Insecta Mundi
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742
In the course of working on new species of North American Phyllophaga Harris, 1827 (Coleoptera: Scarabaeidae: Melolonthinae) some synonyms have been found and are proposed here. New synonymies: Phyllophaga knausii (Schaeffer, 1907) is synonymized with Phyllophaga sociata (Horn, 1878); Phyllophaga chippewa Saylor, 1939 is synonymized with Phyllophaga rugosa (Melsheimer, 1845); and Phyllophaga falta Sanderson, 1950 is synonymized with Phyllophaga bipartita (Horn, 1887). Lectotypes are here designated for the following species: Listrochelus knausii Schaeffer, Listrochelus sociatus Horn, and Lachnosterna bipartita Horn. A neotype for Ancylonycha rugosa Melsheimer is here designated from the Horn Collection.
705
Prothyma (Genoprothyma) thandamoeae Wiesner, Phyu and Hori, new species, Prothyma (Genoprothyma) sotai Wiesner, Phyu and Hori, new species, and Prothyma (Genoprothyma) asamii Wiesner, Phyu and Hori, new species (Coleoptera: Cicindelidae) are described from Myanmar. A key to all members of the genus known to occur in Myanmar is given.
939
Tomoxia bucephala A. Costa (Coleoptera: Mordellidae), a Palearctic tumbling flower beetle native to Europe, Asia, and northernmost Africa, is now known from North America. The first known occurrences were in 2015 in Essex and Union counties, New Jersey, U.S.A. and in 2019 in Passaic County, New Jersey, all in the New York City metropolitan area. An additional collection documents the species in 2016 from Allegheny County, Pennsylvania, in the Pittsburgh metropolitan area. The multiple occurrences, the large distance between those in New Jersey and Pennsylvania, and multiple detections in natural areas indicate T. bucephala is established in North America and apparently invasive. Several morphological features differentiate T. bucephala from the two congeners native to North America, T. inclusa LeConte and T. lineella LeConte, especially coloration patterns of elytral and pronotal vestiture, and coloration of antennae and front legs. This is the first report of a non-native mordellid species established in North America. Tomoxia bucephala does not appear to pose a significant direct economic threat in North America since it feeds in decaying trees. However, T. bucephala occurrences are within the geographic ranges of T. inclusa and T. lineella, and the biology of T. bucephala is similar to these other Tomoxia species. Thus, T. bucephala likely will expand its range within North America, with probable ecological impact on communities of native saproxylic beetles, especially T. lineella and T. inclusa.
ZooBank registration. urn:lsid:zoobank.org:pub:134762B2-9F05-4F02-88F8-4BDCB4231F0F
718
0167
Neolecanium amazonensis Foldi is redescribed and illustrated and is transferred to the new genus Foldilecanium Kondo as Foldilecanium amazonensis (Foldi) comb. nov. A new species, Foldilecanium multisetosus Kondo, is described and illustrated based on specimens collected in Cali, Colombia, on Cananga odorata (Lam.) Hook.f. and Thomson (Annonaceae). An updated taxonomic key to New World Myzolecaniinae and a key to separate the two species of Foldilecanium are provided.
709
Five small species groups of Traumatomutilla André, 1901 are revised. A new species, T. pilkingtoni Bartholomay and Williams sp. nov. (Argentina) is described and placed within its own species group. All known species of the bellica, diabolica, vitelligera, and bifurca species groups are redescribed. New synonymies are proposed for T. vitelligera (Gerstaecker, 1874) = Mutilla fascinata Smith, 1879, syn. nov.; = Traumatomutilla comata André, 1906, syn. nov. and for T. bifurca (Klug, 1821) = T. ira Casal, 1969, syn. nov. The lectotypes of Mutilla bifurca Klug, 1821 and Mutilla comata André, 1906 are designated. The hitherto unknown males of T. vitelligera and T. bifurca are diagnosed and described.
