, new species, described from Great Inagua Island. Eleven species are also known to occur in the USA and/or Cuba. Six species are probably not established based on infrequency of collection.
0139
The five genera and eight species of dynastine scarabs occurring in the Cayman Islands in the West Indies are reviewed. Two new, endemic species are described from Little Cayman, with supporting illustrations: Tomarus adoceteus Ratcliffe and Cave (Pentodontini), new species, and Caymania nitidissima Ratcliffe and Cave (Phileurini), new genus and species.
0433
The fauna of Dynastinae (Scarabaeidae) on the island of Saba, Dutch Caribbean, was investigated through fi eldwork during 2006 to 2015. Three species, belonging to the three tribes Cyclocephalini, Pentodontini and Phileurini, are newly recorded from Saba and are discussed, with summaries of all relevant information from the West Indies. Detailed locality data, temporal distributions, and habitus photographs are presented for each species.
0444
Pandirodesmus rutherfordi, n. sp., represented by 18 individuals including eight adult males, occurs in secondary forests near Charlotteville and Speyside, Tobago, Trinidad and Tobago. Along with the type and second species, P. disparipes Silvestri, from Guyana and known only from females, the segmental legs of P. rutherfordi alternate between long (anterior pairs) and short (posterior ones), spiracular openings are on straw-like tubules, and ozopores are located on paramedian metatergal spines. These features appear to be adaptations for biotopes of loose sand, detritus, or frass, and 17 specimens, including the six juveniles, exhibit coatings of “sand grains” that are loosely cemented together and to the smooth, translucent, grayish-white exoskeleton. The tubules and spines elevate the spiracles and ozopores above the coating, thereby ensuring that they remain open and functional.
The coating, which provides camoufl age and lends strength and rigidity to the poorly sclerotized exoskeleton, is a subuniform “pavement” that covers the entire animal except the labrum/clypeus, tarsal and antennal apices, prozonae, paraprocts, and the gonopods in males. Ramose/dendritic setae, particularly on narrowly rounded podo-/antennomeres, trap “sand grains,” and the ozopore secretions apparently constitute the “glue” that cements the coating, as evidenced circumstantially by layers of “sand” between the spines on the anterior metaterga, where they are physically closest. The alternating segmental leg lengths, in part due to differing ventrolateral and ventromedial origins, appear to be an adaptation for lateral/sideways motion in which the long (anterior) legs extend laterally and pull the body to the level of the short (posterior) ones, which continue the motion while the anterior legs extend to begin the next stroke. The opposing legs perform the complementary pushing motion a fraction after the long legs initiate the pulling stroke and hence are slightly and purposefully out of sync. An adult male paratype lacks the coating, probably because it had just molted and lacked time to amass it; the juvenile female paratype of P. disparipes also is “naked,” as was, according to Silvestri, the now lost adult female holotype. Until fresh material is collected, coatings cannot be confi rmed for P. disparipes even though it shares the anatomical modifi cations that seem adaptions for such. The minute, triramous gonotelopodites of P. rutherfordi are unlike any known for a chelodesmid, so the current generic placement, in a monotypic tribe in the nominate chelodesmid subfamily, is retained. With species in both South America and the southern Antilles, Pandirodesmus/ini had to exist on both the “proto-Antillean” terrane and the adjoining part of Pangaean Gondwana before the former rifted in the Cretaceous/Paleocene, ~66 million years ago, and P. rutherfordi is a remnant of the former population that became isolated on present-day Tobago when the terrane fragmented. Affi nity between Guyanan and southern Antillean platyrhacid millipeds (Polydesmida: Leptodesmidea) suggest that Pandirodesmus/ini may occur sporadically as far north in the island chain as St. Lucia.
724
In two separate occurrences, graduates of the Oregon Forest Pest Detector program discovered the exotic Agrilus cyanescens (Ratzeburg) (Coleoptera: Buprestidae) in Portland, Oregon, damaging a backyard shrub, Lonicera involucrata (Rich.) Banks ex Spreng. (Caprifoliaceae). Although first detected in the USA 99 years ago, the known occurrence nearest to Oregon is in Utah.
