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Brooksia lacromae sp. nov. is described from zooplankton material collected at a marine monitoring station in the South Adriatic in the autumn of 2014. The description of both solitary and aggregate forms is given along with 18S rRNA and mitochondrial cox1 sequence data that provides strong evidence that both forms belong to the same species. The described species is morphologically markedly different from B. rostrata (Traustedt, 1893) and B. berneri van Soest, 1975, previously the only two species in the genus Brooksia. Genetic analysis based on 18S rRNA gene confirmed distinctness of B. lacromae sp. nov. from B. rostrata (1.5% uncorrected pairwise distance). The appendicularian Fritillaria helenae Bückmann, 1924, so far known from the Atlantic only, was found in the same samples as B. lacromae sp. nov. Co-occurrence of B. lacromae sp. nov. with an Atlantic appendicularian suggests an Atlantic or Western Mediterranean origin for this new taxon.
Three new species of Monepidosis Mamaev, 1966, a Holarctic genus of Porricondylinae (Diptera, Cecidomyiidae), are described: M. heterocera sp. nov. from Sweden and Germany, M. scepteroides sp. nov. from Sweden, and M. shikokuensis sp. nov. from Japan. A new porricondyline genus, Antipodosis gen. nov., is introduced for eight new species from New Zealand, named A. australis gen. et sp. nov., A. elongata gen. et sp. nov., A. granvillensis gen. et sp. nov., A. pureora gen. et sp. nov., A. rakiura gen. et sp. nov., A. rotoiti gen. et sp. nov., A. rotoroa gen. et sp. nov., and A. waipapa gen. et sp. nov. Male genitalic morphology indicates that Monepidosis and Antipodosis gen. nov. are closely related, together forming the Monepidosis group of genera, which stands out from the other Porricondylini. Monepidosis spatulata Spungis, 2006, a species originally described from Latvia and Lithuania, is for the first time reported to occur in Sweden.
The nine British and Irish species of Enicospilus are revised, mapped and an identification key provided. One species, Enicospilus myricae sp. nov., is described as new; Enicospilus merdarius (Gravenhorst, 1829) is a senior synonym of E. tournieri (Vollenhoven, 1879) syn. nov.; the only available name for E. merdarius auctt. is Enicospilus adustus (Haller, 1885) stat. rev., and a neotype is designated for Ophion adustus Haller, 1885. Enicospilus cerebrator Aubert, 1969 and E. repentinus (Holmgren, 1860) are newly recorded from Britain. Some host data are available for eight of the nine species.
The new genus Geotypodon gen. nov. is described. It includes two species from the Udzungwa Mountains: G. millemanus gen. et sp. nov. (type species) and G. submontanus gen. et sp. nov., one species from nearby Iringa: G. iringensis gen. et sp. nov., and 18 previously described species hitherto incorrectly assigned to Odontopyge Brandt, 1841.
Two new species from South Africa, Dactylonotus nigricorpus sp. nov. and Dactylonotus tsitsikamma sp. nov., are described and illustrated. D. nigricorpus sp. nov. differs from all other species of the genus in the black body, the smaller size and the shorter antenna. D. tsitsikamma sp. nov. is peculiar in the genus in bearing a flag of long setae on the fifth segment of the fore tarsus. An identification key to 6 Afrotropical species of the genus is provided.
New species of “giant” plume moths of the genus Platyptilia (Lepidoptera, Pterophoridae) from Uganda
(2016)
This paper describes two new species of plume moths from the group of the so-called “giant” Platyptilia Hubner, 1825: Platyptilia fletcheri Ustjuzhanin & Kovtunovich sp. nov. and P. stanleyi Ustjuzhanin & Kovtunovich sp. nov. Both species were collected in the Rwenzori Mountains in Uganda and Rwanda, respectively. Platyptilia stanleyi Ustjuzhanin & Kovtunovich sp. nov. exceeds all the known African species of Pterophoridae in its wingspan of 49 mm.
Fifteen new species of Begonia L. from Sumatra are described and illustrated, in Begonia sect. Bracteibegonia (B. beludruvenea M.Hughes sp. nov. and B. jackiana M.Hughes sp. nov.), B. sect. Petermannia (B. harauensis Girm. sp. nov.), B. sect. Platycentrum (B. leuserensis M.Hughes sp. nov.), B. sect. Reichenheimia (B. fl uvialis M.Hughes sp. nov., B. halabanensis M.Hughes sp. nov., B. karangputihensis Girm. sp. nov., B. kemumuensis M.Hughes sp. nov., B. korthalsiana Miq. ex M.Hughes sp. nov., B. kudoensis Girm. sp. nov., B. lilliputana M.Hughes sp. nov., B. olivacea Ardi sp. nov., B. raoensis M.Hughes sp. nov., B. simolapensis Ardi sp. nov.) and B. sect. Sphenanthera (B. pseudoscottii Girm. sp. nov.). Using the International Union for the Conservation of Nature criteria, 6 are considered to be Least Concern, 5 Vulnerable and 4 Data Deficient. A key to 58 of the 63 currently accepted Begonia species in Sumatra is provided.
The study of subterranean Oniscidea in Portugal has been neglected for nearly 70 years, but recent investigations have revealed high diversity. All the terrestrial isopods known from caves of mainland Portugal, including data from the literature and new material, are listed, revealing new biodiversity patterns. Twenty-seven species, belonging to 15 genera and six families, are known, of which 16 species are considered to be exclusively from subterranean ecosystems, i.e., troglobionts. Trichoniscidae is the most diverse family. Seven species in the family Trichoniscidae and one species in the family Styloniscidae are described as new (all with Reboleira & Taiti as authors): Trichoniscoides bellesi sp. nov. from the Montejunto Massif; T. sicoensis sp. nov. from the Sicó Massif; Metatrichoniscoides salirensis sp. nov. from the typhonic valley of Caldas da Rainha; Troglonethes olissipoensis sp. nov. from the Lisbon Peninsula; T. arrabidaensis sp. nov. from the Arrábida Massif; Miktoniscus longispina sp. nov. from the Sicó Massif and Cesaredas Plateau; Moserius inexpectatus sp. nov. from the Estremenho Massif; and Cordioniscus lusitanicus sp. nov. from Alentejo and Algarve, the southernmost provinces of Portugal. The subgenus Trogleluma Vandel, 1946 (Armadillidiidae) is raised to genus level. In this geographic region terrestrial isopods are the richest group of cave-adapted animals.
