Tuexenia : Mitteilungen der Floristisch-Soziologischen Arbeitsgemeinschaft, Band 33 (2013)
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- Festuco-Brometea (3)
- eutrophication (3)
- vegetation classification (3)
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- Caricion davallianae (2)
- Central Europe (2)
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- Violion caninae (2)
Der diesjährige 8. Trockenrasen-Sonderteil von Tuexenia beginnt mit einen Bericht über die aktuellen Aktivitäten der European Dry Grassland Group (EDGG). Zunächst geben wir einen Überblick über die Entwicklung der Mitgliederzahl und den aktuellen Vorstand, der im Mai 2013 gewählt wurde. Dann berichten wir vom letzten European Dry Grassland Meeting in Prespa (Griechenland, 2012) und informieren über künftige Tagungen und Forschungsexpeditionen der EDGG. Schließlich war und ist die EDGG sehr aktiv darin, Special Features in internationalen Fachzeitschriften herauszugeben. Im zweiten Teil des Vorwortes geben wir eine Einführung zu den sechs Artikeln des diesjährigen Trockenrasen-Sonderteils: Zwei davon beschäftigen sich Biodiversitätsanalyen von Grasland-Ökotonen in einer Flusslandschaft in Lettland bzw. von brachgefallenen Alvar-Trockenrasen in Estland. Der dritte Artikel behandelt die Ökologie und Vergesellschaftung einer Grassippe (Avenula adsurgens subsp. adsurgens) im brachgefallenen, montanen Grasland der Karpaten (Slowakei). Die letzten drei Artikel schließlich sind der Beschreibung und Syntaxonomie von Trockenrasen gewidmet: Zwei davon bilden den Start einer neuen Serie über die pannonischen Trockenrasen Österreichs (Allgemeine Einführung und Trockenrasen des Wienerwalds), während der letzte die Ergebnisse der dritten EDGG-Forschungsexpedition im Jahr 2011 nach Bulgarien präsentiert. Schließlich geben wir einen Ausblick über künftige Pläne für das Special Feature.
Plant communities and environmental gradients in mires of the Ammergauer Alps (Bavaria, Germany)
(2013)
In this study we present the plant communities, their abiotic site conditions and the main environmental gradients underlying variation in species composition of mire sites in three montane mires in the Northern Limestone Alps. For the phytosociological description of the mires we sampled species composition in 115 relevés. In order to characterize the site conditions of the most important plant communities we measured humification as well as water table depth, pH and electrical conductivity at 46 sites during a large part of the growing season. The main environmental gradients were identified by fitting environmental vectors and smoothed surfaces onto NMDS ordination. The most frequent communities on subneutral and neutral sites were the Caricetum davallianae, the Amblystegio stellati-Caricetum dioicae and the Schoenetum ferruginei, whereas acidic sites were mainly covered by the Eriophoro-Trichophoretum cespitosi. The most important factor for floristic variation between sites was pH. However, within bogs and fens, variation in species composition was mainly due to differences in water table depth.