973
Thirteen new fossil eucnemid taxa (Coleoptera: Elateroidea) are described from amber deposits excavated from the vicinity of Santiago, Dominican Republic. Two new genera, Mioxylobius and Paleoquirsfeldia are described. The following 13 new species are described from Dominican amber: Mioxylobius bicolor, Balistica serrulata, Paleoquirsfeldia epicrana, Dyscharachthis dominicana, Idiotarsus poinari, Euryptychus antilliensis, Euryptychus hispaniolus, Plesiofornax caribica, Fornax dominicensis, Fornax serropalpoides, Dromaeolus argenteus, Nematodes miocenensis and Nematodes thoracicus. Each new species are both diagnosed and illustrated. Calyptocerus Guérin-Méneville and Lissantauga Poinar are shown to be congeneic, resulting in a new combination: Calyptocerus epicranis (Poinar, 2013). Summaries of fossil eucnemid discoveries, highlighting differing hypothesis of prehistoric Caribbean island formations/speciation, accounts of ancient Dominican Republic environmental conditions and Dominican Republic amber are provided.
ZooBank registration. urn:lsid:zoobank.org:pub:48A76A23-E48B-46B5-8A35-A27DD6134B6D
0187
Cithaeron praedonius O. P.-Cambridge 1872 (Araneae: Gnaphosoidea: Cithaeronidae) is an Old World species with a distribution from The Gambia, western Africa, and Greece to Malaysia and Australia. In the New World, it was recently found in Brazil, and is now reported for the first time in North America, in the United States. Multiple individuals of both sexes and various life stages, including multiple eggsacs, have been found in a home in Port Richey, Pasco County, Florida. An adult female was found on the outside wall of the house feeding on another spider, suggesting that C. praedonius are no longer contained as a spot introduction in this one house. Observations in captivity indicate that this species may prefer feeding on other spiders. The eggsac and molting nest are described for the first time, and the first records on fecundity are reported.
0036
894
The jumping bristletail Pedetontus gershneri Allen, 1995 (Microcoryphia: Machilidae) is reported from the state of Alabama, USA, for the first time, extending its known range east-southeast by roughly 588 km. The first partial mtDNA barcode was generated for the species. Notes on color variation and sampling needs of Microcoryphia are briefly discussed.
911
Lissomus quisqueya new species and L. woodruffi new species are described. Both species are reported from the Dominican Republic on the island of Hispaniola. These are the first species of Lissomus Dalman reported from the Greater Antilles. Lissomus quisqueya is recorded from montane mesic forests in the Cordillera Central from Dajadon and La Vega provinces. Lissomus woodruffi is recorded from Barahona, Independencia and Pedernales provinces from montane mesic forest in the Sierra de Bahoruco. Morphologically, these species are most similar to members of the L. discedens Bonvouloir species group from Mesoamerica and South America. Known sites of occurrence of L. quisqueya and L. woodruffi are remnants of formerly more extensive mesic forests. Some of the specimens were found in secondary forests and ecotonal areas at sites near to primary forest or remnants, and all these forests are threatened by anthropogenic activities and stochastic climate changes.
0203
Four milliped species, substantiated by preserved voucher samples, are reported from Prince Edward Island, Canada. All are introduced European species that now occur widely in both Canada and the United States, and the panglobal Asian paradoxosomatid, Oxidus gracilis (C. L. Koch, 1847), is listed as probable. Choneiulus palmatus (Némec, 1895) (Julida: Blaniulidae) is newly recorded from New Brunswick, and four representatives of the Julidae are cited from Nova Scotia. Discovery of Cylindroiulus punctatus (Leach, 1815) (Julidae) in this province constitutes the second record from both Canada and North America, the other being in Newfoundland.
0264
Aplagiognathus Thomson, 1861 (Coleoptera, Cerambycidae, Prioninae, Macrotomini) and its two species, A. spinosus (Newman, 1840) and A. hybostoma Bates, 1879, are redescribed and figured. As the type of A. spinosus is apparently lost, a neotype for the species is designated herein as is a lectotype for A. hybostoma. Keys to the North and Central American genera of Macrotomini (excluding the West Indies) and to Aplagiognathus species are provided. Details on the numerous changes in the nomenclatural history of the genus are also chronicled.