A new monospecific genus, Pseudostegopterus gen. nov., endemic to the northwestern region of South Africa, is erected. The type species is described as Pseudospegopterus melonthinoides sp. nov. and is currently known only from male specimens. A provisional dichotomic key of the African Trichiina genera is also provided, in order to facilitate the identification of male specimens to the genus level.
Amage imajimai sp. nov., a new species of Ampharetidae (Annelida: Polychaeta) from Japanese waters
(2015)
A new polychaete species of the family Ampharetidae, Amage imajimai sp. nov., is described from deep waters of Sagami Bay, Japan. It is characterized by the possession of four pairs of branchiae, twelve thoracic uncinigers, eleven abdominal uncinigers, and the lack of thoracic notopodial cirri. The new species is named in honor of the renowned Japanese polychaetologist Minoru Imajima. An identification key for all Amage species from Japanese waters is provided.
Two new junior synonyms, Pseudomesitius Duchaussoy, 1916 syn. nov. and Triglenus Marshall, 1905 syn. nov., for the genus Bradepyris Kieffer, 1905 are proposed and a new diagnosis for the genus is given. Two new species (based on males) from the Saharo-Arabian region are here described: Bradepyris jordanicus, sp. nov. and B. baleariensis, sp. nov. A brief revision, with a key to the males and females of all species of this genus, and the first male genitalia descriptions are also provided.
The genus Aquattuor Frederiksen, 2013 is revised. A. denticulatus Frederiksen, 2013 (type species) from the East Usambara Mts, Tanzania, is redescribed, and six new species are described: A. claudiahempae sp. nov. from Mt. Kilimanjaro, Tanzania, and five species from the Udzungwa Mts, Tanzania: A. longipala Enghoff sp. nov., A. major Enghoff sp. nov., A. stereosathe Enghoff sp. nov., A. submajor Enghoff sp. nov., and A. udzungwensis Enghoff sp. nov.
Four new species of the genus Caccothryptus (Coleoptera: Limnichidae) are described: C. taiwanus from Taiwan; C. orion from Okinawa; C. tibetanus and C. chayuensis from Tibet. All the species belong to the testudo species group (sensu Hernando & Ribera 2014). Additional specimen data and an updated species list are also given, and C. testudo Champion, 1923 is newly recorded from Thailand.
Actaea grimaldii, a new species of reef crab from Papua New Guinea (Crustacea, Brachyura, Xanthidae)
(2015)
A new species of xanthid crab, Actaea grimaldii, is described from the coral reefs of Papua New Guinea. This species has a distinctive red and white coloration and is closest to Actaea spinosissima Borradaile, 1902, from the Indian Ocean. However, the new species can be distinguished by the arrangement of spines on the carapace, chelipeds and ambulatory legs, and the structure of the male gonopods. Actaea grimaldii sp. nov. has also been confused with A. polyacantha (Heller, 1861), but differs markedly in the carapace armature.
New and interesting Surirella taxa (Surirellaceae, Bacillariophyta) from the Congo Basin (DR Congo)
(2015)
Two new diatom taxa belonging to the genus Surirella, S. ebalensis and S. congolensis, are described from material of the Congo Basin, downstream Kisangani, DR Congo. The first taxon is a small, rather common species in the studied material; the second a somewhat larger diatom that was only sporadically observed. The morphology of both taxa is examined with light and scanning electron microscopy. The differences between the new and other closely related taxa such as S. agonaensis and S. bonsaensis, and S. takoradiensis, S. tenuissima and S. pseudotenuissima, respectively, are discussed.
Based on intensive collecting from various sites in Sweden, the genus Dasyhelea Kieffer, 1911 was reviewed and the number of species now known from the country increased from five to twenty. Among the new species recorded there are two species described as new to science, D. dominiakae sp. nov. and D. gothlandica sp. nov., both in the subgenus Dicryptoscena Enderlein, 1936. The following subgenera are now documented from Sweden: Dasyhelea, Dicryptoscena, Pseudoculicoides Malloch, 1915, Prokempia Kieffer, 1913 and Sebessia Remm, 1979, the two latter subgenera being reported for the first time.
A new species of Paracrobeles, P. kelsodunensis sp. nov. is described from the Kelso Dunes area, Mojave National Preserve, southern California. Paracrobeles kelsodunensis sp. nov. is particularly characterised by a body length of 469–626 μm in females and 463–569 μm in males; lateral field with four incisures, extending almost to tail terminus; three pairs of asymmetrical lips, separated by U-shaped primary axils with two long guarding processes, each lip usually with four tines along its margin; three long labial probolae, deeply bifurcated, with slender prongs without tines; metastegostom with a strong anteriorly directed dorsal tooth; pharyngeal corpus anteriorly spindle-shaped, posteriorly elongate bulbous with dilated lumen; spermatheca 24–87 μm long; postvulval uterine sac 60–133 μm long; vulva in a sunken area; spicules 33–38 μm long; and male tail with a 5–8 μm long mucro. The generic diagnosis is emended on the basis of recently described species and a key to the species of Paracrobeles is provided.
Two new species of Elaphropeza Macquart, 1827 are described from the northeast coast of Bali (Indonesia): Elaphropeza triseta sp. nov. belonging to the ephippiata-group and E. balinensis sp. nov. belonging to the biuncinata-group. A COI Neighbour-Joining tree is given for the Southeast Asian Elaphropeza species showing large genetic distances between the species. The morphological characterisation of the ephippiata-group and the biuncinata-group sensu Shamshev & Grootaert 2007 is confirmed.
We report here on recent collections of freshwater crabs from Antsiranana Province, northern Madagascar. The specimens belong to three species, one of which is new to science and is described here. This raises the number of species of freshwater crabs found in Madagascar to 17. All are endemic to the island and all belong to the Afrotropical family Potamonautidae Bott, 1970. The new species, Foza manonae sp. nov., is compared to the other species in this genus, and an updated key is provided. It is distinguished from the other three congeners by characters of the male first gonopod, sternum, carapace, and cheliped. The conservation status of the Malagasy freshwater crab fauna is summarized and discussed in light of the new material reported on here belonging to two other species, Madagapotamon humberti Bott, 1965 and Foza ambohitra Cumberlidge & Meyer, 2009.