Auf dem Plateau des Göttinger Waldes wurden 1980 12 ha eines artenreichen, heute etwa 145 Jahre alten, submontanen Kalkbuchenwaldes (Hordelymo-Fagetum lathyretosum) für ein Ökosystemforschungsprojekt eingezäunt. In diesem Bereich wurde ein großer Transekt (GT) von 2,81 ha mit 281 10x10m-Quadraten als Dauerfläche zur Untersuchung der natürlichen Vegetationsentwicklung ausge-wählt. Von 1981 bis 2011 wurden alle 10 Jahre Flora und Vegetation sehr detailliert in allen Quadraten erfasst (Schichtung, Deckungsgradschätzung aller Arten in %, Vegetationskartierung). Die Ergebnisse werden in Tabellen der Krautschicht, in quantitativen Verteilungskarten einzelner Arten und in Vegetationskarten dargestellt. – Schon in den ersten 10 Jahren hatte sich teilweise eine Strauchschicht, vor-wiegend aus jungen Bäumen, entwickelt. Auch in der Krautschicht gab es deutliche Veränderungen. Eine Frequenztabelle aller 83 gefundenen Arten (Tab. 1) zeigt zahlreiche Pflanzen (33) mit Abnahme-tendenz, dazu einen Grundstock konstanter Arten. Deutlich zugenommen haben nur Allium ursinum, Cardamine bulbifera, Dryopteris carthusiana, Hedera helix und Neottia nidusavis. Für die Vegetationskartierung wurden verschiedene Einheiten nach Dominanz oder Mischung einzelner Arten benutzt, mit den Schlüsselarten Aconitum lycoctonum, Allium ursinum, Anemone nemorosa und Mercurialis perennis. – Insgesamt war lange Zeit der Antagonismus von Allium (deutliche Zunahme) und Mer-curialis (starke Abnahme) besonders auffällig. So hat sich auch der Allium ursinum-Dominanztyp über 30 Jahre stark ausgedehnt. Während sich kleinflächig in den Quadraten (Mikroskala) deutliche Veränderungen der Artenzusammensetzung zeigten, war im gesamten Bestand (Mesoskala) teilweise auch floristische Konstanz zu erkennen. – In der Diskussion werden mögliche Ursachen für die festgestellten Veränderungen erörtert. Neben lokalen Wirkungen wie Einzäunung oder der Konkurrenzkraft von Allium ursinum lassen sich im Literaturvergleich großräumig wirksame Faktoren erkennen. Seit langem vollzieht sich vor allem in forstlich wenig oder gar nicht beeinflussten Laubwäldern eine Verdichtung des Kronendaches mit Ausbildung eines stärker schattig-luftfeuchten Mikroklimas. Seit einigen Jahrzehnten können stärkere Stickstoffeinträge für nährstoffliebende Arten wirksam sein. In den letzten 20 Jahren lassen sich zunehmend Auswirkungen einer Klimaerwärmung erkennen, z. B. eine Verlängerung der Vegetationsperiode. Als neues Phänomen wird das Eschentriebsterben durch Pilzbefall beschrieben.
Within the last 30 years the role of nitrogen in Central European forests has changed fundamentally from limiting resource to environmental problem. As the retrospective tracking of nutrient availability by soil chemical and biogeochemical measurements faces serious problems, bioindication based on understorey species composition is indispensable for monitoring broad-scale eutrophication. Based on a broad survey of more than 100,000 forest vegetation plots accessible in electronic data-bases from Germany and adjacent countries, we calculated unweighted average Ellenberg nutrient values (mN) as a proxy of plant-available macronutrients. Based on the quantiles of the frequency distribution of mN in a regionally stratified sample, we define five trophic classes, which can be used to compare dimensionless mN values. We studied spatial patterns of average nutrient values within 17 regions and compared the periods from 1899 to 1975 and 1976 to 2006. After 1975 eutrophic (mN > 5.67) and hypertrophic (mN > 6.28) conditions were common everywhere except in the Alps and Saxony-Anhalt, but very oligotrophic conditions (mN < 3.44) were still widespread in regions with nutrient-poor bedrock. Before 1975 mN of plots had been lower than after 1975 in all but the southeastern regions. Between the pre- and post-1975 data the proportion of hypertrophic plots increased from 5.7 to 11.8%, and that of very oligo-trophic plots decreased from 14.6 to 8.3%. To remove bias resulting from uneven distribution, the dataset was stratified by five tree layer dominance types, period and region and resampled. In pre-1975 plots medians of mN increased in the order Pinus sylvestris, Quercus spp., Picea abies, Fagus sylvatica and Alnus spp, whereas the increase of mN was highest in forest types with historically low nutrient values. Therefore, the widespread change in mN must be attributed to the pronounced vegetation changes in Quercus and Pinus stands, indicating the importance of land-use change, i.e. recovery of nutrient cycles after hundreds of years of exploitation through coppicing, grazing and litter use. The analysis confirms eutrophication as a megatrend of modern vegetation change and demonstrates the high research potential of linking vegetation plot databases across large regions.