0416
0388
Trixagus steineri (Coleoptera: Throscidae), a new species and first genus record from The Bahamas
(2014)
Trixagus Kugelann 1794 is the second largest genus of Throscidae with 80 valid species presently assigned. Horn (1885, 1890) reviewed the species for the United States and Mesoamerica, and then Blanchard (1917) revised part of the family for Canada and the United States in a posthumous article edited by H.C. Fall. Schenkling (1928) provided the only published worldwide catalog. Yensen (1975) provided a modern revision for the species for Canada and the United States, and then (Yensen 1980) described T. cobosi from Panama and provided a new key to all of the described American species. Among these studies only T. chevrolati (Bonvouloir 1859) was recorded from southern Florida, thus being a species potentially shared with The Bahamas. Aulonothroscus bicarinatus Fleutiaux (1911, 1947) (Blackwelder 1944), from Guadeloupe, is the only other throscid species previously described from the West Indies. Prior to the work of Lawrence and Newton (1995) the subfamily Lissominae received divergent treatment and was usually treated as a subfamily of Throscidae (e.g., Schenkling 1928, Blackwelder 1944, Yensen 1975). This subfamily is represented in The Bahamas by two species of Drapetes Megerle 1821 (Turnbow and Thomas 2008). Representation of the subsequently restricted family in The Bahamas was provided by two undetermined species of Aulonothroscus Horn listed by Turnbow and Thomas (2008) from Andros, Eleuthera, and Great Inagua islands; these will be treated separately from here. The reporting of a new species of Trixagus from New Providence Island provides a second genus from the country and the entire Lucayan Archipelago, and suggests that related species of both genera from Hispaniola and Cuba are probable and await discovery (Peck 2005; Perez-Gelabert 2008).
0240
The mirine plant bug Tropidosteptes forestierae, new species (Hemiptera: Miridae) is described from
Collier County, Florida, where it was found causing serious injury to an extensive ornamental hedge of Florida swampprivet, Forestiera segregata (Jacq.) Krug and Urb. (Oleaceae). Adult male and female, fifth instar, and egg are described. Color images of the adults, nymph, egg, and injury; scanning photomicrographs of selected adult structures; and illustrations of male genitalia are provided. A key to help distinguish the 16 species of Tropidosteptes known to occur in the southeastern United States is given.
0482
512
Two new species of Eburiini (Coleoptera, Cerambycidae, Cerambycinae) are described from Mexico (Guerrero): Susuacanga boteroi; and Eburia (Eburia) girouxae. Both are included in previous keys.
577
Two new genera and species of tiger beetles from Baltic amber (Coleoptera: Carabidae: Cicindelinae)
(2017)
Two fossil tiger beetle species (Coleoptera, Carabidae, Cicindelinae) are described from Eocene Baltic amber using light microscopic and X-ray microscopic techniques. Both species are considered representatives of the subtribe Iresina Rivalier, 1971 due to the shared combination of character states: glabrous head, six labral and four suborbital setae, and glabrous pronotum. Palaeopronyssiformia groehni Wiesner, Will, and Schmidt, new genus, new species, is characterized by a glabrous and furrowed head with six labral setae, large eyes, presence of two supraorbital setae on each side, mandibles with two teeth of the incisor region, and a glabrous and furrowed pronotum. Palaeoiresina cassolai Wiesner, Will, and Schmidt, new genus, new species, is characterized by a unicolored, undentated labrum, mandibles with two teeth of the incisor region, glabrous head with six labral setae, two clypeal setae, two supraorbital setae on each side, and a glabrous pronotum, mesepisternum, mesepimeron, and metepisternum. The species described here represent the only known tiger beetle fossils preserved in Baltic amber.
994
Two new genera, Skelleyus Opitz and Divulgoatus Opitz (Coleoptera: Cleridae: Enopliinae), show unusual antennal development. Funicular antennomeres become shorter as they approach an extensive capitulum. This study involves three taxa, Skelleyus leavengoodi Opitz, new species, Divulgoatus kelleri Opitz, new species, and Divulgoatus discrepans (Gorham).