1041
Members of the genus Centris Fabricius (Hymenoptera: Apoidea: Anthophila) constitute a significant component of the Neotropical (including insular) bee fauna, exhibiting high species richness, a moderate to large body size, and extensive interactions with various important plant groups. Females of most species possess specialized morphology adapted for collecting oils from flowers. This study documents the presence of the genus in Cuba, recognizing six species: C. aethiops Cresson, C. cornuta Cresson, C. fulviventris Cresson, C. poecila Lepeletier, C. taina Genaro and Breto new species, and C. tarsata F. Smith. Detailed information is provided for each species, encompassing a diagnosis, natural history, floral associations, seasonal occurrence, and distribution. Centris taina new species is described from Cuba, based on both sexes, which were previously misidentified as C. versicolor (Fabr.) for females and C. fasciatus F. Smith for males due to sexual dimorphism. Centris tarsata is reported as a new national record for Cuba, possibly introduced by humans from South America and now established and widely distributed across the entire island. A key to differentiate the Cuban species of Centris is presented.
ZooBank registration. urn:lsid:zoobank.org:pub:257916DF-2129-4694-876C-49C858046BF6
0229
Currently, the genus Chimarra Stephens (Trichoptera: Philopotamidae) is represented in the Oriental Region by 259 species. Of these, 61 species have been described or recorded from Vietnam. In this paper, 9 new species from Vietnam are described and illustrated (Chimarra aculeata, C. carinata, C. corneola, C. insolita, C. mina, C. prominens, C. rostrata, C. undulata, and C. ungula). In addition, 3 new country records are noted (Chimarra areli Malicky and Mey, Chimarra pipake Malicky and Chantaramongkol, and Chimarra suthepensis Chantaramongkol and Malicky), and 1 new species group (minuta Group) is proposed and populated. An additional species group (georgensis “Group”), with 1 new species from Vietnam, but otherwise only known from Africa, is discussed, but not formally defined. A table listing all known Vietnamese species of Chimarra is included, along with discussion of variability in the anal veins of the forewing found within this genus, and its relevance for defining subgenera and species groups.
0051
In the eastern United States, the genus Cotinis Burmeister previously contained only C. nitida (L.), the common economic pest known as the "Green June Beetle". A new species from the Florida Keys, Cotinis aliena, is here described and illustrated. A checklist is provided for the genus, which includes 27 valid New World species, and 44 synonyms.
0248
The genus Gromphas Brullé (Coleoptera: Scarabaeidae) comprises four species, of which G. aeruginosa (Perty) and G. amazonica Bates are known to occur in Peru. This paper presents a revised description of Gromphas as well as illustrated diagnoses and distributional and ecological data on the Peruvian species.
805
The genus Ochodaeus in Italy: taxonomy and distribution (Coleoptera: Scarabaeoidea: Ochodaeidae)
(2020)
The author provides a taxonomic, nomenclatural and distributional review of the genus Ochodaeus Dejean, 1821 (Coleoptera: Scarabaeoidea: Ochodaeidae) in Italy. All Italian populations have been confirmed to belong to a single species, O. chrysomeloides (Schrank, 1781). After the study of a syntype, O. cychramoides Reitter, 1892, formerly considered an Italian endemic, is confirmed to be a junior synonym of O.chrysomeloides. Type material of O. chrysomeloides is believed to be destroyed, therefore a neotype is here designated and deposited at the Natural History Museum of Vienna, Austria. A lectotype is here designated for O. cychramoides and deposited in the Hungarian Natural History Museum of Budapest, Hungary. The Italian distribution of O. chrysomeloides is given in detail and illustrated by a map.
0180
Eight species of the genus Psilotreta Banks (Trichoptera: Odontoceridae) are currently known from Vietnam: P. albogera Mey 1997, P. androconiata Mey 1997, P. bidens Mey 1995, P. enikoae Oláh and Johanson 2010, P. frigidaria Mey 1996, P. jaroschi Malicky 1995, P. papaceki Malicky 1995, P. spitzeri Malicky 1995. A new species, Psilotreta kurenschikovorum, from Thua Thien-Hue Province is herein described. The new species differs from other species of the genus by peculiarities in wing venation, by the unusual shape of epicranial suture on the head, and by the bifid apical segment of the inferior appendage. Additional province and collection information for previously recorded species are included.
0326
Five species of the aphid genus Sipha Passerini (Hemiptera: Aphididae) are reported in North America and are reviewed herein. Of these species, three are adventive species and include: Sipha elegans del Guercio, Sipha glyceriae (Kaltenbach), and Sipha maydis Passerini. Sipha maydis was discovered in California in 2007 and now has been found in Georgia. The genus also includes two native species: Sipha agropyronensis (Gillette) and Sipha flava (Forbes). Sipha maydis can be distinguished easily from all the other species in the genus that occur in North America because it is black. All the species except S. agropyronensis have been implicated in damage to crop plants. A key to the apterae and alatae of Sipha found in North America is included.
