European journal of taxonomy : EJT
Paris : Muséum National d'Histoire Naturelle
ISSN: 2118-9773
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871
An extensive survey of Linyphiidae spiders from Wulipo National Nature Reserve (NNR), Chongqing has been conducted, in which 24 species belonging to 16 genera are recorded including two new genera and eight new species described here as: Agyneta Hull, 1911, A. orthogonia sp. nov. (♂♀); Dicristatus gen. nov., D. minutus gen. et sp. nov. (♂♀); Dicymbium Menge, 1868, D. pingqianense sp. nov. (♂♀); Himalaphantes Tanasevitch, 1992, H. azumiensis (Oi, 1979) (♂♀); Indophantes Saaristo & Tanasevitch, 2003, I. wushanensis sp. nov. (♂♀); Ketambea Millidge & Russell-Smith, 1992, K. nigripectoris (Oi, 1960) (♂♀); Molestia Tu, Saaristo & Li, 2006, M. pollicaris sp. nov. (♂♀); Neriene Blackwall, 1833, N. calozonata Chen & Zhu, 1989 (♀), N. cavaleriei (Schenkel, 1963) (♂♀), N. emphana (Walckenaer, 1841) (♂♀), N. japonica (Oi, 1960) (♂♀), N. limbatinella (Bösenberg & Strand, 1906) (♀), N. longipedella (Bösenberg & Strand, 1906) (♂♀), N. oidedicata van Helsdingen, 1969 (♀); Prosoponoides Millidge & Russell-Smith, 1992, P. sinense (Chen, 1991) (♂♀); Ryojius Saito & Ono, 2001, R. simplex sp. nov. (♂♀); Stemonyphantes Menge, 1866, S. bifurcus sp. nov. (♂); Syedra Simon, 1884, S. oii Saito, 1983; Tapinopa Westring, 1851, T. guttata Komatsu, 1937 (♀); Tenuiphantes Saaristo & Tanasevitch, 1996, T. ancatus (Zhu, Li & Sha, 1986) (♂♀); Walckenaeria Blackwall, 1833, Walckenaeria asymmetrica Song & Li, 2011 (♂♀); Wuliphantes gen. nov., W. guanshan (Irfan, Wang & Zhang, 2022) gen. et comb. nov. (♂♀), W. tongluensis (Chen & Song, 1988) gen. et comb. nov. (♂♀), W. trigyrus gen. et sp. nov. (♂♀). Male of Tenuiphantes ancatus (Zhu, Li & Sha, 1986) is described here for the first time as new to science. The taxonomic status of Bathyphantes guanshan Irfan, Wang & Zhang, 2022 and Bathyphantes tongluensis (Chen & Song, 1988) is revised and proposed here as: Wuliphantes tongluensis (Chen& Song, 1988) gen. et comb. nov. (♂♀) and Wuliphantes guanshan (Irfan, Wang & Zhang, 2022) gen. et comb. nov. (♂♀), respectively. Morphological descriptions, photos of body and copulatory organs, as well as the locality map are provided.
864:64-76
Here I describe a new genus, Iviephengus gen. nov., based on a single species, Iviephengus ferreirai gen. et sp. nov., from Peru. This new genus is characterized by the following combination of characters: interantennal distance close to 3 × the antennal socket length; antenna 12-segmented, IV to XI each with two long symmetrical compressed and apically slightly enlarged branches; labrum fused to frontoclypeus; mandibles short, obliquely crossed, each with a notch on the external margin to fit the other mandible and without extra teeth; maxillary palpi 4-segmented, last segment digitiform; labial palpi 2-segmented; posterior tentorial pit consisting of a single small fossa; wing with radial cell closed and transverse, vein r4 interrupted, r3 absent; first tarsomere of pro- and mesotarsus with a ventral comb covering the posterior half of the tarsomere; claws simple, without any teeth; aedeagus with paramere symmetrical, apex unevenly round, toothed inward, with short and sparse bristles. I provide a key to Mastinocerinae genera with 12-segmented antennae and the first pro- and mesotarsomere with ventral combs. I also provide illustrations for the diagnostic features for this new genus. Finally, I discuss the presence and function of some modifications in the mandible and the sternite VIII in Phengodidae and other Coleopteran families.
864:28-63
Three novel species collected from Dinghushan Biosphere Reserve (DHSBR) in southern China, Russula cylindrica Y.Song sp. nov. and R. lacteocarpa Y.Song sp. nov. in subgenus Archaeae and R. reticulofolia Y.Song sp. nov. in subg. Compactae, are described based on morphological and molecular data. In addition, Russula leucobrunnea Y.Song nom. nov. is proposed in replacement of R. leucocarpa nom. illeg. in subg. Brevipedum, as R. leucocarpa (T.Lebel) T.Lebel had been described earlier. Differences between the three novel species and their closely related taxa were analyzed. Another two known species in subg. Brevipedum, R. callainomarginis J.F.Liang & J.Song and R. japonica Hongo were also identified among specimens from DHSBR and are described and illustrated. Phylogenetic analyses of ITS and a five-locus phylogeny (concatenated LSU, mtSSU, rpb1, rpb2 and tef1) support the recognition of these taxa.
861:1-20
Novel data support validity of Phoxinus chrysoprasius (Pallas, 1814) (Actinopterygii, Leuciscidae)
(2023)
The common minnow species Cyprinus chrysoprasius, previously synonymised to Phoxinus phoxinus, was originally described from the Crimean Peninsula (Black Sea – Sea of Azov basin). A genetic analysis of the mitochondrial cytochrome oxydase 1 in the context of a phylogenetic study of European Phoxinus showed that it represents a distinct genetic clade and potentially a valid species. In the present study, we approach the issue following a broader, both genetic and morphological, study in order to check and support the validity of native Crimean Phoxinus under the earliest available name of the species: P. chrysoprasius. Our data demonstrate a reliable genetic distance of this minnow from geographically neighbouring clades and species, and a certain morphological distinctiveness. In order to determine the taxonomic concept of P. chrysoprasius, as a species involved in a genetically well-differentiated, but phenotypically poorly structured complex of east-European Phoxinus, a neotype for the species, based on topotypical material, is herein described and designated. The original type locality of the species is also clarified.
