European journal of taxonomy : EJT
Paris : Muséum National d'Histoire Naturelle
ISSN: 2118-9773
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45
The classification of the largest subfamily of leafhoppers, Deltocephalinae, including 38 tribes, 923 genera, and 6683 valid species, is reviewed and revised. An updated phylogeny of the subfamily based on molecular (28S, Histone H3) and morphological data and an expanded taxon sample (37 taxa not included in previous analyses) is presented. Based on the results of these analyses and on the morphological examination of many representatives of the subfamily, the classification of the tribes and subtribes of Deltocephalinae is revised. Complete morphological descriptions, illustrations, lists of the included genera, and notes on their distribution, ecology, and important vector species are provided for the 38 recognized tribes and 18 subtribes. A dichotomous key to the tribes is provided. All names in the taxonomic treatments are hyperlinked to online resources for individual taxa which are supported by a comprehensive database for Deltocephalinae compiled using the taxonomic database software package 3I. The online functionality includes an interactive key to tribes and subtribes and advanced database searching options. Each taxon (subspecies through subfamily) has a unique taxon webpage providing nomenclatural information, lists of included taxa, an automated description (if available), images (if available), distributional information, bibliographic references and links to outside resources. Some observations and trends regarding the history of taxonomic descriptions in Deltocephalinae are reported. Four new tribes are described: Bahitini tribe nov. (25 genera), Bonsapeiini tribe nov. (21 genera), Phlepsiini tribe nov. (4 genera), and Vartini tribe nov. (7 genera). The circumscription and morphological characterization of Scaphoideini Oman, 1943 (61 genera) is substantially revised. Eleven new species are described: Acostemma stilleri sp. nov., Arrugada linnavuorii sp. nov., Drabescus zhangi sp. nov., Parabolopona webbi sp. nov., Goniagnathus emeljanovi sp. nov., Hecalus hamiltoni sp. nov., Scaphoideus omani sp. nov., Dwightla delongi sp. nov., Abimwa knighti sp. nov., Gannia viraktamathi sp. nov., and Doratulina dmitrievi sp. nov. Some family-group level taxonomic changes are made: Platymetopiini Haupt, 1929, Anoterostemmini Haupt, 1929, and Allygidiina Dmitriev, 2006 are synonymized with Athysanini Van Duzee, 1892, syn. nov.; Procepitini Dmitriev, 2002 is synonymized with Cicadulini Van Duzee, 1892, syn. nov.; Listrophorini Boulard, 1971 is synonymized with Chiasmini Distant, 1908, syn. nov.; Adamini Linnavuori & Al-Ne’amy, 1983, Dwightlini McKamey, 2003, and Ianeirini Linnavuori, 1978 are synonymized with Selenocephalini Fieber, 1872 syn.nov., and all three are now recognized as valid subtribes in their parent tribe. New placements of many genera to tribe and subtribe are made, and these are described in individual taxon treatments.
44
Based on newly designated type material, four poorly known NE Atlantic cheilostome bryozoan species are redescribed and imaged: Cellaria harmelini d’Hondt from the northern Bay of Biscay, Hippomenella mucronelliformis (Waters) from Madeira, Myriapora bugei d’Hondt from the Azores, and Characodoma strangulatum, occurring from Mauritania to southern Portugal. Moreover, Notoplites saojorgensis sp. nov. from the Azores, formerly reported as Notoplites marsupiatus (Jullien), is newly described. The genus Hippomenella Canu & Bassler is transferred from the lepraliomorph family Escharinidae Tilbrook to the umbonulomorph family Romancheinidae Jullien.
43
Two new species of Thomisidae are described (Mecaphesa reddelli sp. nov. and Tmarus galapagosensis sp. nov.). Of a third species, Mecaphesa inclusa (Banks, 1902), three colour variations are described. Tmarus specimens previously listed from the islands have always erroneously been called T. stolzmanni Keyserling, 1880. The Philodromidae are mentioned for the first time for the archipelago and are represented by two new species: Apollophanes fitzroyi sp. nov. and Apollophanes (?) lonesomegeorgei sp. nov.