547
The eight species in the genus Tomarus Erichson (Coleoptera: Scarabaeidae) in Argentina, Chile, and Uruguay are reviewed. Tomarus roigjunenti new species and Tomarus spinipenis new species are described from Argentina. We include a key to species, representative habitus illustrations for all species, character illustrations, and distribution maps for each, as well as commentary about the natural history and distributions for each species. Diagnostic characters are discussed for each species, and species relationships are hypothesized based on the analysis of internal and external morphological characters. The male of T. bidentulus (Fairmaire) is described for fi rst time. The following taxonomic changes are made: Tomarus guianucai Dechambre and Lumaret, 1985 is a new junior synonym of Tomarus rubripes (Boheman, 1858), which was formerly and incorrectly cited as occurring in Argentina.
1039
U.S. port of entry interception data revealed that Frankliniella Karny (Thysanoptera: Thripidae) was the most frequently intercepted genus of Thripidae. No published identification resource for intercepted Frankliniella is available to USDA port of entry entomology identifiers. A morphological review of intercepted species was conducted. A dichotomous key for the identification of intercepted slide mounted adult females was created. Morphological diagnosis, description of each species with their geographic region(s) of origin and frequency of interception is provided.
ZooBank registration. urn:lsid:zoobank.org:pub:403B362E-9A7F-4385-A0F1-9DB87FE09AD2
747
The adventive mole cricket species (Orthoptera: Gryllotalpidae) in Hawaii is apparently Gryllotalpa krishnani Arun Prasanna et al., 2012. The adventive was long thought to be Gryllotalpa africana Palisot de Beauvois, 1805. From 1983 until 2008 it seemed that Gryllotalpa orientalis Burmeister, 1839 might be the adventive’s correct name. However, male genitalia and tegmental characters of Hawaiian specimens match features of G. krishnani and not G. orientalis.
0079
The formerly monotypic weevil genus Platytenes Pascoe (Coleoptera: Curculionidae: Cryptorhynchinae: Cryptorhynchini: Cryptorhynchina) as treated here, includes two commonly encountered and broadly distributed species in the eastern Indo-Australian region. Label data indicate that both species are associated with semicultivated betel palm (Areca catechu L., Arecaceae) and human mediated dispersal is suggested to have influenced their present-day distribution. Herein, we provide a diagnosis for Platytenes and redescribe its type species, P. varius Pascoe. We also describe P. occultus Setliff and Larson, a new species from the Solomon Islands. New host and locality records are provided, including the first records of the genus occurring on Ambon Island, the Bismarck Archipelago, D’Entrecasteaux Islands, and many previously unreported localities on New Guinea. A full bibliography, key to species, and distribution map for the genus are also provided.
0136
Orthosiphon stamineus Bentham, a medicinal plant in the family Lamiaceae, is used to make a well known herbal tea in many countries including Malaysia. Since its establishment as an important cash crop, the herb has been relatively free from any serious insect problems until recently. In Selangor, Malaysia we observed the herb heavily infested by the lace bug Cochlochila bullita Stål (Heteroptera: Tingidae). This is the first record of its occurrence in Malaysia and also the first record on the host plant, O. stamineus. The lace bug damages the host plant by piercing and sucking young leaves and shoots, resulting in the curling and drying of the leaves and shoots. The infestation pattern and survival of C. bullita on O. stamineus indicates this lace bug has the potential to be a serious pest of this medicinal plant.
861
Peru’s Cosñipata Region in Cuzco and Madre de Dios Departments is a valley between Manu National Park and the Amarakaeri Communal Reserve that ranges from 400 to 4,000 m elevation. A team of experienced lepidopterists sampled the butterfly fauna of this valley for more than a decade (7,440 field person-hours). We analyze the data for Lycaenidae (Lepidoptera: Papilionoidea), a family for which we have taxonomic expertise. After adding data on the fauna from museums and the literature, we present an annotated checklist of the 340 Lycaenidae species recorded from the Cosñipata Region with notes for each species on the elevations and seasons at which it occurs, adult behavior, and sampled relative abundance. Species richness is twice that recorded for Trinidad or Brazil’s Parque Nacional do Itatiaia, each of which also has a mix of low, mid, and high elevation habitats. There was an average of 8.3 adult specimens per species in the fieldwork sample. For those species with more than 8 specimens, the median elevational range was 1,100 m. Species richness in low elevation habitats was greater than that at mid or high elevations, which is contrary to findings for some other Neotropical insects. We present evidence why further sampling is likely to increase this difference. Maximal adult species richness occurs during the transition from dry to wet seasons (September to November) at all elevations, but there is little evidence that adults of species occur only during this season. Sampled relative abundances were skewed so that 70% of the species were encountered less frequently than average (1/340). These results are consistent with the observation that most species are rarely encountered using standard sampling methods.