In Kiefernbeständen des Naturschutzgebietes Mallertshofer Holz wurden, stratifiziert nach Besto-ckungstypen, Vegetationsaufnahmen angefertigt, klassifiziert und mittels Ordination und Zeiger-wertanalyse standörtlich und dynamisch interpretiert. Bei homogenen primären Standortbedingungen folgt die Vegetation einem starken Nährstoffgradienten, bedingt durch unterschiedliche extensive Vornutzungen, Selbstmelioration und Stickstoffeintrag. Für das Management der Wälder ergeben sich daraus drei Optionen: 1. Fortsetzung der selbstgesteuerten Entwicklung eutropher Kiefernforste; 2. aktiver Waldumbau durch Einbringen von Schattbaumarten der potenziellen natürlichen Vegetation; 3. gezielte Auflichtung und Ausmagerung durch starke Eingriffe in Gehölzbestand (Ganzbaumernte) und Bodenvegetation (Beweidung). Der Naturschutzwert des Gebietes kann durch ein Nebeneinander der Varianten 2 und 3 gesichert und optimiert werden.
Summed frequency data of apomictic taxa derived from vegetation relevées from Central Europe were correlated with ecological traits – water content of soil, calcium and magnesium content of soil or water, nutrient content of soil or water, environmental dynamics, hemeroby, frequency of stress-tolerant species, salt tolerance, altitude, height, and frequency of annual taxa – of alliances of the Central European vegetation. The partly problematic identification of taxa where apomixis is important for seed production was based on a literature review. In Central Europe, high altitudes favour apomictic taxa. Otherwise, no correlations between ecological factors attributed to the alliances of the European phytosociological system and frequencies of apomictic taxa in the alliances of the European phytosociological system were found. Analyses could not confirm theories on ecological preferences of apomictic taxa; e.g., that competition penalizes apomicts, apomicts have better colonizing abilities than sexual plants, or apomicts have larger ranges than their sexual relatives could not be confirmed.
Only a small proportion of introduced plant species become invasive and may eventually create ecological or economic problems. In many species it is still not clear which traits cause biological inva-sions. As a case study we focussed on the fast-spreading Epilobium brachycarpum in Central Europe to investigate the potential of this species to become a transformer or agricultural weed. We (1) documented the spread of the species in Central Europe, (2) modelled its range and (3) seed dispersal, (4) described its phytosociological alignment, (5) analysed the traits of invaded vegetation types, (6) described seed production, population densities and life cycle, (7) did competition and germination tests, and (8) drafted a risk assessment. Relevant traits and characteristics of E. brachycarpum are (i) formation of dense stands under ruderal conditions, (ii) high seed production, (iii) effective seed dispersal, (iv) high competitiveness on bare soils against other ruderal plants, and (v) ecological niche shift com-pared to its native range. We expect E. brachycarpum to settle in the Mediterranean, sub-Mediterranean and many parts of temperate Europe within the next decades in habitats strongly altered by human activities, especially open stands of the alliance Sisymbrion. We predict that E. brachycarpum will become a noxious weed in vineyards, and that it will also colonise vegetation of the alliances Bidention and Carici-Epilobion.
Evidence for increasing homogenization and de-ruralization of the Central European village flora
(2013)
In this study, the spontaneous flora of North Rhine-Westphalian villages (NW Germany) recorded in the periods 1980–1984 and 2004–2005 was analyzed. We asked: (i) Did the similarity of the village flora increase with time and to what extent did non-native species contribute to this? (ii) Is the ongoing trend of urbanization reflected in the village flora? (iii) Regarding the species composition and β-diversity: are there differences between the results of a repeated survey and of a type comparison? In both periods, 200 villages were visited once, using consistent criteria for village selection and floristic investigation. Of these, a subset of 61 villages was investigated in both time periods and the rest were selected randomly for the second survey but with respect to the same criteria as used in the first. Using different β-diversity indices (Sørensen index of dissimilarity, Simpson index of dissimilarity and nestedness resultant index) to disentangle pure species turnover and nestedness of species assemblages, we tested whether the similarity among the villages increased over the years. This was done by calculating all pairwise comparisons among the villages of each time period, i.e. their spatial turnover. Additionally, different ecological groups were defined (e.g. "neophytes", "C-strategists" or "urbanophilous species") using the subgroups of species traits (floristic status, Grime's life strategy and urbanity) to indentify urbanization and compositional changes. Our results show that the traditional village flora (species of the Arction lappae, Bidention tripartitae and Potentillion anserinae) is further blurred towards unspecific ubiquitous assemblages found in the periphery of man and that the floristic urbanization is still ongoing. Supported by nearly all ecological groups regarded, our results corroborate that the species composition of Central European villages underwent a considerable degree of homogenization. To some extent this is caused by structural homogenization of villages. Regarding the considerable increase of common ubiquists and α-diversity in general, however, changes in weeding practices seem to be the main reason. Based on the reinvestigated subset we could show that comparable results can be obtained by a consistent use of criteria for the selection of villages, even if unpaired samples were used.