42
We report the rediscovery of the Pied Butterfl y Bat, Glauconycteris superba Hayman, 1939, 40 years after this species was last recorded. The new specimen from Mbiye Island, Democratic Republic of the Congo, is compared with the type specimens of G. s. superba and G. superba sheila Hayman, 1947 and a specimen from Matonguiné, Ivory Coast. The variation in the striking colouration of the pelage as well as in morphometric data is considered to be individual rather than geographic variation and we tentatively regard G. s. sheila as a synonym of the nominate form. Despite the wide distribution of this species in the tropical forest zone of West and Central Africa, only four specimens from four localities are known to date, which might indicate very specific habitat preferences. Contemporary land cover information around historic collection sites shows degraded landscapes. Given the highly uncertain area of occupancy of this species, we suggest changing the status of G. superba in the IUCN Red List of Threatened Species from “Least Concern” to “Data Defi cient”.
41
The late Katian, Late Ordovician Boda Limestone of Dalarna, Sweden contains a rich cephalopod assemblage. The assemblage consists of 61 species, in 31 genera, comprising almost all major Ordovician cephalopod orders. Most common and diverse are the Orthocerida. The Ascocerida are also remarkably common and diverse. The new ascocerid species, Redpathoceras bullatum sp. nov., R. depressum sp. nov., R. magnum sp. nov., and Probillingsites scandinavicum sp. nov., give reason to revise current hypotheses on the origin and evolution of this group. An ascocerid origin from barrandeoceratids or aspidoceratids is hypothesised. The absence of actinocerids in the Boda Limestone is notable, and is interpreted as an indication of relatively cool and/or deep depositional environments. The dominance of orthocerids is provisionally interpreted as evidence for nutrient-rich waters during the time of the deposition of the Boda Limestone. Additionally, the assemblage contains the new barrandeocerids Schuchertoceras fryi sp. nov., Siljanoceras varians gen. et sp. nov., Warburgoceras gen. nov. (for Cyrtoceras longitudinale Angelin in Angelin & Lindström, 1880), the new endocerid Cameroceras turrisoides sp. nov., the new oncocerid Cyrtorizoceras thorslundi sp. nov., and the new orthocerids Dawsonoceras stumburi sp. nov., Isorthoceras angelini sp. nov., I. curvilineatum sp. nov., Nathorstoceras adnatum gen. et sp. nov., N. kallholnense gen. et sp. nov., Palaeodawsonocerina? nicolletoides sp. nov., Pleurorthoceras osmundsbergense sp. nov., and Striatocycloceras isbergi sp. nov.
40
Cteniogaster, a new genus of small ground spiders is described from Kenya and Tanzania. It encompasses seven new species, three of which are known from both sexes: C. toxarchus sp. nov., the type species, C. conviva sp. nov. and C. hexomma sp. nov. Three species are known from females only: C. lampropus sp. nov., C. sangarawe sp. nov. and C. taxorchis sp. nov. and one only from males: C. nana sp. nov. The new genus can be recognised by the presence of a posterior ventral abdominal f eld of strong setae and anterior lateral spinnerets with enlarged piriform gland spigots in males. A cladistic analysis attributes the genus to Liocranidae, Cybaeodinae. The results of the analysis performed do not produce an unequivocal autapomorphy for Liocranidae, but provide a combination of non-homoplasious character changes that offers significant potential for recognising genera as Liocranidae. Moreover, robust apomorphies are determined within Liocranidae for the subfamilies Liocraninae and Cybaeodinae. Based on these fi ndings Toxoniella Warui & Jocqué, 2002 is transferred from Gallieniellidae to Liocranidae, Cybaeodinae. Jacaena Thorell, 1897, Plynnon Deeleman-Reinhold, 2001 and Teutamus Thorell, 1890 are transferred to Corinnidae, Phrurolithinae and Montebello Hogg, 1914 to Gnaphosidae. Itatsina Kishida, 1930 is synonymised with Prochora Simon, 1886.