559
The Antillean fauna of Mantispidae (Neuroptera) consists of six species in four genera, all in the subfamily Mantispinae. Two new species are here described: Leptomantispa antillesensis Hoffman, n. sp., is described from Puerto Rico, Anguilla, British Virgin Islands (Grand Camanoe, Guana, Tortola), United States Virgin Islands (St. Croix, St. John, St. Thomas), Dominica, and Guadeloupe, and Leptomantispa hispaniolaensis Hoffman, n. sp. is described from Hispaniola (Dominican Republic). Mantispilla zayasi Alayo is synonymized under Leptomantispa pulchella (Banks), new synonomy, and Mantispilla taina Alayo is synonymized under Zeugomantispa minuta (Fabricius), new synonomy. Climaciella cubana Enderlein is recorded for the fi rst time from Hispaniola (Dominican Republic). Dicromantispa sayi (Banks) is newly reported from the Cayman Islands (Cayman Brac and Little Cayman), and Hispaniola (Dominican Republic). Zeugomantispa minuta (Fabricius) is now recorded from the Bahamas (Andros) and Hispaniola (Dominican Republic).
810
Matsucoccus krystalae Ahmed and Miller, new species, (Hemiptera:Coccomorpha: Matsucoccidae) is described based on morphological characters of adult females and third-instar males. We designate the lectotype of Matsucoccus alabamae Morrison. Matsucoccus alabamae Morrison and M. gallicolus Morrison are redescribed, also based on adult females and third-instar males. Detailed illustrations and descriptions are presented for each species and an identification key for the five species occurring in the eastern U.S. is provided. Analysis of 18S, 28S D2/D3, and 28S D10 loci were performed to support morphological determination. Barcodes using 5′ COI of M. alabamae and M. krystalae were generated, the first such data from authoritatively identified Matsucoccus species. Of particular interest is that most of the specimens in the study were taken in Lindgren funnel traps.
999
The Egyptian fauna of mealybugs (Hemiptera: Coccoidea: Pseudococcidae) is reviewed and an illustrated key to the 30 genera and 54 species is provided. Phenacoccus madeirensis Green is reported for the first time in Egypt. A new genus, Ezzatacoccus Evans and Abd-Rabou, is described and illustrated with Amonostherium arabicum Ezzat, 1960 designated as its type species. Octococcus salicicola Priesner and Hosny, 1935 is reinstated as a valid taxon and transferred to Misericoccus Ferris, new combination. Ripersia cressae Hall is transferred to Maconellicoccus Ezzat, new combination and Planococcus lindingeri (Bodenheimer) is transferred back to Formicococcus Takahashi, revised status.
ZooBank registration. urn:lsid:zoobank.org:pub:8CA7B000-E8D4-463D-95B0-431BA0A7BA57
0083
The class Diplopoda, represented by the families Spirostreptidae (Spirostreptida) and Paradoxosomatidae (Polydesmida), is recorded from Saudi Arabia for the first time. Archispirostreptus transmarinus Hoffman, 1965 (Spirostreptidae) inhabits the Jabal Al-Hijaz Mountains in the southwest, and the Paradoxosomatidae, represented by an unidentifiable, indigenous female, occurs in a “wadi” in the center of the country. Other Middle Eastern familial records are documented, and occurrences in the Arabian Peninsula are mapped. Males, necessary to identify the paradoxosomatid, may be encountered if samplings are timed to coincide with seasonal rains.
0105
With documentation of an unidentifiable adult female and juvenile Tingupidae (Chordeumatida), Kodiak Island, Alaska, becomes the westernmost indigenous diplopod locality in North America including continental islands. The northernmost and most proximate locality, Yakutat, lies ca. 935 mi (1,496 km) to the eastnortheast, while Haines, the type locality of Tingupa tlingitorum Shear and Shelley, some 1,196 mi (1,914 km) in this direction, is the most proximate familial site. Kodiak is also one of the most remote indigenous milliped localities in the Pacific, the most proximate ones to the west and south, Kamchatka, Russia, and the Hawaiian Islands, United States, being over 3,300 mi (5, 280 km) distant. Tingupidae is recorded for the first time from Canada excluding the Queen Charlotte Islands, and geographically remote, ostensibly indigenous records from the North Pacific Ocean and environs are tabulated.