861:168-184
A new species, Clitopilus cretoalbus A.Izhar, Zaman, M.Asif, H.Bashir, Niazi & Khalid sp. nov., is described herein based on several collections from Punjab, Pakistan. It is characterized by a clitocyboid to somewhat omphaloid stature combined with a white pileus, decurrent lamellae, an almost central to slightly eccentric whitish relatively long stipe, the occurrence of cheilocystidia, and basidiospores with 6 to 9 ridges in polar view. Molecular phylogenetic analyses of nrITS and nrLSU performed using the maximum likelihood method supported the novelty of this Pakistani species and its placement within the genus Clitopilus section Scyphoides. A comparison with other morpho-anatomically close species confirmed that the newly described species is distinct from others.
860:116-140
Specimens of munidid squat lobsters belonging to the genera Heteronida Baba & de Saint Laurent, 1996 and Torbenella Baba, 2008 were collected in scientific expeditions made by several French cruises along the southwestern Pacific. These specimens were identified as two species (one new) of Heteronida and six species (four new) of Torbenella. The present paper provides systematic accounts of the five new species, along with new locality records of known species shown by this material and color information where available. Molecular data is provided to support the systematic status of each new species. A key to species for each of the genera is also presented.
847:73-103
The description in 1891 of the sea pen genus Gyrophyllum Studer, 1891 and also the type species G. hirondellei Studer, 1891 was based on a single colony collected in the Azores Archipelago. During the 19th and 20th centuries, the family placement of this genus became controversial as the set of morphological features present in Gyrophyllum could justify its assignation to both the families Pennatulidae Ehrenberg, 1834 and Pteroeididae Kölliker, 1880. Deliberations over this intermediate set of characters finally ended in the reunification of the genera and species of both families under Pennatulidae by principle of priority. The use of molecular sources of information based on a series of sequencing techniques presents a different but promising phylogenetic scenario in order to go further in the understanding of pennatulacean systematics. In this paper, a complementary morphological and molecular study (multiloci sequences with three mitochondrial and one nuclear markers) based mainly on newly collected material is carried out. This study re-confirms from a molecular point of view previously published results that indicate the position of Gyrophyllum as being distant from Pennatula Linnaeus, 1758 and Pteroeides Herklots, 1858 (type genera of the families Pennatulidae and Pteroeididae, respectively). This fact together with the results of a detailed morphological examination strongly supports the placement of the enigmatic genus Gyrophyllum in a separate family: Gyrophyllidae fam. nov. and resolves the nomenclatural uncertainty at family level for this genus. Moreover, the characters previously considered useful in the distinction of the two currently recognised species G. hirondellei in the Atlantic and G. sibogae Hickson, 1916 in the Indo-western Pacific are revisited.
846:1-41
We provide the first distribution pattern of the Microhyla heymonsi group based on available molecular and morphological data collected from East and Southeast Asia. Our analyses show a high level of genetic diversity in the M. heymonsi group with nine distinct lineages from China, Myanmar, Vietnam, Laos, Malaysia, Thailand, Cambodia, as well as Singapore, and Indonesia. The study also reveals the discovery of two new species in Vietnam, Microhyla hmongorum sp. nov. from Lai Chau Province and Microhyla xodangorum sp. nov. from Kon Tum Province. When comparing the 12S–16S rRNA gene, the genetic divergence between Microhyla xodangorum and other congeners of the Microhyla heymonsi group ranges from 7.5-- to 8.9% (M. cf. heymonsi) and approximately 8.4% between the new species and M. heymonsi s. str. from Taiwan, China. The genetic divergence between Microhyla hmongorum and its congeners ranges from 4.5–5.6% (M. cf. heymonsi) to 8.7% (Microhyla xodangorum). These new findings bring the total number of known species in the genus Microhyla to 48 and the recorded species of Microhyla from Vietnam to 14.
845:1-29
Based on morphological and molecular data we investigated the species boundaries in the genus Ochraethes Chevrolat, 1860. The species delimitation with cytochrome c oxidase subunit 1 (COI) data supports some synonymies and highlights the existence of a high phenotypic plasticity in some species. The following species are considered as new junior synonyms: O. virescens (Chevrolat, 1860) syn. nov. of O. brevicornis (Chevrolat, 1860), O. octomaculata Chemsak & Noguera 2001 syn. nov. of Ochraethes cinereolus (Bates, 1892) comb. nov., O. zebratus Bates, 1885 syn. nov. of O. obliquus (Chevrolat, 1860), Trichoxys giesberti Botero, Santos-Silva & Wappes, 2019 syn. nov. of O. sommeri (Chevrolat, 1835), O. clerinus (Bates, 1892) of O. viridiventris (Chevrolat, 1860), O. cristoforii (Chevrolat, 1860) syn. nov. and O. litura Bates, 1885 syn. nov. of O. z-littera (Chevrolat, 1860). Two new species are described from Mexico: Ochraethes confusus sp. nov., and Ochraethes nigroapicalis sp. nov.
835
The genus Cyparium Erichson, 1845 (Staphylinidae, Scaphidiinae, Cypariini) comprises 55 species, distributed mainly in the Neotropical and Oriental regions. Twenty-four species are known from the Neotropical region, but only eight species are reported from Brazil. In this paper we describe five new species and redescribe two species of Brazilian Cyparium, as follows: Cyparium achardi sp. nov., C. lescheni sp. nov., C. loebli sp. nov., C. newtoni sp. nov., C. pici sp. nov.; Cyparium collare Pic, 1920; and Cyparium oberthueri Pic, 1956. We provide images of adult males and females and their dissected parts, and information on host fungi whenever available. We also provide a comparative plate of dorsal colour patterns of Neotropical Cyparium.