ZooBank registration. urn:lsid:zoobank.org:pub:E059A916-DDE2-4193-A090-8CDD46EDC859
0232
0197
The Mesoamerican species of Telephanus distinguished by the presence eight lateral pronotal spines
and long temples are reviewed. The group includes T. serratus Nevermann and two previously undescribed species
that are described herein: T. bellus Thomas, new species, from Costa Rica, and the flightless T. monstrosus
Thomas, new species, from Mexico.
855
Phyllophaga (Listrochelus) benwarneri new species and Cinacanthus cunninghami new species(Coleoptera: Scarabaeidae) are described from dune systems in northern Arizona and extreme southern Nevada, USA. Habitus and diagnostic characters of these species are illustrated, and couplets from the respective, most recent (sub)generic keys are modified to incorporate the new species. Available habits and habitat information for each species, and a discussion of the Phyllophaga “senex complex” of species are provided.
0360
Two new species and a new genus of Cerambycidae are described from South America: Cotyclytus arriagadai sp. nov., from Bolivia; and Lembu dieguezi, gen. nov., sp. nov., from Paraguay. Orthomegas irroratus (Lameere, 1915) is redescribed, based on the second and third known specimens, and its distribution is expanded to include Ecuador. The male of Jamesia fuscofasciata Dillon and Dillon, 1952 is described and illustrated for the fi rst time, and the distribution of the species is expanded to Peru. Thirty-two new country records (twelve for Paraguay, fi fteen for Peru, two for Ecuador, three for Bolivia) and one new province record (Argentina) are presented.
738
Saphenista bartellae Brown, new species (TL: Colorado), and S. powelli Brown, new species (TL: California) (Lepidoptera: Tortricidae), are described and illustrated. Two other western North American species, S. latipunctana (Walsingham, 1879), new combination, and S. dilutana (Walsingham, 1879), new combination, are transferred to Saphenista based on morphology of the genitalia.
0380
Two new species of pselaphine staphylinids in the genus Batrisodes are described: B. (Declivodes) dorothae Ferro and Carlton from Feliciana Preserve, Louisiana; and B. (Babnormodes) spretoides Ferro and Carlton from Great Smoky Mountains National Park, Tennessee. They differ from all other described Batrisodes species in secondary male characters, especially details of the frontal region of the head. These two species bring the total diversity of the genus in North America to 88 species. Specimens were imaged using light microscopy, scanning electron microscopy, and X-ray microtomography (micro-CT) techniques. Utility of multiple imaging techniques, especially micro-CT, is discussed.
1033
The genus Lichnanthe Burmeister (Coleoptera: Scarabaeoidea: Glaphyridae) has been comprised of eight Nearctic species and contains the only New World members of their family. Here, two new species of arenicolous bumblebee scarabs are described from the central United States. The recent discovery of an undescribed Wyoming species led to the reevaluation of a dubious 134-year-old Nebraska Lichnanthe specimen, indicating that it also was an undescribed species. These two species, Lichnanthe brusti new species from central Wyoming and L. bruneri new species from central Nebraska are immediately distinguishable from all existing species of the genus by their square mandibles.
ZooBank registration. urn:lsid:zoobank.org:pub:401927DA-66D9-47A1-AE68-965210824713
534
813
0195
0454
Two new species of Corimbion Martins, 1970 are described from Bolivia: Corimbion kuckartzi and Corimbion ledezmae. A previous key to the South American species of Corimbion (Martins 2009) is herein modified to include the new species. Dorsal, ventral and lateral habitus illustrations, as well as variation in color and dorsal pattern for C. kuckartzi, are also presented.
811
900
1027
Identifications of recently collected Eucnemidae (Coleoptera: Elateroidea) borrowed from two collections have resulted in the discovery of two new species: Entomophthalmus abbreviatus Otto (Cuba) and Trigonopleurus cordobaalfaroi Otto (Guatemala and Louisiana, USA). Images of the two newly described species along with two New World Entomophthalmus Bonvouloir species for comparative purposes are included.
ZooBank registration. urn:lsid:zoobank.org:pub:E83B2AC6-33ED-4680-8F2E-4E455A26D4C8
1003
893
778
Nothochodaeus yeti Huchet, new species, from Nepal and Sikkim (north India) and N. martensi Huchet, new species,from Nepal, are described and illustrated (Coleoptera: Scarabaeoidea: Ochodaeidae). A distribution map and an updated catalogue of the Ochodaeidae occurring on the Indian subcontinent are provided.
