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The North Arnerican species of the genus Cremastocheilus are reviewed. These belong to 5 subgenera, Macropodina, Trinodea, Anatinodia, Mymcotonus, and Cremastocheilus. Taxonomie changes are: She inclusion of Crernastocheilus nitens and C. chapini in the subgenus Cremastocheilus rather than Myrmecotonus. Also Anatinodia is elevated to subgeneric status. A key to the subgenera is provided, as is a key to the species of the 5 subgenera, recognizing that the 35 species in the subgenus Cremastocheilus are in need of revision. A critical review of the host records, geographic distribution, and ecology of the Tribe Crernastocheilini (Family Scarabaeidae. subfamily Cetoniinae) is provided. This contains enormous numbers of new records for both the genera Genuchinus and CremastocheiLus both from the literature and from the extensive field work that is reported here for the first time. A Summary of the host records is presented in tabular form. This table shows the association of all species of Cremastocheilus with ants as adults and the larvae either associated with the vegetable material of the ant nests or with vegetable material in rodent burrows. Genuchinus is shown to be a general predator on soft bodied insects while the other genera of the Cremastocheilini are associated with plants, particularly bromeliads. A detailed study of the external morphology and sexual dimorphism of the genera Genuchinus and Crernastocheilus is presented. All species of Cremastocheilus can be sexed with the naked eye by the difference in the shapes of the abdominal terminal Segments, wherein males have the posterior border of the last ventral abdominal segment either straight or slightly bowed, while females have this border broadly rounded. There are other microscopic sexual differences in the structure of the legs. The rest of the external morphology is also presented, particularly from the point of view of adaptations to either a predaceous or rnyrmecophilous existente. Particularly adapted for predation are the pointed maxillae which are used for piercing prey. Particularly adapted for myrmecophily are the mentum, the maxillae, the generally thick exoskeleton, trichomes on both the anterior and posterior angles of the pronotum, the elytra, and the legs (which are adapted to the nest substrate of the host ant nests. Exocrine glands are described for Genuchinus ineptus and at least 1 species of each of the 5 subgenera of Cremastocheilus. In general, there are no gland cells nor glandular areas in Genuchinuc that are comparable to those of Cremastocheilus. The gland cells and glandular areas are quite extensive andvariable arnong species of Cremastocheilus. The frontal gland of some Cremastocheilus (strongly developed in C. castaneus and the C. canaliculatus species group, but weakly developed in the C. wheeleri species group) is described for the first time. Because these glands are not found in Genuchinus ineptuc, a species with general predatory habits, it is thought that these play a role, as yet unknown, in interactions with ants. The life cycles of the subgenera of Cremastocheilus are described. The general life cycle entails adult beetles eclosing in ant nests during the summer and then undertaking dispersal flights. The adults then enter ant nests and ovenivinter there, eating ant larvae during the Winter. Another dispersal flight occurs in the spring during which the adults mate and enter ant nests again. The females then lay eggs and the adults die. The eggs hatch and the larvae spend 3 instars feeding upon vegetable material in the nests. The lmae then pupate in typical scarabaeine earthen cells made of fecal material and soil. These eclose in the summer and the cycle is repeated. Variation from species to species is largely in the timing. Leaving the nest in late Summer, mating seems to be triggered by rainfall in all the species studied. Mating of C. (Macropodina) beameri takes place in rodent burrows. Males seem attracted to females from a distance but the mechanism of this remains obscure. In the subgenus Trinodia, mating takes place on sandy washes or roadsides where females land. In the subgenus Myrmecotonus, maüng also takes place in sandy areas. In C. (Cremastocheilus) mating takes place on sand bars along rivers in the southeastern U.S. and in sand dunes in northeastern U.S. The femaies dig down into the sand. Males locate these places by some unknown mechanism and then dig down to copulate with the females. Field experiments showed unequivocaily that males dig only into areas occupied by females. No sex-specific Sex attractant glands have been located in females so far. Dispersal to ant nests occurs after mating except for C. (Macropodina) beameri which lays its eggs in the rodent burrows and then probably disperses to ant nests. Beetle activity going in and out of nests was studied using wire hardware cloth screens over entrances to Mynnecocystus nests. The mesh size was such that the ants could move freely in or out but the beetles got stuck by their thoraces. The direction then could be interpreted by the direction in which they got stuck. By this method, C. stathamae was shown to leave nests from 23 June to 1 September with a peak on 6 July, just after the beginning of the summer rains. Beetles entered nests from June 23 to August 3, however 39% entered on July 16, probably pulsed by the leaving time which was correlated with the rains. Life cycle timing: C. (Macropodina) develop in the nests of Wood rats (Neotoma sp.]