834:38-57
The genus Cottus contains more than 60 species that are common in freshwater bodies of northern Eurasia and North America. Despite the abundance of species, this genus has been insufficiently studied in Eastern European rivers and lakes. The new taxon Cottus cyclophthalmus sp. nov. was found in the Neman/Nemunas and Venta river systems (Baltic Sea Basin). Cottus cyclophthalmus is sister and most morphologically similar to Cottus gobio. The new species has the following diagnostic characters: round, protruding (tubular) eyes near front of head; dermal papillae on top and sides of head, naked body, absence of bony prickles, full trunk canal with 32–36 pores. The description of Cottus cyclophthalmus and a new finding of Cottus microstomus improved understanding of the Eastern European biodiversity.
831:45-66
Five new species of the genus Andes Stål, 1866 from China (Hemiptera, Fulgoromorpha, Cixiidae)
(2022)
Five new species of the genus Andes Stål, 1866, A. balteiformis Wang, Zhi & Chen sp. nov., A. bifidus Wang, Zhi & Chen sp. nov., A. furcutus Wang, Zhang & Chen sp. nov., A. latanalus Wang & Chen sp. nov. and A. pallidus Wang & Chen sp. nov. from China, are described and illustrated. A key to the species of Andes in China is provided.
831:67-108
Coiled nautiloids of the Tournaisian and early to middle Viséan (Early Carboniferous) have so far only become known from a few regions. Here we describe material from five localities in southern Algeria; these belong to four stratigraphic horizons (two horizons in the late Tournaisian, one horizon near the Tournaisian–Viséan boundary, one horizon in the early to middle Viséan). From these, the new genera Stroborineceras gen. nov. and Trilobitoceras gen. nov. and the following new species are described: Rineceras tenerum sp. nov., Stroborineceras insalahensis gen. et sp. nov., Stroborineceras felis gen. et sp. nov., Stroboceras mane sp. nov., Stroboceras ancilis sp. nov., Vestinautilus angulatus sp. nov., Vestinautilus papilio sp. nov., Vestinautilus inflexus sp. nov., Vestinautilus bicristatus sp. nov., Trilobitoceras peculiaris gen. et sp. nov., Aphelaeceras azzelmattiense sp. nov., Maccoyoceras saharensis sp. nov., Maccoyoceras habadraense sp. nov. and Maccoyoceras concavum sp. nov.
828:109-137
Eight new Neotropical species of Dexosarcophaga Townsend, 1917 are described, five from Brazil, Dexosarcophaga phoenix sp. nov., Dexosarcophaga jandainae sp. nov., Dexosarcophaga patiuorum sp. nov., Dexosarcophaga petra sp. nov., and Dexosarcophaga sphaera sp. nov., one from Costa Rica, Dexosarcophaga limon sp. nov., one from Ecuador, Dexosarcophaga napo sp. nov., and one from Colombia, Dexosarcophaga pallida sp. nov. Male and female morphology is documented with photographs and illustrations, including details of the male terminalia for all new species and female terminalia of Dexosarcophaga phoenix sp. nov. and Dexosarcophaga sphaera sp. nov. With the addition of these new species, 58 species of Dexosarcophaga are now known, with records from the American continent spanning from the southern United States to northern Argentina.
826:1-32
Two new species of subgenus Heterophyllidiae subsection Cyanoxanthinae, Russula fusiformata Y.Song sp. nov. and R. purpureorosea Y.Song sp. nov., collected from the Dinghushan Biosphere Reserve (DHSBR), are described based on both morphology and a phylogenetic analysis of the internal transcribed spacer (ITS), further increasing Heterophyllidiae species diversity in the area. Differences between the two new species and related taxa are analyzed. The other 17 reported species of Russula subgenus Heterophyllidiae that have been collected from DHSBR during mushroom explorations since 2014 are also summarized. The dominant species and the ecological distribution of all 19 species are briefly discussed, and most species are presented in macrofungal plates.
826:80-93
Three new species of the genus Superciliaris Meng, Qin & Wang, 2020 are described from southern Vietnam, Malaysia (northern Borneo), and Indonesia (northern Sulawesi). The genus is recorded for the first time from mainland Asia and Sunda Archipelago. A key to species of Superciliaris is given. The relationships of the species are discussed.
826:94-134
The present study examines whether the chaetotaxy of the costal vein in the calyptrate families Fanniidae and Muscidae deserves more attention in phylogenetic and taxonomic contexts. An overview of the macrotrichia and their arrangement on wing vein C is given. Special attention is given to the presence/absence of ventral and dorsal setulae on the costal sectors CS1‒3. This is described as one variable character (A) with nine states (A0‒A8). Specimens of both sexes (when possible) of each species belonging to 4 of a total of 5 fanniid genera and 115 of a total of 179 muscid genus-group taxa were examined and scored for character A. It was found that the presumed ancestral state of character A differs between the two families. It is further shown that the main transformational trend in character A in Muscidae has been bi-directional, leading either to the loss of ventral setulae or the gain of dorsal setulae. The utility of character A in the Fanniidae and Muscidae is many-sided and involves taxa ranging from species to family. It is concluded that character A and other aspects of costal chaetotaxy deserve more attention in morphology-based studies of calyptrate flies.