This paper focuses on ecological requirements and phytosociological affinity of Avenula adsurgens subsp. adsurgens. Although this grass is widely distributed in central and south-eastern Europe reaching dominance in certain grassland types, the knowledge on its ecology and coenology is very poor. Moreover, some of the published data on its distribution are wrongly related to Avenula praeusta. We studied the taxon within an area of about 300 km2 (Central Slovakia) where it occurs in diverse habitats. Data from a systematic phytosociological survey were used to assess interspecific associations and ecological indicator values of the taxon. Detailed measurements from a transect along a spruce colonisation gradient were used to evaluate its relationship to a set of topographical, microclimatical, pedological and soil-microbiological characteristics. Tillers of A. adsurgens subsp. adsurgens were cultivated for two growing seasons to estimate characteristics of its clonal morphology and growth and its ability of spatial spreading. In the studied area, the taxon occurred mainly over the volcanic bedrock along a wide range of altitudes. It was concentrated in slightly managed or abandoned montane grasslands (800–1100 m) of the Violion caninae and Nardo strictae-Agrostion tenuis alliances. Phytosociologically the taxon seems not to be an important diagnostic species of these communities, rather an indicator of their successional development. Its highest cover was recorded in early to mid-successional stages without significant proportions of woody species. In stands with traditional management by mowing, grazing or their combination, the taxon was common (albeit with lower percentage cover), but more intensive ma-nagement including amelioration or fencing led to its retreat. Ellenberg ecological indicator values of A. adsurgens subsp. adsurgens were set as follows: light 7, temperature 5, continentality 4, moisture 4, soil reaction 5 and nutrients 3. Along a successional gradient, the taxon cover was positively correlated with percentage cover of herb litter, catalase activity, canopy openness and potassium content and negatively correlated with percentage cover of needles and canopy index reflecting the cumulative effects of spruce colonisation. The horizontal spreading of the taxon was enabled by the formation of spacers with a maximum length of 25.5 cm. We suppose that the ability to spread horizontally by spacers, together with the observed clonal plasticity, tall stature and fast growth by intensive tillering, belong to important attributes of A. adsurgens subsp. adsurgens promoting its success and dominant role in the abandoned grasslands.
The article deals with Norway spruce (Picea abies), silver fir (Abies alba) and larch (Larix decidua) forests of the orders Piceetalia and Athyrio-Piceetalia (Vaccinio-Piceetea) in Slovenia. A total of 934 relevés of spruce, larch and fir forests within the Piceetalia and Athyrio-Piceetalia were collected from the literature. After resampling, 319 relevés remained, originating from 28 associations. Classification revealed two major groups of spruce, larch and fir forests, one thriving on non-carbonate bedrock and the other on carbonate. We further divided forests on non-carbonate substrate into two major groups – one thriving at high altitudes and the other in lowlands, both assigned to Piceion abietis. The second major group contains mesophilous, species-rich communities found mostly on carbonate bedrock. This major group can be further divided into four groups. The group of secondary forest on sites of beech forests, as well as the group of forest found in frost hollows can be assigned to Abieti-Piceion. The group of herb-rich, mesophilous communities forming zonal spruce and larch forests can be assigned to Chrysanthemo-Piceion and the group of forests on boulder scree to Calamagrostio-Abietion. Analysis of structural, functional, phytogeographical and geomorphological features, as well as ecological condi-tions estimated by bioindicator values, demonstrated the usefulness of this classification.