Species of Mortoniella are revised for the northern and Andean part of the South American continent, including the countries of Bolivia, Peru, Ecuador, Colombia, Venezuela, and Guyana. All previously described species from the region are reillustrated and redescribed, except for Mortoniella santiaga Sykora, 1999 and M. quinuas Harper and Turcotte, 1985, whose types could not be located, and M. tranquilla Martynov, 1912, whose type is based on a female specimen and thus is currently unidentifiable. Included in the revision are 35 described species and 59 new species. Mortoniella similis Sykora, 1999 is considered a junior synonym of M. roldani Flint, 1991, and M. macuta (Botosaneanu, 1998) is considered a junior synonym of M. limona (Flint, 1981). A new subgenus, Nanotrichia, is recognized to accommodate species previously referred to as members of the ormina and velasquezi groups. Mexitrichia pacuara Flint, 1974 is designated the type species for the subgenus. Species previously referred to as members of the bilineata and leroda species groups are retained in the nominate subgenus, along with additional taxa not previously placed to species group, and treated within a number of subgroups. Previously described species of M. (Mortoniella) which are redescribed and reillustrated include: M. angulata Flint, 1963; M. apiculata Flint, 1963; M. atenuata (Flint, 1963); M. bifurcata Sykora, 1999; M. bilineata Ulmer, 1906; M. bolivica (Schmid, 1958); M. chicana Sykora, 1999; M. denticulata Sykora, 1999; M. elongata (Flint, 1963); M. enchrysa Flint, 1991; M. flinti Sykora, 1999; M. foersteri (Schmid, 1964); M. hodgesi Flint, 1963; M. iridescens Flint, 1991; M. leei (Flint, 1974); M. limona (Flint, 1981); M. marini (Rueda Martín and Gibon, 2008); M. paralineata Sykora, 1999; M. paraenchrysa Sykora, 1999; M. pocita (Flint, 1983); M. punensis (Flint, 1983); M. roldani Flint, 1991; M. simla (Flint, 1974); M. spinulata (Flint, 1991); M. squamata Sykora, 1999; M. unilineata Sykora, 1999; and M. wygodzinskii (Schmid, 1958). New species described in M. (Mortoniella), followed by their respective areas of distribution, include: M. acutiterga (Ecuador); M. adamsae (Peru); M. akrogeneios (Ecuador); M. applanata (Peru); M. auricularis (Colombia); M. barinasi (Venezuela); M. biramosa (Venezuela); M. bothrops (Peru); M. brevis (Ecuador, Venezuela); M. bulbosa (Peru); M. catherinae (Peru); M. chalalan (Peru); M. cornuta (Peru); M. cressae (Venezuela); M. croca (Peru); M. curtispina (Venezuela); M. curvistylus (Ecuador); M. dentiterga (Ecuador); M. dinotes (Peru); M. draconis (Ecuador); M. emarginata (Ecuador, Colombia); M. esrossi (Colombia); M. flexuosa (Colombia); M. furcula (Ecuador); M. gilli (Ecuador); M. gracilis (Venezuela); M. grandiloba (Venezuela); M. guyanensis (Guyana); M. hamata (Colombia); M. langleyae (Ecuador); M. longiterga (Ecuador); M. membranacea (Bolivia); M. monopodis (Colombia, Ecuador); M. parameralda (Ecuador); M. pica (Ecuador); M. proakantha (Ecuador); M. prolata (Peru); M. quadrispina (Ecuador); M. rectiflexa (Ecuador); M. ruedae (Bolivia); M. schlingeri (Colombia); M. silacea (Colombia, Ecuador); M. sinuosa (Bolivia, Peru); M. spatulata (Venezuela); M. tanyrhabdos (Venezuela); M. tridens (Peru); M. triramosa (Bolivia); M. tusci (Venezuela); and M. variabilis (Venezuela, Colombia). Species assigned to the subgenus M. (Nanotrichia) which are redescribed and reillustrated include: M. aequalis (Flint, 1963); M. aries (Flint, 1963); M. collegarum (Rueda Martín and Gibon, 2008); M. eduardoi (Rueda Martín and Gibon, 2008); M. macarenica (Flint, 1974); M. pacuara (Flint, 1974); M. usseglioi (Rueda Martín and Gibon, 2008); and M. velasquezi (Flint, 1991). Previously described species of Mortoniella, outside the area of coverage, that are transferred to the subgenus M. (Nanotrichia) include: M. alicula Blahnik and Holzenthal, 2011; M. bocaina Blahnik and Holzenthal, 2011; M. catarinensis (Flint, 1974); M. froehlichi Blahnik and Holzenthal, 2011; M. ormina (Mosely, 1939); M. rodmani Blahnik and Holzenthal, 2008; and M. tripuiensis Blahnik and Holzenthal, 2011. New species in the subgenus M. (Nanotrichia), followed by their respective areas of distribution, include: Mortoniella cognata (Ecuador, Venezuela); M. coheni (Ecuador); M. licina (Ecuador); M. paucispina (Peru); M. quadridactyla (Venezuela); M. simplicis (Venezuela); M. spangleri (Ecuador); M. triangularis (Ecuador); M. venezuelensis (Venezuela); and M. zamora (Ecuador). A key to the males of species from the region is also provided, as well as a key to females for the major subgroups and a species key to females of the velasquezi group. Finally, a partially resolved phylogeny of the species is presented, along with a discussion of evolutionary trends within the genus.