. Females lay about 40 eggs each. The 3 larval instars to pupation take about 1 month. Pupae are found from late August to weil into September. In other subgenera as well, larvae are found in parts of the nest devoid of ants, The timing is similar in all the subgenera found with ants. Mortality factors: While ants attack Cremastocheilus adults, there is no evidence that they are ever killed by ants nor is there evidence that ants kill larvae nor hard earthen pupae cases which protect the pupae. During dispersal fiights and mating, the adults are exposed to predation and evidence is presented that shows predation by horned toads, spiders, magpies, and tiger beetles. Probably most mortality occurs in the larval and pupd stages where the beetles are attacked by internal parasites and fungus. Further rnortality is caused by limitation of the food supply during the larval stage. Reentering nests: Females of C. (Macropodina) beameri select specific rodent and other burrows, attract males for rnating. and then enter the burrow for oviposition. C. stathamae are carried into the ants nests from as far away as 25ft. The beetles appear to land spontaneously after flying randomly over M. depilis nesting areas. Then the wander about waiting for the ants to carry them into the nests. Cremastocheilus hirsutus fly low over the ground searching for Pogonomyrrnex barbatus nests, land. and move straight for the nest entrances which they enter unhindered. Among all species, the ants frequently eject beetles but the net rnovement is in. Ants frequently attacked Cremastocheilus in laboratory observation nests when they were introduced. These attacks seldom resulted in the death of the beetles and the beetles were eventually ignored. When the beetles entered brood chambers, where they fed upon larvae, they were mostly ignored and even licked assiduously by the ants. A principle defensive behavior by the beetles is feigning death (letisimulation). The beetles give off an unpleasant "dead fish odor when collected in the I field. Experiments show that this substance functions to fend off some predators but further experiments indicated that these substances were ineffective against both ants and kangaroo rats. Experiments with various species of Cremastocheilus adults indicate that the adults eat only ant larvae. The beetles will eat larvae of non-host ants but show preferences for the larvae of their normal hosts. Under the same experimental conditions. Genuchinus ineptus adults will feed on a variety of insect adults and larvae. Field experiments on the function of trichome secretions did not indicate that they function to attract ants at a distance nor are they involved in worker acceptance. Laboratory experiments in which areas with a high concentration of gland cells were presented to ants showed that no ants were attracted. Laboratory introduction of Cremastocheilus hamisii adults into Fomica schau.si nests yielded many interactions including ants licking the anterior pronotal angles, the mentum area where the frontal glands empty and a carina over the eye with a dense pad of short setae. These are areas of concentration of gland cells and these are the first observations of licking by ants in specific sites containing exocrine glands. Radioisotope experiments showed food exchange among ants but never from ants to beetles. Other experiments showed that ants can pick up radioactivity from the beetles without feeding on trichome secretions. Evolutionary pathways: Adult Cremastocheilini probably followed the evolutionary route from adult predation on soft bodied insects to specialized feeding upon ant brood and the subsequent development of the beetle larvae in vegetable material in the ant colonies. Thus Genuchininseptus makes a logical outgroup in that they are general predators probably feeding mostly on Diptera larvae associated with Sotol plants in the field. The rnajor evolutionary step taken by Cremastocheiluswas to specialize on ant brood. Then the species radiated into ant colonies inhabiting southwestem North Arnenca. Most of the ant hosts invaded have quantities of vegetable material in their nests sufficient to support several developing scarab larvae. Host colonies are large, contain accessible brood, and are usually dominant foragers Evidence supports the idea that the species of Cremastocheilus have differentes in behavior and morphology that reflect adaptation to the behavioral ecology of different species of ants rather than different evolutionary levels of integration into ant colonies.
All over the world meat plays an important role in the nutrition of people. Mostly it is considered to be a special source of strength and health. In many peoples' minds the consumption of animal products, such as muscle, fat, blood, inner organs and bones, is much more associated with vital strength than a vegetarian meal. A reason for this may be the inherent physical similarity between human being and animal, especially mammals. There are other ways of producing meat, such as hunting and fishing, but today the most common method is butchering. The people in Tenkodogo consider beef to be an excellent meat. We will focus our comparative studies on special occasions, specialised butchers, locations, times, technical methods, distribution and ideas connected with the production and consumption of beef. Two fundamental reasons for the butchering of cattle can be identified: firstly, bulls are killed during the rituals of the year and secondly, cattle is slaughtered for daily commercial purposes on the market. In both cases almost the entire carcass of the butchered animal is consumed by people. In Tenkodogo we can actually compare those two different reasons, which have at least one common impact.
Tenkodogo, a township situated in the south-eastern part of Burkina Faso on the road leading from the capital Ouagadougou to the Togo border, has approximately 29,000 inhabitants. It is Burkina's seventh largest town and is the location of the regional government of the Boulgou-Province. This regional government is represented by a high-commissioner and a "préfet" as it is the residence of a traditional ruler, otherwise known as Tenkodogo-naaba. His sphere of influence covers many villages and hamlets in the region: in total he is the sovereign of nearly 120,000 people. The power of the traditional rulers was curtailed first by the arrival and following overrule of the French colonialists and then after independence by Sankara and his revolutionary government. The kings ceased to be the ultimate judges who were able to determine life and death of their subjects. Henceforth they were no longer allowed to recruit subjects for certain work on their fields, and they no longer could claim control over the allocation of resources. Their position was strengthened anew by Sankara's successor in office, Blaise Campaore, who quickly recognized that collaborating with the traditional rulers could only be of advantage: in fact they later proved to be his best supporters in the election campaign.