823:64-81
Tardigrades reports from British Columbia (Canada) trace back to 1908 and numerous species have been recorded from this region, despite the relatively few published sampling studies. We describe by integrative taxonomy (light microscopy morphology, morphometrics, and DNA sequencing) a new tardigrade species, Sisubiotus hakaiensis sp. nov. from the British Columbia central coast. The new species has been found in moss collected from a vertical rock outcrop near the Hakai Institute Calvert Island Field Station. Sisubiotus hakaiensis sp. nov. differs from all the other known species in the genus by the presence of a labyrinthine layer inside the egg process walls, whereas no consistent differences in the animals were found. This unique egg characteristic therefore required the amendment of the Sisubiotus generic diagnosis to account for the presence of the labyrinthine layer inside the egg process walls.
821:1-15
A new woodlouse species of the genus Buchnerillo Verhoeff, 1942 is described and illustrated from the Cantabrian Coast of Asturias (Eastern Atlantic Ocean of the Iberian Peninsula). Buchnerillo atlanticus sp. nov. is a halophilic woodlouse that lives under embedded rocks in fine-grained sand areas of its type locality beach. Its morphological features, including secondary sexual characteristics, allow it to be distinguished from the other three known species of the genus Buchnerillo. Biological, ecological and ethological data of the new species are commented. To facilitate the separation of the four known species of Buchnerillo, the main diagnostic features are summarized and their known distribution is commented.
821:40-57
Acantholeberis smirnovi Paggi & Herrera-Martinez, 2020 (Cladocera: Acantholeberidae) was recently described from Andean habitats in South America. The presence of a population on the eastern coast of the continent brought the hypothesis of a new Neotropical species different from A. smirnovi. This hypothesis was confirmed from morphological, ecological and biogeographic evidence. Acantholeberis accolismaris Sousa, Elmoor-Loureiro & Álvarez-Silva sp. nov. differs from A. smirnovi in the morphology of the head, the valves and the limbs (especially the second and third limbs). Species of Acantholeberis are adapted to live in acid water bodies, with A. accolismaris sp. nov. presenting the same ecological requirements. However, the new species is adapted to live in temporary ponds near the Atlantic Ocean in Brazil in altitudes ranging between 4 and 15 meters above sea level. Considering that Acantholeberidae is an ancient taxon with a pre-Mesozoic origin, geological and environmental changes may have been relevant to speciation and to the observed biogeographic pattern of species of Acantholeberis in Neotropics.
819 :1-22
Acrostilicus Hubbard, 1896 and Pachystilicus Casey, 1905 are North American genera known from only one and two species, respectively, and have never been a subject of a modern revision. In fact, Acrostilicus was not even properly described as its author provided only a sketchy diagnosis of the genus and species. Here, we provide a redescription of the genus Acrostilicus and its species and illustrate the habitus and male genital features. For the first time, we also redescribe Pachystilicus and its two species, and provide their differential diagnoses. Additionally, we tested the phylogenetic position of both genera. They were scored into a morphological matrix supplemented with molecular data and the analyses were run using Bayesian inference and maximum likelihood methods. A total of 119 morphological characters and 4859 bp of nuclear (28S, TP, Wg, CADA, CADC, ArgK) and mitochondrial (COI) sequences were analysed for 46 taxa. The results confirmed that both Acrostilicus and Pachystilicus are members of the subtribe Stilicina, but at the same time challenged the monophyly of the subtribe in its current composition. Additionally, we provided further evidence for non-monophyly of the subtribe Medonina and discussed the biology of Acrostilicus and Pachystilicus.
807
The marine annelid family Sigalionidae is little known in the Grand Caribbean Region; there are few records of these worms in the area, and some of the recorded species have uncertain taxonomic status. In this contribution, the subfamily Pelogeniinae was addressed through a faunistic study, aiming to improve the knowledge of Sigalionidae in the region. In order to do this, material deposited in the three following institutions was examined: University of Miami Deep Sea Expeditions; Florida Museum of Natural History, University of Florida, Gainesville; and El Colegio de la Frontera Sur, Chetumal. Eleven species are recognized, including six newly described: Dayipsammolyce paulayi sp. nov., Hartmanipsammolyce pettiboneae sp. nov., Neopsammolyce fragilis sp. nov., Pelogenia brevipalpata sp. nov., P. capitata sp. nov. and P. salazarvallejoi sp. nov. Four other species are confirmed for this region: N. floccifera (Augener, 1906), P. hartmanae Pettibone, 1997, P. kinbergi (Hansen, 1882) and Psammolyce flava Kinberg, 1856; and one is indeterminate: N. aff. floccifera. A standardized terminology of neurochaetae is proposed, along with notes on the notochaetal morphology and elytral structures. For all genera covered in this study, identification keys are also provided.
806:177–186
A new species of the Neotropical genus Platygonia Melichar, 1925 is described and illustrated from the municipality of Ipixuna, State of Amazonas, Northern Brazil. Platygonia nigra sp. nov. can be distinguished from the other species of the genus by the following combination of features: (1) dark brown to black ground color of dorsum; (2) presence of a white to pale yellow spot at the distal portion of corium; (3) male pygofer with a conspicuous diagonal cleft; (4) connective Y-shaped, keeled, with the stem longer than the arms; and (5) aedeagus with an unpaired basiventral process directed anteriorly. This is the first record of the genus from the Brazilian Amazon Rainforest. Notes on P. undecimmaculata (Fowler, 1899), which is a taxon of uncertain taxonomic position, a key to the species of Platygonia, and a map showing their distribution are added.
797
Descriptions of the following 23 species of Macrotomoderus Pic, 1901 new to science, from continental China, are provided as an addition to the recently published review of the genus from China and Taiwan (Telnov 2018): M. angelinii, M. belousovi, M. bicrispus, M. boops, M. bordonii, M. dali, M. daxiangling, M. femoridens, M. hajeki, M. hartmanni, M. hengduan, M. imitator, M. kabaki, M. korolevi, M. lapidarius, M. muli, M. palaung, M. similis, M. tenuis, M. transitans, M. truncatulus, M. usitatus, and M. wudu spp. nov. Additional records are provided for some poorly known species. The identification key to the species of Macrotomoderus from China, the Japanese Archipelago, and Taiwan is herewith significantly supplemented and updated. Biogeographical peculiarities and altitudinal gradient of Macrotomoderus distribution in continental China are briefly discussed.