0207
The milliped genus Euryurus Koch, 1847, and the species, E. leachii (Gray, 1832) (Polydesmida: Euryuridae), are recorded from three sites on the northern part of Crowley’s Ridge (Cross, Lee, and Poinsett counties), Arkansas, where the only prior familial records are of Auturus evides (Bollman, 1887). Coupled with the published locality of E. leachii in Phillips Co., at the southern extremity of the Ridge, the only known occurrences of both the genus and species in Arkansas and west of the Mississippi River are in this physiographic feature. The Arkansas population is geographically peripheral but anatomically intermediate between the two recognized subspecies, E. l. leachii and E. l. fraternus Hoffman, 1978, and we do not assign it to a race. Molecular investigations seem necessary to resolve relationships in the “E. leachii complex.”
0196
The taxonomically neglected milliped order Glomeridesmida and family Glomeridesmidae (infraclass
Pentazonia, superorder Limacomorpha) inhabit 21, rather than seven, regions of the world, being newly recorded
from Thailand; Cambodia; the Republics of Palau, the Philippines, and Vanuatu; New Britain, Bismarck Archipelago;
the Island of New Guinea (both West Papua [formerly Irian Jaya], Indonesia, and Papua New Guinea);
and Sulawesi and Borneo, Indonesia. Occurrence in Fiji is confirmed with two additional samples, and discovery is
predicted in southern China, Myanmar, and perhaps Madagascar. Coupled with published localities, these records
suggest subcontinuous (super)ordinal and familial ranges extending some 12,480 km (7,800 mi) southeastward from
northwestern Thailand to Fiji. Though infrequently encountered, the taxa may actually be diverse and abundant
within this area, which encompasses all of the Indochina and Malay peninsulas, the Philippines, Palau, the Island
of Borneo and Indonesia, Papua New Guinea, the Solomon and Santa Cruz Islands, Vanuatu, and Fiji; it excludes
Taiwan, Australia, New Caledonia, and the Loyalty Islands. The paucity of preserved individuals probably results
from their dark pigmentations and minute sizes, adults being <6.5 mm long; Berlese extractions and sieved litter
techniques are recommended over hand collecting. Glomeridesmida are much more continuous, widespread, and
abundant in the “east” than previously believed and clearly do not comprise a minor, insignificant taxon. The first
glomeridesmidan photos are published.
0132
0283
The first reports of the multicolored Asian lady beetle Harmonia axyridis (Pallas, 1773) (Coleoptera: Coccinellidae) in Colombia appeared in 2011. However, based on museum insect specimens, the introduction of H. axyridis in Colombia occurred in 1989 or earlier, making it the second oldest record of the species in South America after the deliberate releases of the species in Argentina in 1986. Currently in Colombia, H. axyridis is well established and is here recorded from the States of Antioquia, Caldas, Cauca, Cundinamarca, Nariño, Tolima and Valle del Cauca.
953
Megapsyrassa Linsley, 1961 is synonymized with Psyrassa Pascoe, 1866 (Coleoptera: Cerambycidae). Aneflomorpha martini Chemsak and Linsley, 1968 is synonymized with Psyrassa sinaloae Linsley, 1935, and the species is transferred to Aneflomorpha Casey, 1912, new combination. Morphological and chromatic variations in Psyrassa cylindricollis Linsley, 1935 are reported, and a new state record is provided. New records are provided for the following species: Psyrassa atkinsoni (Chemsak and Giesbert, 1986) new combination; P. ebenina Linsley, 1935; and P. nigripes Linsley, 1935. Lastly, four new species of Psyrassa Pascoe, 1866 are described: Psyrassa wappesi García and Santos-Silva, from Mexico (Michoacán); P. sonorensis García and Santos-Silva, from Mexico (Sonora); P. obscuriventris García and Santos-Silva, from Mexico (Jalisco); and P. ocularis García and Santos-Silva, from Guatemala (Zacapa).