859
Two species of palm-leaf skeletonizers (Lepidoptera: Pterolonchidae: Homaledra Busck) that are invasive in Florida, USA are newly described. Homaledra howardi new species is described from Florida and the Dominican Republic. Homaledra knudsoni new species is described from Florida, southern Texas, and Mexico. A lectotype is designated for Homaledra sabalella (Chambers). The three are diagnosed and compared with other congeners, and Homaledra is compared with other Pterolonchidae. The larvae of the three common species feed on epidermis of palm foliage, causing significant damage.
837
Two species of the early-diverging lineages of Pharaxonotha Reitter (Coleoptera: Erotylidae: Pharaxonothinae) are described: Pharaxonotha taylori Skelley and Tang, new species, and Pharaxonotha thomasi Skelley and Tang, new species. A new key to described species of Pharaxonotha, based on previously unused characters, is presented.
827
Phyllophaga (Phyllophaga) badbunnyi Cano new species, and Phyllophaga (Phyllophaga) sechi Cano new species, of the Schizorhina species group (Coleoptera: Scarabaeidae: Melolonthinae), are described and illustrated from specimens collected at cloud forests between 1300-1750m elevation of southeastern Chiapas, México andwestern Guatemala. A key to the Guatemalan and Chiapanecan species of Phyllophaga Harris, group Schizorhina, with dorsal finger-like projections of genital capsule, is provided.
699
Two new species of Onciderini Thomson, 1860 (Coleoptera: Cerambycidae: Lamiinae) are described and illustrated: Neocherentes adrianoi Nearns and Monné, from Brazil, and Neocherentes pergeri Nearns and Monné, from Bolivia. The male of Neocherentes dilloniorum Tippmann, 1960 is redescribed and the female is described for the first time. Neocherentes dilloniorum is excluded from the Brazilian fauna. A key to the known species of Neocherentes Tippmann, 1960 is provided.
516
Himaloaesalus gaoligongshanus Huang and Chen, new species (Coleoptera: Lucanidae: Aesalinae) is described from the Gaoligongshan Mts., Yunnan, China. It is the fi fth species of the genus Himaloaesalus Huang and Chen. This new species is similar to the Himalayan species Himaloaesalus himalayicus Kurosawa and H. saburoi Araya et al., from which it is distinguished. The male and female genitalia of all the known species of Himaloaesalus are illustrated. Dorcus yongreni Huang and Chen, new species (Coleoptera: Lucanidae: Lucaninae) is described from Ruili area, Yunnan, China. It belongs to the elegans group (Huang and Chen 2013). This new species is similar to the Indian species Dorcus apatani (Okuda and Maeda), new combination, originally described in Digonophorus Waterhouse. The male genitalia of both species are compared and illustrated.
0225
Two new species of the genus Pahamunaya Schmid (Trichoptera: Polycentropodidae), P. talon sp. n. and P. spinifera sp. n., from Vietnam are described and illustrated. Examination of the holotype male of P. khoii Oláh and Johanson, in combination with an additional specimen of the same species, revealed new characters. New illustrations for this species are provided.
0412
Two new species of Aulonothroscus Horn are described from The Bahamas and a third species is newly reported. Aulonothroscus inawa new species is described from Great Inagua and Aulonothroscus sibateo new species is described from Eleuthera. Aulonothroscus convergens (Horn) is reported from Andros, providing an island and new country record. These are the fi rst Aulonothroscus identifi ed to species from the Lucayan Archipelago and from a West Indies locality other than Guadeloupe. A key to the species of Bahamian Aulonothroscus is provided.