Species of Mortoniella are revised for the northern and Andean part of the South American continent, including the countries of Bolivia, Peru, Ecuador, Colombia, Venezuela, and Guyana. All previously described species from the region are reillustrated and redescribed, except for Mortoniella santiaga Sykora, 1999 and M. quinuas Harper and Turcotte, 1985, whose types could not be located, and M. tranquilla Martynov, 1912, whose type is based on a female specimen and thus is currently unidentifiable. Included in the revision are 35 described species and 59 new species. Mortoniella similis Sykora, 1999 is considered a junior synonym of M. roldani Flint, 1991, and M. macuta (Botosaneanu, 1998) is considered a junior synonym of M. limona (Flint, 1981). A new subgenus, Nanotrichia, is recognized to accommodate species previously referred to as members of the ormina and velasquezi groups. Mexitrichia pacuara Flint, 1974 is designated the type species for the subgenus. Species previously referred to as members of the bilineata and leroda species groups are retained in the nominate subgenus, along with additional taxa not previously placed to species group, and treated within a number of subgroups. Previously described species of M. (Mortoniella) which are redescribed and reillustrated include: M. angulata Flint, 1963; M. apiculata Flint, 1963; M. atenuata (Flint, 1963); M. bifurcata Sykora, 1999; M. bilineata Ulmer, 1906; M. bolivica (Schmid, 1958); M. chicana Sykora, 1999; M. denticulata Sykora, 1999; M. elongata (Flint, 1963); M. enchrysa Flint, 1991; M. flinti Sykora, 1999; M. foersteri (Schmid, 1964); M. hodgesi Flint, 1963; M. iridescens Flint, 1991; M. leei (Flint, 1974); M. limona (Flint, 1981); M. marini (Rueda Martín and Gibon, 2008); M. paralineata Sykora, 1999; M. paraenchrysa Sykora, 1999; M. pocita (Flint, 1983); M. punensis (Flint, 1983); M. roldani Flint, 1991; M. simla (Flint, 1974); M. spinulata (Flint, 1991); M. squamata Sykora, 1999; M. unilineata Sykora, 1999; and M. wygodzinskii (Schmid, 1958). New species described in M. (Mortoniella), followed by their respective areas of distribution, include: M. acutiterga (Ecuador); M. adamsae (Peru); M. akrogeneios (Ecuador); M. applanata (Peru); M. auricularis (Colombia); M. barinasi (Venezuela); M. biramosa (Venezuela); M. bothrops (Peru); M. brevis (Ecuador, Venezuela); M. bulbosa (Peru); M. catherinae (Peru); M. chalalan (Peru); M. cornuta (Peru); M. cressae (Venezuela); M. croca (Peru); M. curtispina (Venezuela); M. curvistylus (Ecuador); M. dentiterga (Ecuador); M. dinotes (Peru); M. draconis (Ecuador); M. emarginata (Ecuador, Colombia); M. esrossi (Colombia); M. flexuosa (Colombia); M. furcula (Ecuador); M. gilli (Ecuador); M. gracilis (Venezuela); M. grandiloba (Venezuela); M. guyanensis (Guyana); M. hamata (Colombia); M. langleyae (Ecuador); M. longiterga (Ecuador); M. membranacea (Bolivia); M. monopodis (Colombia, Ecuador); M. parameralda (Ecuador); M. pica (Ecuador); M. proakantha (Ecuador); M. prolata (Peru); M. quadrispina (Ecuador); M. rectiflexa (Ecuador); M. ruedae (Bolivia); M. schlingeri (Colombia); M. silacea (Colombia, Ecuador); M. sinuosa (Bolivia, Peru); M. spatulata (Venezuela); M. tanyrhabdos (Venezuela); M. tridens (Peru); M. triramosa (Bolivia); M. tusci (Venezuela); and M. variabilis (Venezuela, Colombia). Species assigned to the subgenus M. (Nanotrichia) which are redescribed and reillustrated include: M. aequalis (Flint, 1963); M. aries (Flint, 1963); M. collegarum (Rueda Martín and Gibon, 2008); M. eduardoi (Rueda Martín and Gibon, 2008); M. macarenica (Flint, 1974); M. pacuara (Flint, 1974); M. usseglioi (Rueda Martín and Gibon, 2008); and M. velasquezi (Flint, 1991). Previously described species of Mortoniella, outside the area of coverage, that are transferred to the subgenus M. (Nanotrichia) include: M. alicula Blahnik and Holzenthal, 2011; M. bocaina Blahnik and Holzenthal, 2011; M. catarinensis (Flint, 1974); M. froehlichi Blahnik and Holzenthal, 2011; M. ormina (Mosely, 1939); M. rodmani Blahnik and Holzenthal, 2008; and M. tripuiensis Blahnik and Holzenthal, 2011. New species in the subgenus M. (Nanotrichia), followed by their respective areas of distribution, include: Mortoniella cognata (Ecuador, Venezuela); M. coheni (Ecuador); M. licina (Ecuador); M. paucispina (Peru); M. quadridactyla (Venezuela); M. simplicis (Venezuela); M. spangleri (Ecuador); M. triangularis (Ecuador); M. venezuelensis (Venezuela); and M. zamora (Ecuador). A key to the males of species from the region is also provided, as well as a key to females for the major subgroups and a species key to females of the velasquezi group. Finally, a partially resolved phylogeny of the species is presented, along with a discussion of evolutionary trends within the genus.
Many nomenclatural changes are implemented in the beetle families Georissidae, Histeridae, Hydraenidae, Hydrochidae, Hydrophilidae, Ptiliidae, Leiodidae and especially Staphylinidae, of the beetle series Staphyliniformia (Coleoptera), in preparation for making a world catalog of this group available online. Limited taxonomic changes are also made in the staphylinid subfamilies Osoriinae and Staphylininae.
At the level of family-group taxa, Article 29.4 of the current (1999) Zoological Code is reviewed and the original spellings of two tribal names, Nymphisterini Tishechkin (Histeridae) and Cryptonotopsisini Pace (Staphylinidae), are resurrected. The tribal name Stictocraniini Jakobson (Staphylinidae) is also resurrected as the valid name for its new synonym Fenderiini Scheerpeltz.