789:11-48
The diversity of Palpigradi is not evenly distributed among its different branches. The widespread genus Eukoenenia includes 80% of the species, while the other genera are poorly known. Allokoenenia certainly is the most understudied genus because it is represented only by the African species Allokoenenia afra Silvestri, 1913. Its description is short and does not include many features depicted in modern taxonomy of Palpigradi. In this paper, we describe two troglobitic species of Allokoenenia, report the occurrence of a third species represented by an immature specimen from Brazilian caves, and provide brief notes on the morphology of A. afra. Allokoenenia canhembora sp. nov., A. stygia sp. nov., and Allokoenenia sp. differ from A. afra by several morphological features, including more elongated appendages and a greater number of blades on lateral organs. Thus, they are considered troglomorphic. These new species are vulnerable to extinction because they are endemic to a single or few caves directly impacted by mining activities and groundwater exploitation. This study represents the first step for the conservation of these species and their habitats, since Brazilian caves with rare troglobites cannot be irreversibly impacted. Also, it brings important contributions on the distribution and morphology of this enigmatic genus.
789:104-129
Early Carboniferous coiled nautiloids from North Africa are virtually unknown. An assemblage of nine species, all from the family Trigonoceratidae, from the Dalle à Merocanites (Tournaisian-Viséan boundary interval) of Timimoun in western Algeria is described, being the most diverse Carboniferous nautiloid assemblage known from North Africa but much less diverse than the time-equivalent assemblages from Belgium and Ireland. The assemblage consists of the species Maccoyoceras pentagonum sp. nov., Lispoceras orbis sp. nov., Thrincoceras devolvere sp. nov., Rineceras multituberculatum sp. nov., Rineceras rectangulatum sp. nov., Vestinautilus padus sp. nov., Vestinautilus concinnus sp. nov., Planetoceras destrictum sp. nov. and Planetoceras transforme sp. nov. A morphometric analysis of Maccoyoceras pentagonum sp. nov. and Lispoceras orbis sp. nov. shows that the intraspecific variation in these species is within rather narrow limits.
782:21-54
The unique Neotropical species of the alderfly genus Protosialis van der Weele, 1909, P. bifasciata (Hagen, 1861), is herein transferred to the newly described genus Caribesialis gen. nov. This new taxon is proposed to be sister to the clade Protosialis + Sialis Latreille, 1802, after a phylogenetic analysis that included male genital characters scored on a previous morphological matrix of the family. Also, Ilyobius nigrocephalus sp. nov., a remarkable new species from Ecuador, is described and its phylogenetic position is discussed. Furthermore, Ilyobius bimaculatus (Banks, 1920) from Bolivia, known solely from the female holotype, is redescribed. Based on the present study, the Neotropical fauna of Sialidae is proposed to be constituted by two genera, one insular (Cuba) and one continental (Mexico to Chile and Argentina).
775:15-33
Two novel species of Russula (Russulaceae, Russulales), R. coronaspora Y.Song sp. nov. and R. minor Y.Song sp. nov. belonging to subgenus Russula crown clade, are described based on both morphological and phylogenetic evidence. In morphology, R. coronaspora sp. nov. is mainly characterized by its distinct spores ornamented with sparse, cylindrical and isolated spines, which resemble coronavirus, and gelatinized pileipellis with pileocystidia mostly septate and sometimes branched; R. minor sp. nov. possesses a very small basidiocarp with pileus less than 2.5 cm in diameter., small basidia, easily peeling and gelatinized pileipellis with slender terminal cells and abundant SV+ pileocystidia. Positions of the two new species in both phylogenetic trees based on ITS and 5-locus sequences (nLSU, mtSSU, rpb1, rpb2 and tef1) confirm their distinct taxonomic status.
773:80-119
Simulium (Trichodagmia) Enderlein, 1934 has an unstable classification system. The broader concept of the subgenus includes five species-groups resulting from an extensive history of synonymisations, often outside of a phylogenetic framework. This concept also ignores relationship hypotheses with the Afrotropical subgenera S. (Anasolen) Enderlein, 1930, S. (Freemanellum) Crosskey, 1969, S. (Xenosimulium) Crosskey, 1969, and the Oviedoi species-group, with several Neotropical species of S. (Trichodagmia). We performed a morphological phylogenetic analysis to test the monophyly of S. (Trichodagmia), its species-groups, and their relationship with the above-mentioned subgenera and Oviedoi species-group. We analysed a data matrix with 69 terminal taxa and 62 characters under parsimony implied weights, with a range of concavities (k1–100), finding three categories of k. Our analysis concludes that S. (Trichodagmia) is not monophyletic, since Oviedoi and the Afrotropical subgenera group with its species-groups Tarsatum and Orbitale. Therefore, we propose a new classification for S. (Trichodagmia) by restricting it to the Orbitale species-group, revalidating S. (Hearlea) Vargas et al., 1946, S. (Hemicnetha) Enderlein, 1934, S. (Obuchovia) Rubtsov, 1947 and S. (Shewellomyia) Peterson, 1975, synonymysing S. (Xenosimulium) with S. (Anasolen), and erecting a new subgenus, S. (Disculter) subgen. nov. for Oviedoi. The geographical distribution of the groups involved is discussed.