The tale portrays the unhappy life of a dove. Constantly surrounded by enemies, hunted by human beings and animals, disappointed by friends and separated from her family, the dove despairs of her life. She ponders over her unjust fate in this world and in a monologue she begins to consider, whether it would not be better to end her own life. This tragic theme forms the climax of several episodes, in which the tension between life and death is described. The elaborate development of dramatic acts demonstrates the intertwining of guilt and innocence in human existence.
In a recent article dealing with the Tangale Peak or Kilang, as it is called in the local Tangale language, Herrmann JUNGRAITHMAYR presents an account narrated by a Tangale elder about the attempted ascent of that characteristic mountain by a British colonial officer and his subsequent death.1 Kilang mountain is a basaltic cone approximately 1300 m high, about 8 km southwest of Kaltungo, one of the principal settlements of the Tangale people, in southern Bauchi State, northeastern Nigeria. During a research stay at the National Archives in Kaduna in November 1993 I was able to consult a file containing various documents relating to this incident in detail.2 In the following note I present an outline of the events based on the evidence in the colonial records. By doing this I not only intend to shed more light on a tragic event from the very early years of the colonial era. The picture of the circumstances emerging from the investigations of the colonial authorities may serve as a background to the narrative by the Tangale elder presented in JUNGRAITHMAYR's publication.
A calculation of the vacuum-polarization contribution to the hyperfine splitting for hydrogenlike atoms is presented. The extended nuclear charge distribution is taken into account. For the experimentally interesting case 209Bi82+ we predict a delta-lambda- -1.6 nm shift for the transition wavelength of the ground-state hyperfine splitting.
The electron-positron pairs observed in heavy-ion collisions at Gesellschaft für Schwerionen-forschung Darmstadt mbH have been interpreted as the decay products of yet unknown particles with masses around 1.8 MeV. The negative results of resonant Bhabha scattering experiments, however, do not support such an interpretation. Therefore we focus on a more complex decay scenario, where the e+e- lines result from a two-collision process. We discuss the induced decay of a metastable 1++ state into e+e- pairs. For most realizations of a 1++ state such a decay in leading order can only take place in the Coulomb field of a target atom. This fact has the attractive consequence that for such a state the Bhabha bounds are no longer valid. However, the absolute value of the e+e- production cross section turns out to be unacceptably small.
We investigate the possibility that high-energy photons are channeled, when passing through an oriented single crystal, due to Delbrück scattering. For this purpose the exact electron propagator for the single-string model is constructed. Starting from a separation of variables, we solve the Dirac equation for a cylindrical electrostatic potential. The propagator for such external fields is constructed from solutions of the radial Dirac equation. This propagator is applied to a calculation of the S matrix for Delbrück scattering. We specify the conditions under which photon channeling takes place. Unfortunately these conditions are only matched for a very small fraction of those photons being produced by channeled electrons.
Gitter sind diskrete additive Untergruppen des Rn. Praktische Bedeutung erlangte die Gittertheorie durch effziente Algorithmen zur Gitterbasenreduktion, mit deren Hilfe Optimierungsprobleme gelöst werden können. Der erste dieser Algorithmen wurde von Lenstra, Lenstra und Lovasz entwickelt. Schnorr und Euchner entwickelten effizientere Algorithmen. Sie untersuchten die Güte der Reduktion anhand von Rucksack-Problemen. Bei einem Rucksack-Problem der Dimension n müssen aus einer gegebenen Menge von n Gewichten diejenigen bestimmt werden, die zusammen einen gegeben Rucksack genau ausfüllen. Die Algorithmen von Schnorr und Euchner lösen fast alle Rucksack-Probleme der Dimensionen 42 bis 66. Meine neuen verbesserten Algorithmen lösen einen noch größeren Anteil der Rucksack-Probleme in kürzerer Rechenzeit. Gleichzeitig sind sie in Dimensionen 103 bis 151. Coster, Joux, LaMacchia. Odlyzko, Schnorr und Stern geben eine untere Schranke für die Größe der Gewichte von Rucksack-Problemen an, die fast immer gelöst werden können. Die Gewichte werden zufällig aus einem Intervall natüurlicher Zahlen gewählt. Dieses Ergebnis erweitere ich auf k-fache Rucksack-Probleme. Weiterhin kann für für die Wahl jedes Gewichtes eine beliebige Menge ganzer Zahlen festgelegt werden. Ebenso sind Mengen mit nur einem Element zulässig.