0346
Examination of the type series of Schizotus gibbifrons Lewis 1887 and Pyrochroa higoniae Lewis 1895, together with examination of the historical Lewis collection of Pseudopyrochroa japonica (Heyden 1879), provide hints as to the conspecifi city of these binomials. This evidence, together with concurrent collection events spanning more than 100 years and general anatomy suggests Pseudopyrochroa japonica (Heyden 1879) and Pseudopyrochroa gibbifrons (Lewis 1887) are conspecifi c. Schizotus gibbifrons Lewis is proposed as a new junior synonym of Pyrochroa japonica Heyden, where it joins the established synonym, Pyrochroa higoniae Lewis (1895). Schizotus theresae Pic 1911 is also proposed as a new junior synonym of Pyrochroa basalis Pic 1906, where it joins a long list of synonyms. Complete synonymies are presented for both species.
704
Cylindera (Ifasina) thitarooae Wiesner and Hori, new species, Jansenia phyuae Wiesner and Hori, new species, and the previously unknown male of Jansenia myanmarensis Wiesner, 2004 (Coleoptera: Cicindelidae) are described. Cylindera (I.) thitarooae may be recognized by shape of labrum and aedeagus. Jansenia phyuae may be recognized by its elytral contour and shape of elytral maculation.
696
0477
Two new species of the mexicanus group of Vaejovis C.L. Koch are described from the Madrean pine-oak forests of the Sierra Madre Occidental in the state of Durango, Mexico. These species, Vaejovis sierrae sp. nov. and Vaejovis mcwesti sp. nov., are distinguished from each other and the only other species of the mexicanus group known from this mountain range, Vaejovis montanus Graham and Bryson, by morphometrics, carinal development of the pedipalps, granulation of the metasoma, and body size. A key to the species of the mexicanus group from
the Sierra Madre Occidental is provided.
0227
Two species of the weevil genus Compsus Schoenherr (Coleoptera: Curculionidae: Entiminae) from Colombia are redescribed: C. obliquatus Hustache and C.viridivittatus (Guérin-Méneville). A key by Hustache in 1938, to 33 of the 34 recognized species of Colombian Compsus then known, is modified to include the one additional species. Habitus illustrations of males and females of the two species and illustrations of selected parts of the male and female genitalia are included. Nearly all of the specimens of these two species were collected on various species or varieties of citrus, indicating their potential as citrus pests in the future.
0340
Tynommatidae, n. stat., elevated from Tynommatinae, is established as a schizopetalidean family encompassing the western North American callipodidans previously assigned to the Mediterranean Schizopetalidae. It is considered a valid taxon despite somewhat anatomically dissimilar subfamilies, and Colactidinae, Texophoninae, Diactidinae, and Aspidiophoninae constitute tribal elevations and additional new statuses. With a subbasal telopodal prefemoral process, Diactis hedini, n. sp., requires rediagnoses of all three diactidine genera, Diactis Loomis, 1937, and Florea and Caliactis, both by Shelley, 1996, and suggests that telopodal branches ‘B’ in congeners and Florea represent distal relocations of the process along the stem. Similarities in the sizes and shapes of the pleurotergal carinae suggest a sister-group relationship with the other, and partly sympatric, New World family, Abacionidae, which is supported by gonopodal similarities between Colactidinae and Abacion Rafi nesque, 1820. The Western Interior Seaway of the Cretaceous Period, Mesozoic Era, ~141–66 million years ago, appears to have fueled divergence by isolating “proto-abacionid stock” in “Appalachia,” the Eastern North American land mass, which has subsequently spread well into previously inundated areas. The allopatric position of Texophoninae, on the Gulf Coast of south Texas around 1,136 km (710 mi) east of the most proximate familial records, is attributed to this waterway, which eradicated faunal linkages with “proto-Tynommatidae” in “Laramidia,” the Western North American land mass. Texophoninae probably survived the Cretaceous on insular refugia; however, it is rarely encountered anymore and seems destined for imminent extinction. Representatives of the east-Asian families, Caspiopetalidae, Paracortinidae, and Sinocallipodidae, also possess demarcated pleurotergal crests and, implausible though it seems, may share ancestry with the North American taxa vis-à-vis the “Asiamerica” and or “Boreotropic” concepts.