767:94–116
The genus Afronurus has several very common mayfly species in China and they are widely distributed in this country. Some of them are quite similar to each other in both imaginal and nymphal stages. However, these species have not been systematically compared and reviewed so far. In this study, six species are recognized. All nymphs of them share the following characters: gills V–VI with additional arrow-like accessory lobes, branched dentisetae, two rows of bristles and setae on hindtibiae and spotted abdominal terga. The males have divergent penes and clearly expressed titillators. The nymphs of the new species A. drepanophyllus sp. nov. have sickle-like gills I, spotted and striped body color, and males have unique genitalia. The nymphal stages of A. furcatus and A. hunanensis, which are associated and described for the first time, have similar body color to A. drepanophyllus sp. nov., but their pale dots on the head capsules and the shape of the hypopharynx are different. Keys to males and nymphs of the six species are provided.
767:83–93
Two new species of the genus Eponisiella Emeljanov, 1984 are described and illustrated from China (Guizhou and Shandong Provinces). These are E. dafangensis sp. nov. and E. shandongensis sp. nov., giving the genus eight species in total. A key to the species of Eponisiella is provided as well as a map of their geographic distribution, which is briefly discussed.
766
Ophiuroidea is the largest class among extant echinoderms, with over 2000 described species assigned to 33 families. Here, the first identification key to the recently revised classification was developed, and revised morphological descriptions were derived from it, expanding the previous short diagnoses. The key was built by analyzing internal and external skeletal characters of predominantly the type species of each family, including at least two mutually exclusive attributes per family. Various numeric and multistate characters were used to create a traditional as well as an interactive key using the DELTA and Xper software programs. Illustrations (SEM and digital photos) are included in the key to facilitate the assessment of character states by users. Not only is it the first identification key to the families, according to the recently proposed new classification and the examined species, but this interactive key also assists users in understanding the family level taxonomy of brittle stars. The interactive key allows new characters and states to be added, when more species will have been analyzed, without the need to reconfigure the complete key (as may be necessary with conventional keys).
764:85-118
The genus Salvadora has not been subjected to a modern phylogenetic analysis. Described in 1853, its taxonomic history is complex and confusing. In this study, we evaluate the monophyly of the genus and present the first phylogenetic hypothesis based on an analysis of 66 qualitative and quantitative morphological characters of over 1000 specimens representing all described taxa across their entire distribution. Morphological characters were processed in Fast Morphology for subsequent phylogenetic analysis in PAUP under the maximum parsimony criterion. We obtained a single tree in which Salvadora appears as a monophyletic group with two clearly defined geographic species groups: a southern mexicana group and a northern grahamiae group. Based on our phylogenetic hypothesis, we evaluate the taxonomic status of all described taxa. Additionally, we include a diagnosis for all species, distribution maps, and an illustrated dichotomous taxonomic key of the genus.
764:18-36
The genus Trypheridium Brancucci, 1985 is endemic to the Hindu Kush Himalayan Region, and is currently known from a single species, T. nuristanicum (Wittmer, 1956). Here, the genus is reviewed, T. nuristanicum nom. emend. is re-described and T. kashmiricum sp. nov. is described from Kashmir Himalayan Region of India. Descriptions, diagnoses, high quality images, distribution maps and identification keys are presented. The morphology and distribution of Trypheridium are discussed and compared with those of the closely related genus Trypherus LeConte, 1851.
762:49-95
Five species of the subgenus Clessiniella of Cochlostoma (Cochlostomatidae) are recognized, viz. Cochlostoma (Clessiniella) villae (Strobel, 1851), Cochlostoma (Clessiniella) tergestinum (Westerlund, 1878), Cochlostoma (Clessiniella) waldemari (A.J. Wagner, 1897), Cochlostoma (Clessiniella) anomphale Boeckel, 1939 and Cochlostoma (Clessiniella) stelucarum sp. nov. The shells and the genitalia are described and the distributional data are summarized for all the species.
760:160-185
A new species of abyssal Neanthes Kinberg, 1865, N. goodayi sp. nov., is described from the Clarion-Clipperton Zone in the central Pacific Ocean, a region targeted for seabed mineral exploration for polymetallic nodules. It is a relatively large animal found living inside polymetallic nodules and in xenophyophores (giant Foraminifera) growing on nodules, highlighting the importance of the mineral resource itself as a distinct microhabitat. Neanthes goodayi sp. nov. can be distinguished from its congeners primarily by its distinctive, enlarged anterior pair of eyes in addition to characters of the head, pharynx and parapodia. Widespread, abundant, and easily recognisable, N. goodayi sp. nov. is also considered to be a suitable candidate as a potential indicator taxon for future monitoring of the impacts of seabed mining.
760:32-60
In the Pacific Ocean, the taxonomy of the family Zosimeidae Seifried, 2003 is poorly understood and to date only five species of the genus Zosime Boeck, 1873 are known. During oceanographic cruises exploring the species diversity of harpacticoids, two undescribed zosimeid copepods were sampled from shallow Korean waters and the deep northwestern Pacific. A detailed morphological examination has led us to propose two new genera, Heterozosime gen. nov. for the Korean zosimeid H. tenuis gen. et sp. nov. and Acritozosime gen. nov. for the deep-sea zosimeid A. spinesco gen. et sp. nov. Both new genera exhibit a distinctive feature in that the first thoracic leg has a two-segmented exopod, in contrast to the three-segmented exopod of this leg in all known zosimeid genera. Furthermore, Acritozosime gen. nov. can also be discriminated from other genera by the two-segmented endopod in second to fourth thoracic legs and the reduced setal armatures of the second exopodal segment of antenna, the first endopodal segment of first to third thoracic legs and the third exopodal segment in second to fourth thoracic legs. A comparison of the fundamental structures of appendages suggests that A. spinesco gen. et sp. nov. experienced a unique evolutionary history within the Zosimeidae.