ZooBank registration. urn:lsid:zoobank.org:pub:194F7545-EF7C-44B7-9783-286A8BDEB1EC
721
The Palearctic flea beetle Epitrix pubescens (Koch) (Coleoptera: Chrysomelidae: Galerucinae: Alticini) is reported as established in North America. It is recorded in the United States from Illinois, Massachusetts, New Hampshire, New York, Oregon, and Wisconsin, and in Canada from the Provinces of Nova Scotia, Ontario, and Quebec. The oldest records are from 1975. The species has been reported feeding on Solanum dulcamara L. (Solanaceae). Full distributional information and reference photos for identification are provided. Epitrix pubescens may have avoided detection for such a long time due to its size, difficulty of identification, and affinity for a weed rather than agricultural crops. The more than 40-year lag between arrival and discovery of an immigrant species illustrates the importance of routine biodiversity monitoring efforts and taxonomic works.
1009
Hurd (1952), in revising the Nearctic species of Pepsis Fabricius, separated P. cerberus Lucas from P. elegans Lepeletier based on external morphology and geography. Vardy (2005), in his Western Hemisphere Pepsis revision, combined these taxa and several Neotropical color and structural variants in a broad definition of P. menechma Lepeletier extending across ~11,250 km and two continents. Vardy (2005) synonymized the familiar and well-documented, 160-year-old P. elegans under P. menechma probably because it appeared several pages later in Lepeletier’s (1845) Histoire Naturelle des Insectes. Hyménoptères. Vardy’s (2005) interpretation of Pepsis menechma as a viable species presents a taxonomic and nomenclatural problem. He violated the principle of nomenclatural stability in synonymizing the widely and established species names P. elegans and P. cerberus under P. menechma, a name that had not been used for 160 years. Recent discoveries warrant a re-evaluation of the problematic taxonomy of this species complex. Morphological and ecological divergence of P. elegans and its sister taxon, P. cerberus, combined with their narrow sympatric distribution justifies species recognition. Hurd’s (1952) two species concept for P. elegans and P. cerberus is more practicable, useful, and nomenclaturally acceptable than Vardy’s (2005) P. menechma. Pepsis cerberus Lucas and P. elegans Lepeletier should be reinstated as species and removed from the synonymy of Pepsis menechma Lepeletier.
ZooBank registration. urn:lsid:zoobank.org:pub:F59B3131-74DE-4704-9936-337E380BF3E0
537
The Republic of Panama currently has 21 recorded species of stoneflies, all in the genus Anacroneuria (Plecoptera: Perlidae). Herein, we record five species of this genus from the Mount Totumas Cloud Forest and Biological Reserve, in the upper reaches of the Río Chiriquí Viejo watershed. One of these species, A. plutonis (Banks), represents a new country record for Panama. These results are part of an ongoing effort to characterize the aquatic insect fauna of Panama, and to evaluate that country’s major watersheds.
788
Psyllids are an economically important group of insects. Several species are serious emerging pests with regulatory significance. About 20 adventive species have been discovered in Florida in the past 20 years, including several pests. Additionally, five species new to science have been found. We provide an annotated checklist of Florida species with taxonomic information and identification tools, including keys to Florida genera and known species. Seventy species of Psylloidea currently are reported from Florida. Forty-one are native to Florida, with 12 endemic to the state. Twenty are adventive, the majority being from the Neotropics. One was introduced deliberately for biological control, seven represent temporary populations (eradicated, reared in quarantine), and one is a dubious record. Craspedolepta euthamiae Burckhardt and Halbert, new species, Katacephala wineriterae Burckhardt and Halbert, new species, Pseudophacopteron gumbolimbo Burckhardt and Halbert, new species, Nothotrioza longipedis Burckhardt and Halbert, new species, and Trioza myresae Burckhardt and Halbert, new species are described from Florida and are native endemic species. Aphalara persicaria Caldwell is redescribed and separated from similar species. Aphalara persicaria var. cubana Caldwell is confirmed as a junior synonym of Aphalara persicaria. The Florida records of Craspedolepta spp. are revisited and revised, including Craspedolepta euthamiae Burckhardt and Halbert, new species. Bactericera nigrilla (Crawford), new combination, revived status is recognized from Florida, redescribed, and distinguished from similar species. Rhinopsylla caldwelli Tuthill is transferred to Kuwayama Crawford and becomes Kuwayama caldwelli (Tuthill), new combination.Trioza maritima Tuthill is transferred to Leuronota Crawford and becomes Leuronota maritima (Tuthill), new combination. Species of Bactericera Puton on Salix L. (Salicaceae) in North America are reviewed. Bactericera flori (Crawford), new combination, new status. is determined to be the correct name for Trioza assimilis Crawford nec Flor (= Trioza flori Crawford, replacement name, = Trioza pomonae Aulmann, replacement name), and Trioza dubia Patch, new synonym. Lectotypes are designated for Trioza marginata Crawford, Trioza minuta Crawford, Trioza minuta similis Crawford, and Trioza nigra Crawford.