0458
524
A new species of Autostichidae, Eraina beechei nov. sp., is described from specimens collected in the Andean mountains of Ñuble, Biobío. Diagnostic characters for the species are presented; photographs of adults, illustrations of male and female genitalia, and a key for identifi cation of Eraina species are provided.
727
The taxonomy of Nearctic tiger beetles (Coleoptera: Carabidae: Cicindelinae) is reviewed in light of modern systematics research. Despite decades of published molecular phylogenies, the taxonomic nomenclature has not been formally updated since the 1950s. We generated a maximum likelihood phylogenetic tree based on three mitochondrial gene fragments (16S, COX3 and CytB) to address the placement of Nearctic taxa that were not included in recent studies; these species were historically contained within Cylindera Westwood, 1831, a polyphyletic genus. Here we describe Parvindela Duran and Gough, new genus, and propose 20 new combinations based on a plurality of data, including our topology, prior molecular phylogenetic studies, morphology and ecology: Apterodela unipunctata (Fabricius, 1775) new combination; Brasiella praecisa (Bates, 1890) new combination; Brasiella viridisticta (Bates, 1881) new combination; Cicindela amargosae (Dahl, 1939) new combination; Cicindela senilis (G. Horn, 1866) new combination; Cicindela willistoni (LeConte, 1879) new combination; Eunota californica (Menetries, 1883) new combination; Eunota circumpicta (LaFerte, 1841) new combination; Eunota fulgoris (Casey, 1913) new combination; Eunota gabbii (G. Horn, 1866) new combination; Eunota pamphila (LeConte, 1873) new combination; Eunota praetextata (LeConte, 1854) new combination; Eunota severa (LaFerte, 1841) new combination; Eunota striga (LeConte, 1875) new combination; Parvindela debilis (Bates, 1890) new combination; Parvindela celeripes (LeConte, 1848) new combination; Parvindela cursitans (LeConte, 1860) new combination; Parvindela terricola (Say, 1824) new combination; Parvindela nephelota (Bates, 1882) new combination; Parvindela lunalonga (Schaupp, 1884) new combination.
497
The previously published list of Coleoptera holotypes in the Museo de Historia Natural, Noel Kempff Mercado (MNKM), Santa Cruz, Bolivia is updated to include those deposited in the Museo since then though the end of 2015. Literature citations for the original descriptions of each listed holotype are also provided along with summary comments regarding family composition and authorship of included species. Photographs of collector, authors, habitats, and major Bolivian type repositories are provided.
975
An updated list is given of 25 species of soft scale insects (Hemiptera: Coccidae) which have been intercepted on plants imported into South Korea during the period of 1996 to 2021. Information on the number of interceptions, host plants, distribution and origin of species intercepted at South Korean ports of entry is provided. In addition, data on intercepted species was analyzed to determine potential invasive species of soft scales that could threaten South Korean plants.
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0347
Pinnaspis chamaecyparidis Takagi, Pinnaspis hikosana Takagi and Pinnaspis uniloba (Kuwana), occurring on Chamaecyparis obtusa Endl., Styrax japonica S. et Z. and Cleyera japonica Thunb. are newly documented in the Korean fauna of armored scales (Diaspididae). The characters of these species are here redescribed with illustrative photographs and information on distribution and hosts along with a dichotomous key to the species of Pinnaspis for correct species identifi cation. In addition, the paper discusses the current status of Pinnaspis buxi (Bouché) and Pinnaspis strachani (Cooley) which are known as native armored scale insects of Korea by analyzing information on the result of the survey.
816
Quick and accurate identification of intercepted psyllids (Hemiptera: Psylloidea) is an essential requirement for effective pest management and phytosanitary procedures. However, due to lack of morphological characters in the immature stages that can be used to distinguish species, other molecular methods, such as DNA barcoding are proving to be useful. The current study was to designed to generate comprehensive information on the identification of all developmental stages of eight species of psyllids intercepted on consignments of infested fresh cut flowers at the ports of entry in South Korea using DNA barcoding. It is considered that DNA barcoding is a reliable technique for identification of intercepted psyllids for immature stages and will be helpful in the development of more effective pest management options for regulating pest species.