Changes at the genus-group level in Histeridae include placing Contipus Marseul as a new synonym of Hister Linnaeus due to the current placement of its validly designated type species C. subquadratus Marseul; proposal of Contipides Newton gen. nov. (type species Contipus digitatus Marseul) for the 10 species that had remained in Contipus of authors; and new designation of Idolia laevigata Lewis as type species of Idolia Lewis. In Ptiliidae, Rodwayia ovata Lea is newly designated as type species of Rodwayia Lea, and Throscidium germainii Matthews is newly designated as type species of Throscidium Matthews. In Staphylinidae, Paramichrotus Naomi is resurrected as a valid subgenus of Hesperosoma Scheerpeltz with Hemihesperosoma Hayashi placed as a new synonym of it; Sonoma corticina Casey is reaffi rmed as the type species of Sonoma Casey in place of Faronus tolulae LeConte; Stanosthetus Dejean is recognized as an available name and junior synonym of Euplectus Kirby; Taplandria Pace (type species T. guyanensis Pace) is recognized as a junior homonym and new synonym of Taplandria Pace (type species T. fl ava Pace); and Termitobiella Wasmann is resurrected as the valid name for the genus Felda Blackwelder. Replacement names for preoccupied generic or subgeneric names include in Histeridae Bellatricides Newton nom. nov. for Pachylister (Bellatrix) Mazur, junior homonym of Bellatrix Boie; and in Staphylinidae Foxiides Newton nom. nov. for Foxia Pace, junior homonym of Foxia Ashmead, and Xenasterides Newton nom. nov. for Xenaster Bierig, junior homonym of Xenaster Simonwitsch. Taxonomic changes at the generic level in Staphylinidae include proposal of Prolibia Newton gen. nov. (type species Lispinus californicus LeConte) for four Nearctic species recently placed in Clavilispinus Bernhauer; placement of Heterotrochinus Coiffait and its synonym Heterotrochus Coiffait as new synonyms of Eulibia Cameron; placement of the generic or subgeneric names Chapmaniella Bernhauer, Glenothorax Bierig, Euryolinus Bernhauer and Plesiolinus Bernhauer as new synonyms of Platydracus Thomson; and transfer of the subgenus Poikilodracus Scheerpeltz from Staphylinus Linnaeus to Platydracus. First reviser actions are used to select Georissites Ponomarenko (Georissidae) as the correct original spelling over the alternate original spelling Georyssites, and Kyrtusa Pace (Staphylinidae) as correct original spelling over Kirtusa.
Several hundred nomenclatural and taxonomic changes at the species group level are briefl y summarized here but are too numerous to list completely. Replacement names for preoccupied species or subspecies names in current use are proposed in Histeridae (3), Hydrochidae (1), Hydrophilidae (1), Leiodidae (2), Ptiliidae (3) and Staphylinidae (180); an additional staphylinid replacement name, Phloeopora nilgiriensis, is newly proposed by G. Paśnik. New or resurrected combinations are proposed for either nomenclatural or taxonomic reasons in the following genera (with indication of how many names in each genus): in Histeridae, Contipides Newton (10); in Staphylinidae, Abemus Mulsant and Rey (4), Allotrochus Fagel (6), Atheta Thomson (1), Cheilocolpus Solier (4), Eulibia Cameron (4), Foxiides Newton (1), Lispinus Erichson (3), Loncovilius Germain (2), Nacaeus Blackwelder (119), Naddia Fauvel (1), Neohypnus Coiffait and Sáiz (8), Neolosus Blackwelder (1), Ocypus Leach (2), Ontholestes Ganglbauer (1), Platydracus Thomson (59), Prolibia Newton (4) Termitobiella Wasmann (10), Thyreocephalus Guérin-Méneville (4), Xenasterides Newton (1), and Zeoleusis Steel (3). First reviser actions are used to resolve the correct original spellings (of two or more original spellings) of two species of Hydraena Kugelann (Hydraenidae) and 21 species of Staphylinidae. Changes in priority or availability of names are cited to establish the following names as valid over one or more new synonyms each: Acrotrichis rotundata (Haldeman) and Acrotrichis glabricollides Newton sp. nov. in Ptiliidae, Nemadiopsis franki Perreau in Leiodidae, and Gyrophaena nigra Kraatz, Heterothops fumigatus LeConte, Loncovilius germaini (Scheerpeltz), Philonthus upotovus Newton, sp. nov., Stenus fulviventris Rougemont, and nine species of Homalota Mannerheim in Staphylinidae. Finally, the species Eleusis lata Coiffait and Eleusis microlestiformis Coiffait are noted as not belonging to the genus Eleusis Laporte de Castelnau or to Staphylinidae, and are transferred without generic assignment to the subfamily Inopeplinae of the family Salpingidae.
A systematic redefinition of the species belonging to the genus Geomyphilus Gordon and Skelley, 2007 (Coleoptera: Scarabaeidae: Aphodiinae) of Mexico and neighboring countries is presented. The new species G. tuzincola of Mexico is described and figured. The new combination Coelotrachelus macgregori (Islas, 1955) is proposed.
Two new genera and species of tiger beetles from Baltic amber (Coleoptera: Carabidae: Cicindelinae)
(2017)
Two fossil tiger beetle species (Coleoptera, Carabidae, Cicindelinae) are described from Eocene Baltic amber using light microscopic and X-ray microscopic techniques. Both species are considered representatives of the subtribe Iresina Rivalier, 1971 due to the shared combination of character states: glabrous head, six labral and four suborbital setae, and glabrous pronotum. Palaeopronyssiformia groehni Wiesner, Will, and Schmidt, new genus, new species, is characterized by a glabrous and furrowed head with six labral setae, large eyes, presence of two supraorbital setae on each side, mandibles with two teeth of the incisor region, and a glabrous and furrowed pronotum. Palaeoiresina cassolai Wiesner, Will, and Schmidt, new genus, new species, is characterized by a unicolored, undentated labrum, mandibles with two teeth of the incisor region, glabrous head with six labral setae, two clypeal setae, two supraorbital setae on each side, and a glabrous pronotum, mesepisternum, mesepimeron, and metepisternum. The species described here represent the only known tiger beetle fossils preserved in Baltic amber.