886
Platystasius transversus (Thomson) (Hymenoptera: Platygastridae) is a rarely collected egg parasitoid of Leptura aurulenta Fabricius (Coleoptera: Cerambycidae). Four female specimens were found in Germany, a new country record for the genus and species. Illustrations, DNA barcodes, and an updated distribution are provided. We review its taxonomic history, biology, and ecological associations.
995
We studied the slides of Nipaecoccus viridis (Newstead) (Hemiptera: Pseudococcidae) deposited in the Florida State Collection of Arthropods and provided specimen label data, including geographical location, global positioning system coordinates when available, host plant, collector name, adult females/immature stages, sex of specimens on respective slides, number of slides, and collection date. In addition, we discuss its first record from Florida ornamental landscape and two most recent new host records of the species on commercial crops, including blueberries and hemp. These data will help regulatory agencies slow the spread of this pest inside and outside of Florida.
ZooBank registration. urn:lsid:zoobank.org:pub:EEB2767C-9B59-4BA9-A5B5-D5B14FF7B38F
949
A list of abbreviations regarding literature, collections and persons as used by early authors (1758–1779) of scarabaeoid beetles (Insecta: Coleoptera) is given together with modern referrals to the Literature Cited. Notes regarding referential errors are included. Hyperlinks to all mentioned and freely online available publications are provided.
ZooBank registration. urn:lsid:zoobank.org:pub:8508A5D0-CA65-4BBB-9FD4-8D14AC261F72
787
An annotated checklist of 227 species of Scarabaeoidea (Coleoptera) known to occur in Iowa is pre-sented, including seven families and 79 genera. This list was developed from information gleaned during literature review, from examination of specimens deposited in 22 regional and comprehensive collections, as well as from several years of active collecting conducted by the authors around the state. Thirty-six species included here rep-resent new state records.
0309
The six species of Siricidae (Hymenoptera: Symphyta) occurring in Florida and one species of regulatory concern introduced to North America are reviewed. Included are diagnoses of Eriotremex formosanus (Matsumura), Sirex areolatus (Cresson), Sirex nigricornis Fabricius, Tremex columba (Linnaeus), Urocerus cressoni Norton, Urocerus taxodii (Ashmead) and Sirex noctilio (Fabricius). A key to species, photographs of morphological features, biological notes and distribution data are provided. For the species T. columba, S. nigricornis, U. taxodii, and U. cressoni, a total of eight new state records are presented.
1020
A faunal study for the order Phasmatodea of the Chicaque Natural Park is presented, including a list of species found, descriptions, redescriptions, and biological notes. A total of nine species were found and studied; two new genera: Ramandeun new genus, Nubilophasma new genus, and four new species: Atratomorpha jorgei new species, Isagoras franciscoverai new species, Nubilophasma chicaquensis new genus and new species, and Ramandeum coronatum new genus and new species are described. The description of the eggs of the new taxa, of the previously unknown eggs of Paraceroys quadrispinosus (Redtenbacher, 1906), and the redescription of the eggs of Libethra rabdota Stål, 1875, and Libethra inchoata Brunner von Wattenwyl, 1907 are provided. Additional comments on the ecology and morphological variation of the reviewed taxa are included. Finally, further studies on the stick insect fauna of the Colombian Andes are discussed and recommended to provide more information to broaden the understanding of the species that inhabit this complex mountain system.
ZooBank registration. urn:lsid:zoobank.org:pub:7D34FF9A-CDEE-4DD4-A643-E0F467E00A5B
857
0435
The Republic of Panama currently has 245 recorded species of Trichoptera distributed among 13 families. Herein we add 32 new country records for Panama, including one new family (Limnephilidae) and one new genus (Limnephilus). We also provide the first collection information for Oxyethira maya Denning for Panama. These results are part of an ongoing effort to characterize the caddisfly fauna of Panama, and to evaluate that country’s major cuencas (water basins).