39
A taxonomic revision of the cichlid genus Astatoreochromis is presented. Eighteen meristic and 23 morphometric measurements were taken on 185 individuals, including type specimens. While fin counts separate populations from the Lake Victoria region (Astatoreochromis alluaudi) from those of the Rusizi and Malagarazi rivers in the Lake Tanganyika basin (A. vanderhorsti and A. straeleni respectively), clear differentiation was not detected between the latter two. Mann-Whitney U-tests on specimens of comparable size from the two Tanganyika populations revealed significant differences in specimens < 75 mm SL, mainly in pharyngeal jaw proportions, but also in some other measurements on the head and body. However, these differences were not detected in specimens > 75 mm SL and Astatoreochromis vanderhorsti is herein considered a junior synonym of A. straeleni. A redescription of the two valid species of Astatoreochromis, A. alluaudi and A.straeleni, is provided.
38
37
The genus Dadagulella gen. nov. is described to include 16 species of small, dentate, ovateacuminate Afrotropical snails. An identification key is provided and biogeography, anatomy and systematics are discussed. The type species is the Kenyan D. radius (Preston, 1910) comb. nov., whose name has informally been used for part of the group in the past. Substantial intraspecific variation occurs in three species: D. radius itself, D. browni (van Bruggen, 1969) comb. nov. and D. minuscula (Morelet, 1877) comb. nov. (= Ennea fi scheriana Morelet, 1881) (non Gulella minuscula Emberton & Pearce, 2000) . We recognise subspecies within each of these: D.radius radius (Preston, 1910) comb. nov., D. r. calva (Connolly, 1922) comb. et stat. nov., D. browni browni (van Bruggen, 1969) comb. nov., D. b. mafi ensis subsp. nov., D. b. semulikiensis subsp. nov., D. minuscula minuscula (Morelet, 1877) comb. nov., D. m. mahorana subsp. nov. Six new Tanzanian species are described: D. cresswelli sp. nov., D. delta sp. nov., D. ecclesiola sp. nov., D. frontierarum sp. nov., D. minareta sp. nov., and D. pembensis sp. nov. The genus includes seven other previously described species: D. cuspidata (Verdcourt, 1962) comb. nov.; D. rondoensis (Verdcourt, 1994) comb. nov.; D. conoidea (Verdcourt, 1996) comb. nov.; D. selene (van Bruggen & Van Goethem, 1999) comb. nov.; D. meredithae (van Bruggen, 2000) comb. nov.; D. nictitans (Rowson & Lange, 2007) comb. nov.; and D. delgada (Muratov, 2010) comb. nov.
36
Ostracods belonging to the genus Bennelongia differ much in valve morphology between adults and juveniles. Adult valves are asymmetrical, characterised by a beak-like feature in the anteroventral region of the left valve, and, with some notable exceptions, mostly have smooth or weaklyornamented valves. Juvenile specimens, on the other hand, have valves that are almost symmetrical, with no beak-like feature and are often heavily ornamented. We have examined the last 3 - 4 juvenile stages of 6 Bennelongia species from 5 different lineages, in order to decipher the types of external valve ornamentation and their recurrences during ontogeny and across lineages. It is clear that ornamentation is more prevalent at the early instar stages compared to the last 2 pre-adult stages, and especially when compared to the adult stage itself. We also examined the surprising presence of a calcified inner lamella with a prominent inner list in the pre-adult stages of Bennelongia species, that is usually absent in juveniles of other ostracods, thus questioning if heterochronic processes have provided an intermediate valve morphology between the simple (normal) cypridinid juvenile state and the heavily derived and modifi ed state of adult Bennelongia. We discuss the possible (speculative) functionality of the ornamentation in juveniles.