The New World genus Chariessa Forster (Coleoptera: Cleroidea: Cleridae) is revised and includes C. catalina Opitz, new species, C. elegans Horn, C. dichroa (LeConte), C. floridana Schaeffer, C. pilosa (Forster), C. texana Wolcott, C. ramicornis Perty, C. vestita (Chevrolat), and C. duponti (Spinola). Enoplium pilosa var. marginata Say is synonymized with Chariessa pilosa Forster. Lectotypes are designated for C. pilosa (Forster), C. ramicornis Perty, and C. vestita (Chevrolat). Available information indicates that Chariessa adult and immature individuals are predatory on lignicolous insects with a particular affinity for cerambycids and buprestids that infest species of oak. It is postulated that Pleistocene speciation generated the North American components of Chariessa with more ancient southern species generated during the Middle Tertiary; after closures of the Middle American portals and orogeny of the South American Andes. Included in this treatise is a discussion of natural history, key to species, narratives of zoogeography and phylogeny, one diagram of a phylogenetic tree, 35 line drawings, eight SEM micrographs, twelve habitus photographs, nine photographs of male genitalia, and five distributional maps.
The eight species in the genus Tomarus Erichson (Coleoptera: Scarabaeidae) in Argentina, Chile, and Uruguay are reviewed. Tomarus roigjunenti new species and Tomarus spinipenis new species are described from Argentina. We include a key to species, representative habitus illustrations for all species, character illustrations, and distribution maps for each, as well as commentary about the natural history and distributions for each species. Diagnostic characters are discussed for each species, and species relationships are hypothesized based on the analysis of internal and external morphological characters. The male of T. bidentulus (Fairmaire) is described for fi rst time. The following taxonomic changes are made: Tomarus guianucai Dechambre and Lumaret, 1985 is a new junior synonym of Tomarus rubripes (Boheman, 1858), which was formerly and incorrectly cited as occurring in Argentina.
Species descriptions, keys to genera and species, and geographical distributions are presented for 43 species of the family Bruchidae (Coleoptera: Chrysomeloidea) for Chile. Of these species, seven are described as new:
Acanthoscelides aricae sp. nov., Lithraeus chillan sp. nov., L. comptus sp. nov., L. elguetai sp. nov., L. limari sp. nov., L. lonquimay sp. nov., and L. penai sp. nov. Eight species are endemic to Chile. A list of true host plants and floral records for those with known host associations is presented. Habitus photographs and drawings of pertinent body parts, including male genitalia, are provided. References pertaining to the previously described species are listed.
The first myrmecophilous fl ea beetle genus (Myrmeconycha Konstantinov and Tishechkin, new genus) with four new species (M. erwini Konstantinov and Tishechkin, new species – Ecuador, M. gordoni Konstantinov and Tishechkin, new species – Brazil, M. pakaluki Konstantinov and Tishechkin, new species – Panama, and M. pheidole Konstantinov and Tishechkin, new species – Costa Rica) is described and illustrated. It is compared with fl ea beetles of the subtribe Disonychina (Coleoptera: Chrysomelidae: Galerucinae: Alticini) and may be easily differentiated based on the external and internal features, which include the waxy surface of the head and pronotum, reticulated surface of the pronotum, and four longitudinal ridges on each elytron.
New data on Odonata of the Guadalcanal Island, Solomon Islands are provided following a recently completed Rapid Biodiversity Assessment of the Tetena Haiaja ridge. Two new species, Lieftinckia ulunorum and Procordulia valevahalo are described.
The first is a new member of the Solomon Islands endemic genus while the second is a new genus for the country and the second validated species from the Corduliidae family known from this Pacific archipelago. As L. ulunorum is found to be very closely related to formerly known L. lairdi Lieftinck, 1963, which was also collected during the field trip, both are described in detail based on mature adults and teneral specimens. Comparison with L. salomonis Kimmins, 1957 (investigated only from figures published in the original species description) and Salomoncnemis gerdae Lieftinck, 1987 (also sampled during this study) were provided as well.
Additional morphological data is given on the following species: Teinobasis bradleyi Kimmins, 1957, female is illustrated here for the first time; Anax sp. cf. gibbosulus, second record of the genus for the country and Gynacantha amphora Marinov & Theischinger, 2012, originally described by a single male, here the description of the female is provided.
All other species collected during the field trip will be published separately in the final expedition report.
Six new species of Platypalpus Macquart, 1827 are described from tropical forest at Yangambi (Democratic Republic of the Congo): Platypalpus bolikoi sp. nov., P. ikoso sp. nov., P. lokonda sp. nov., P. manjano sp. nov., P. saffradi sp. nov. and P. yangambensis sp. nov. All species are photographed and, except for P. saffradi sp. nov. known only from females, male terminalia are illustrated for all. A key is provided for the six species of DR Congo. COI barcodes are available for all species at GenBank.
Twenty new species of the millipede genus Chaleponcus Attems, 1914, are described from the Udzungwa Mountains: C. netus sp. nov., C. quasimodo sp. nov., C. malleolus sp. nov., C. scopus sp. nov., C. nikolajscharffi sp. nov., C. mwanihanensis sp. nov., C. basiliscus sp. nov., C. krai sp. nov., C. nectarinia sp. nov., C. circumvallatus sp. nov., C. ibis sp. nov., C. vandenspiegeli sp. nov., C. vilici sp. nov., C. teres sp. nov., C. hamerae sp. nov., C. termini sp. nov., C. gracilior sp. nov., C. mwabvui sp. nov., C. howelli sp. nov. and C. tintin sp. nov. Together with C. dabagaensis Kraus, 1958, they constitute the Chaleponcus dabagaensis-group, well characterized by apparently apomorphic gonopodal characters, presumably monophyletic, and the first example of a major radiation within the Udzungwas. All species are restricted to altitudes >1390 m, all but one were found in only one, rarely two forest reserves, and the vast majority of specimens were collected in montane forest. Chaleponcus gracilior sp. nov. was collected in four forest reserves, often in secondary habitats where other species were only exceptionally found. Co-occurrence of multiple species, inter-specific differences in body size and unusual tarsal setation of a few species tentatively suggest adaptive radiation.