760:101-135
The present study redescribes four species of Neanthes Kinberg, 1865 (Nereididae de Blainville, 1818) based on their type specimens collected from different worldwide localities: Neanthes chilkaensis (Southern, 1921) from India, N. galetae (Fauchald, 1977) from Panama, N. helenae (Kinberg, 1865) from St Helena Island, and N. mossambica (Day, 1957) from Mozambique. The morphology of the types was re-examined for the first time after the species were originally described, and incorporated the recent improvements in the standards and terminology for describing nereidid features. The arrangement of paragnaths on area VI stood out among the diagnostic features used to distinguish these four species. Neanthes chilkaensis and N. helenae are the unique nereidids bearing p-bar paragnaths on the area VI. Both species are also distinctive as the former species only exhibited p-bar paragnaths on the area VII–VIII and the latter ventrolateral projections on the apodous segment. Further examination revealed that N. nanciae (Day, 1949) from St Helena is a junior synonym of N. helenae. Moreover, N. galetae and N. mossambica are distinguishable from other species also by the development of dorsal cirri, neuropodial postchaetal lobe and ventral ligule, the presence/absence of merged paragnaths on area IV, paired oesophageal caeca, among other features. This study has further contributed to the morphological delimitation of the species in Neanthes as a first step towards revising the genus.
751:38-67
A morphological and molecular review of the genus Goniurosaurus, including an identification key
(2021)
The genus Goniurosaurus (tiger geckos) currently consists of 23 species distributed in China, Japan and Vietnam. Several species complexes and recent discoveries of cryptic species pose challenges to the species identification, which is crucial to effectively implement the recent listing of the species from China and Vietnam in CITES Appendix II and the species from Japan in CITES Appendix III. Based on the results of our field work in northern Vietnam and data compiled from literature, we herein provide a taxonomic review of the genus Goniurosaurus. Our phylogenetic analyses showed that all recorded populations of tiger geckos from Vietnam, which were found to be monophyletic with low intra-specific genetic divergences, are assigned to one of the four species: G. catbaensis, G. huuliensis, G. lichtenfelderi or G. luii. Both genetic and morphological analyses confirm that the species from China and Vietnam can be split into three major groups. Based on the newly collected data, we provide an extended morphological description of the Vietnamese species. In addition, we provide an identification key for all Goniurosaurus species from China, Japan and Vietnam in order to assist authorities in the enforcement of the recent CITES listing.
750:140-155
Most valvatiform genera of the gastropod family Hydrobiidae are narrow-range taxa. One exception is the genus Arganiella, which is comprised of three congeners: the type species A. pescei from the Apennine Peninsula, A. wolfi from the Iberian Peninsula and A. tabanensis from the Balkans. The genus assignment of the latter two species was based on morphological similarities with A. pescei in the shell, operculum, radula and genitalia. Given that the morphology of hydrobiids is sometimes susceptible to convergence, this study re-evaluates the taxonomic status of species of Arganiella by analysing mitochondrial (mtCOI) and nuclear (18S rRNA) sequences of topotypes or near topotypes to infer their phylogenetic position. Our phylogenetic analyses depicted Arganiella as a non-monophyletic group within Hydrobiidae, and sequence divergence among the three species ranged from 14.5 to 16.7% for mtCOI and 2.0 to 3.8% for 18S. We also re-examined the extent of morphological variation among species of Arganiella and found a few differences among them and other valvatiform genera. Consequently, we propose two new genera for A. wolfi and A. tabanensis. Our results conflict with the classification of valvatiform hydrobiid species solely based on traditional phenotypical methods and suggest further taxonomic evaluation within a molecular framework.
750:156-196
The Smicridea (Smicridea) fasciatella species group occurs from the southwestern USA, throughout Central America, the Greater Antilles islands, and most of South America, except for the Chilean subregion. It is characterized by the phallic apparatus being a simple tube with eversible internal sclerites at the apex. The fasciatella group is composed of 61 species, of which only 11 occur in Brazil, mainly in the Atlantic Forest biome in the southeastern region. In order to reduce the Linnean and Wallacean shortfalls for the Smicridea Brazilian fauna, we diagnose, describe, and illustrate males of six new species in the fasciatella group: Smicridea (Smicridea) blahniki Desiderio, Pes & Hamada sp. nov., S. (Smicridea) brevitruncata Desiderio, Pes & Hamada sp. nov., S. (Smicridea) caaguara Desiderio, Pes & Hamada sp. nov., S. (Smicridea) ipiranga Desiderio, Pes & Hamada sp. nov., S. (Smicridea) jeaneae Desiderio, Pes & Hamada sp. nov., and S. (Smicridea) polyacantha Desiderio, Pes &; Hamada sp. nov. Additionally, we provide distributional data for S. (Smicridea) albosignata Ulmer, 1907, S. (Smicridea) bivittata (Hagen, 1861), S. (Smicridea) erecta Flint, 1974, S. (Smicridea) obliqua Flint, 1974, S. (Smicridea) paranensis Flint, 1983, and S. (Smicridea) sattleri Denning & Sykora, 1968. The number of S. (Smicridea) species in Brazil increases from 21 to 27 and Smicridea is recorded from the states of Acre, Amapá, and Sergipe for the first time.
749
The genus Miridiba Reitter, 1902, of phytophagous chafers from Asia, is revised based on external morphological and genital (male and female) characters. In this study, a total fifty-eight species of Miridiba were examined, and the genus is redescribed. Male genital characters of Miridiba, especially the morphology of parameres and endophallus, are studied in depth herein. The female genitalia of Miridiba are studied and described in detail for the first time. According to genital characters, nine genital morphotypes, including fifty-four species, are established under the genus Miridiba. Sixty-seven type specimens are studied. Miridiba gressitti (Frey, 1970) comb. nov., Miridiba borneensis (Moser, 1918) comb. nov., Miridiba coxalis (Arrow, 1944) comb. nov., Miridiba rugaticollis (Moser, 1913) comb. nov., Miridiba nigrescens (Moser, 1916) comb. nov., Miridiba scutata (Reitter, 1902) comb. nov., Miridiba ciliatipennis (Moser, 1903) comb. nov. and Miridiba brunneipennis (Moser, 1916) comb. nov. are transferred from Holotrichia. Holotrichia dalatensis Frey, 1970 is transferred to Miridiba as a synonym of Miridiba sinensis (Hope, 1842). Four junior subjective synonyms are proposed: Miridiba pilosella (Moser, 1908) (= Holotrichia formosana Moser, 1909), Miridiba sinensis (= Holotrichia dalatensis Frey, 1970), Miridiba scutata (= Holotrichia scutulata Dalla Torre, 1912 and Miridiba lassallei Keith, 2010). Miridiba frontalis (Fairmaire, 1886) is resurrected as a valid species. A key to 51 of the 58 valid species of Miridiba is presented.
748:36-50
A new avian chewing louse genus Apomyrsidea gen. nov. is described based on species parasitizing birds in the family Formicariidae. Diagnostic characteristics and phylogenetic analyses were used to evaluate and confirm the generic status and merit its recognition as unique and different from Myrsidea Waterston, 1915. Three species previously belonging to the genus Myrsidea are placed in the new genus Apomyrsidea gen. nov. and are discussed: Apomyrsidea circumsternata (Valim & Weckstein, 2013) gen. et comb. nov., Apomyrsidea isacantha (Valim & Weckstein, 2013) gen. et comb. nov. and Apomyrsidea klimesi (Sychra in Sychra et al., 2006) gen. et comb. nov.
748:1-14
746:112-129
Comanopa Blocker, 1979 and Gehundra Blocker, 1976 are small and poorly known genera of leafhoppers, previously comprising four and five species, respectively. In this study, two new species are proposed: Comanopa mananciensis sp. nov. from the state of Paraná, Southern Brazil, and Gehundra cristalinensis sp. nov. from the state of Mato Grosso, Centralwestern Brazil. The male of Gehundra sordida (Baker, 1900) is studied by the first time. Detailed descriptions and illustrations of males and females are provided and keys to males of Comanopa and Gehundra are given.
746:50-61
A new scale insect genus and species, Gompholopium quercicola gen. et sp. nov. is described and illustrated, based on material from China (Yunnan Prov.). Hemaspidoproctus cinereus (Green, 1922) is redescribed, based on material from India (Allahabad). Walkeriana tosariensis Reyne, 1957 is transferred to the genus Hemaspidoproctus Morrison, 1927 – H. tosariensis (Reyne, 1957) comb. nov. Monophlebus neglectus Gavrilov-Zimin, 2018 syn. nov. is formally synonymized with the unrecognizable Monophlebus atripennis Burmeister, 1835 in order to resolve an old taxonomic and nomenclatural problem in the subfamily Monophlebinae.
739:158-167
A new species of verrucid barnacle, Gibbosaverruca weijiai sp. nov., is described and illustrated based on specimens collected from a deep-sea seamount, Weijia Guyot, in the West Pacific. This is the third barnacle species reported from Weijia Guyot. The new species is distinguished from its congeners by its extremely long caudal appendage. In terms of shell morphology, G. weijiai sp. nov. is most similar to G. gibbosa and G. sulcata but differs from them by the intermediate articular ridges of its movable tergum and scutum, which are distinctly wider than the axial ridges. The COI and 16S rRNA sequences of the holotype of G. weijiai sp. nov. are presented to support future research, and a key to extant species of the genus Gibbosaverruca Young, 2002 is provided.
733:125-145
Thanks to newly collected material from the Terra Nova Bay area (Ross Sea, Antarctica), we discuss the taxonomy of the ampharetid genera Amage Malmgren, 1866 and Amythas Benham, 1921. A new species of Amage, A. giacomobovei sp. nov., is described based on morpho-anatomical data. This is the second new species described from an area which appears to be rich in ampharetids, a coastal embayment at ~500 m depth near the Italian “Mario Zucchelli” research station. The new species is characterized by having 16 abdominal uncinigers and four pairs of branchiae that readily distinguish it from its congeners. Tubes of A. giacomobovei sp. nov. are also characteristic in showing a large amount of embedded sponge spicules, suggesting a possible close association to spicule mats. Based on the amended diagnoses of the two genera, Amage septemdecima Schüller & Jirkov, 2013 is transferred to the genus Amythas. Finally, to simplify the task of ampharetid genera recognition for untrained people, we provide a dichotomic key for ampharetid genera found in Antarctica and a checklist of species occurring in Terra Nova Bay.
724:1-33
On eight species of the spider genus Synagelides Strand, 1906 from China (Araneae: Salticidae)
(2020)
Six new species of Synagelides Strand, 1906 are diagnosed and described: S. bohdanowiczi sp. nov. (♂♀), S. leigongensis sp. nov. (♂♀), S. logunovi sp. nov. (♂♀), S. subgambosus sp. nov. (♂♀), S. wuliangensis sp. nov. (♂♀) and S. xingdouensis sp. nov. (♂♀). The female of S. forkiforma Yang, Zhu & Song, 2007 and the male of S. longus Song & Chai, 1992 are described for the first time. Photos of the habitus and copulatory organs, as well as a distributional map, are provided.
724:93-108
The genus Parasogata Zhou, Yang & Chen, 2018 is here reported from India represented by the new species Parasogata sexpartita sp. nov. collected in a recent exploration and survey of delphacids from Nagaland in northeastern India. A second species of Eoeurysa Muir, 1913 from India, the new species Eoeurysa sagittaria sp. nov., was found in Rampur, Una, Himachal Pradesh. Both new species are described with illustrations, and a molecular identification is given with the mtCOI gene sequence. A modified key to species of the genera is also provided.