The endophallic structure of the genus Laius is studied and discussed based on the examination of 19 species from Asia to the Indian Ocean. The structure contains two primary sclerites (named gonoporal piece and ligula), a secondary sclerite on the basal part of the gonoporal piece (named additional sclerite) in some species, and a membranous basal area closely covered with many spines (named spinous area). Five species groups are recognized based on the morphology of the endophallic sclerites. The sympatric species have different body sizes and quite distinguishable endophallic sclerites (= different species group), while the allopatric species have overlapping body sizes and similar endophallic sclerites (= same species group). Three new species are described and six previously known species are redescribed with endophallic sclerites, and the descriptions of endophallic sclerites of the remaining ten species are added. The larva of Laius rodriguesensis sp. nov. is also described. The genus
Nossibeus Evers, 1994 is synonymised with Laius Guérin-Méneville, 1830.
The South African endemic bees of the "euryglossiform" species of the genus Scrapter Lepeletier & Serville, 1828 are revised and illustrated. The species-group is defined for the first time and comprises 20 species, 16 of which are described here as new: Scrapter exiguus sp. nov. ♀, ♂, S. gessorum sp. nov. ♀, S. inexpectatus sp. nov. ♀, S. luteistigma sp. nov. ♀, ♂, S. minutissimus sp. nov. ♂, S. minutuloides sp. nov. ♀, S. minutus sp. nov. ♀, S. nanus sp. nov. ♀, ♂, S. nigerrimus sp. nov. ♀, S. nigritarsis sp. nov. ♀, S. papkuilsi sp. nov. ♀, ♂, S. punctatus sp. nov. ♀, ♂, S. pygmaeus sp. nov. ♀, S. roggeveldi sp. nov. ♀, ♂, S. spinipes sp. nov. ♀, ♂ and S. ulrikae sp. nov. ♀, ♂. For S. acanthophorus Davies, 2005 and S. sittybon Davies, 2005 the female is here described for the first time. A key to all species is provided.
Three new species of Pachygnatha, P. bispiralis sp. nov., P. intermedia sp. nov. and P. ventricosa sp. nov., are described from forest areas in western Burundi. The presence of P. procincta Bosmans & Bosselaers, 1994 in Burundi confirms its very wide distribution spanning most of Africa.
Pachygnatha appears to be an important element of the afromontane spider fauna.
Populations of Stegelleta are described from California, New Zealand and Senegal. An amphimictic population from California is identified as belonging to S. incisa and compared with type specimens from Utah and an amphimictic population from Italy. One population from New Zealand is close to S. incisa but considered to represent a new species, Stegelleta laterocornuta sp. nov. It is particularly characterised by a 379–512 μm long body in females and 365–476 μm in males; cuticle divided into 16 rows of blocks at midbody (excluding lateral field); lateral field with four incisures; three pairs of asymmetrical lips, U-shaped primary axils without guarding processes, each lip asymmetrically rectangular with a smooth margin, only lateral lips have slender acute tines; three labial probolae, bifurcated at half of their length; vulva without flap; spermatheca 17–31 μm long; postuterine sac 7–24 μm long; spicules 21.5–23.5 μm long. Other specimens from New Zealand are identified as belonging to S. tuarua. A parthenogenetic population from Senegal is identified as belonging to S. ophioglossa and compared with type specimens from Mongolia and records of several other populations of S. ophioglossa. The generic diagnosis is emended and a key to the species of Stegelleta is provided.
Two new species of the mexicanus group of Vaejovis C.L. Koch are described from the Madrean pine-oak forests of the Sierra Madre Occidental in the state of Durango, Mexico. These species, Vaejovis sierrae sp. nov. and Vaejovis mcwesti sp. nov., are distinguished from each other and the only other species of the mexicanus group known from this mountain range, Vaejovis montanus Graham and Bryson, by morphometrics, carinal development of the pedipalps, granulation of the metasoma, and body size. A key to the species of the mexicanus group from
the Sierra Madre Occidental is provided.
Fifth instars for eight of the nine species of Microrhagus Dejean (Coleoptera: Eucnemidae: Melasinae:
Dirhagini), all endemic in the Nearctic region, are described and illustrated. Biological information is provided, along with a brief discussion of the larval morphology among the eight Nearctic species of Microrhagus. Adult descriptions
are provided for four new species. These new species include Microrhagus breviangularis new species, Microrhagus carinicollis new species, Microrhagus lecontei new species and Microrhagus opacus new species.
A checklist and identifi cation keys for both adults and most known larvae are provided.
The genus Bembidion Latreille (Carabidae: Bembidiini) is reviewed for New Zealand. Thirty-six species-group taxa are recognized. Seven species are described as new: Bembidion (Zecillenus) karikari new species, Bembidion (Zecillenus) puponga new species, Bembidion (Zecillenus) tepaki new species, Bembidion (Zecillenus) waimarama new species, Bembidion (Zemetallina) bullerense new species, Bembidion (Zemetallina) mangamuka new species, Bembidion (Zemetallina) waiho new species. The taxonomic status of two species-group taxa is changed (valid names listed after equal sign): Bembidion (Zeactedium) orbiferum giachinoi Toledano, 2005 = Bembidion (Zeactedium) giachinoi Toledano, 2005; Bembidion (Zeperyphodes) nesophilum Broun, 1886 (previously synonymized with Bembidion (Zeperyphodes) callipeplum Bates, 1878) is resurrected from synonymy. A new synonymy is established (valid name listed after equal sign): Bembidion (Ananotaphus) rotundicolle eustictum Bates, 1878 = Bembidion (Ananotaphus) rotundicolle Bates, 1874. A concise review of the taxonomy of all taxa is provided.
Descriptions, identifi cation keys, illustrations of male genitalia, habitus photos, as well a s distributional data and
maps are given. Extensive information on ecology, biology, dispersal power, and collecting techniques is included for each species.
Two new species of Aulonothroscus Horn are described from The Bahamas and a third species is newly reported. Aulonothroscus inawa new species is described from Great Inagua and Aulonothroscus sibateo new species is described from Eleuthera. Aulonothroscus convergens (Horn) is reported from Andros, providing an island and new country record. These are the fi rst Aulonothroscus identifi ed to species from the Lucayan Archipelago and from a West Indies locality other than Guadeloupe. A key to the species of Bahamian Aulonothroscus is provided.
Within the tribe Coelidiini, subfamily Coelidiinae (Cicadellidae: Hemiptera), fragmentation of the genera Calodia Nielson, Olidiana McKamey and Taharana Nielson established the following 13 new genera: Cladolidia, type-species, Lodiana cladopenis Zhang; Creberulidia, type-species, Calodia paucita Nielson; Glaberana, type-species, Glaberana spadix, sp. nov.; Hamusolidia, type-species, Hamusolidia introrsa, sp. nov.; Hiatusorus, typespecies, Taharana schonhorsti Nielson; Laosolidia, type-species, Laosolidia complexa, sp. nov.; Orbisolidia, typespecies, Calodia spinocava Nielson; Singillatus, type-species, Lodiana furcata Nielson; Trinoridia, type-species, Trinoridia calcaris, sp. nov.; Tripesidia, type-species, Calodia warei Nielson; Tumidorus, type-species, Lodiana nielsoni Zhang; Webbolidia, type-species, Taharana webbi Nielson and Zhangolidia, type-species, Lodiana polyspinata Zhang. Nineteen genera in the tribe are treated.
The following 62 new species in 12 genera are described, illustrated and photographed: Calodia bicompressa (India); C. birama (Philippines); C. propennata (India); C. sichuanensis (China); C. sinuata (Laos); C. vincula (China, Vietnam); Creberulidia corniger (Laos); C. inflata (Thailand); C. multipenicula (Cambodia); C. ordospinosa (Thailand); C. penicula (Thailand); Glaberana ampla (Thailand); G. dentilamina (Thailand); G. longilamina (Thailand); G. penita (Laos); G. spadix (Laos); G. stylafurcata (Indonesia); Hamusolidia introrsa (Laos); Hiatusorus aviformus (Laos); H. robustus (China); H. supraspinosus (Thailand); Laosolidia complexa (Laos); L. tuberis (Laos); L. longiserrata (Laos); Olidiana tuberis (Vietnam); O. bispiculata (Laos); O. filiata (Thailand); O. implicata (Thailand); O. inaequabilia (Thailand); O. lata (Laos); O. parafringa (Laos); O. pennata (Laos); O. tonkinensis (Vietnam); O. vincula (Vietnam); Singillatus gracilius (Indonesia); S. ventrospinatus (India); Taharana abstrusa (Thailand); T. angusta (Vietnam); T. biavicula (Thailand); T. biunca (Thailand); T. brevicutata (Thailand); T. caverna (Malaysia); T. exiquitas (Thailand); T. forcipia (Thailand); T. gracilata (Thailand); T. incisura (Thailand); T. intimacalcara (Thailand); T. lacertosa (Thailand); T. mediolata (Thailand); T. minutura (Thailand); T. oblongiserrata (Laos); T. subspinata (Thailand); T. sublamina (Thailand); T. phetchahabunesis (Thailand); T. protriangulata (Thailand); T. subtumida (Thailand); T. truncata (Thailand); Trinoridia calcaris (India); T. trifida (Malaysia); Tripesidia kubani (Laos); Webbolidia kristenseni (Thailand); W. magna (Laos).
Taxonomy of all the genera is elucidated with a revised key to genera and species. The following formerly suppressed species are herein reinstated: Olidiana (Lodiana) flavofasciana Li, 1989, Olidiana (Lodiana) nigritibiana Li, 1987, Olidiana rufofasciana Li and Wang, 1989 and Webbolidia (Taharana) uniaristata Zhang, 1990. The following 3 species are new junior synonyms: Calodia flavinota Cai and Kuoh, 1993: 220 [= Calodia patricia (Jacobi), 1944:49], Olidiana yangi McKamey 2006: 502 [= Lodiana (Olidiana) hamularis Xu, 2000: 220] and Taharana yinggenensis Zhang and Zhang, 1994: 96 [= Taharana (Coelidia) sparsa (Stål) 1854: 254]. Taharana hainana Zhang 1994: 132 is a nomen nudum based on the same name in Zhang’s thesis (1988) which name was cited later by Li and Wang 1991: 275. Lodiana hainana Cai and He 2002: 139 is also a nomen nudum. A replacement name proposed herein is caii, nom. nov. in the genus Olidiana; Zhang’s 1994: 71 illustrations of subgenital plate (I) and aedeagus (M) of “fasciana Li” does not appear to represent the respective illustrations in Li 1991: 357 and may represent a new species in the genus Glaberana. The name of Olidiana nigridorsum (Cai and Shen) is changed to Olidiana nigridorsa (Cai and Shen) to agree with gender. Among 12 genera, 102 species are proposed in new combinations. Lodiana reductusi Xu and Kuoh, 1997 and Lodiana spicata Xu and Kuoh, 1997 are declared incertae sedis after attempts failed to locate the original descriptions and type specimens. Both species are provisionally assigned to the genus Olidiana. Two syntype specimens of Jassus egregius Schumacher, previously thought to be lost, were located in the Senckenberg Deutsches Entomologisches Institut, Müncheberg, Germany. The male specimen is designated lectotype herein. Six species in the Olidiana brevis (Walker) interspecific variation complex are elucidated with illustrations of male genitalia features.
New records, an updated checklist and a synoptic catalogue are also provided. All taxa including 264 valid species and 304 names are indexed.
An updated list of the mosses of the Inner Seychelles is given based on the previous literature and collections of the first author in 2008. It includes data on the frequency of species as well as distributional data for the individual islands. The moss flora of the islands is characterized. Campylopus brevirameus Dixon is regarded as synonym of C. julaceus ssp. arbogastii (Renauld & Cardot) J.-P.Frahm. Brachymenium dicranoides, Bryum alpinum, Campylopus flaccidus, C. flexuosus, Ectropothecium brachycladulum, E. chenagonii and E. perrotii are reported as new to the Seychelles. Garckea flexuosa, Syrrhopodon involutus and S. prolifer are reported as new to La Digue, Bryum leptospeiron, Brachymenium exile and Calymperes afzelii as new to Praslin.
Contribution to the bryophyte flora of India: the Aralam
Wildlife Sanctuary in the Western Ghats
(2009)
The bryophyte flora of the Aralam Wildlife Sanctuary in the Western Ghats of India is catalogued for the first time. The catalogue consists of 116 taxa (89 mosses, 27 liverworts), of which two are new for India (Plagiochila singularis, Vesicularia dubyana), 21 species are newly reported for Peninsular India (Clastobryopsis planula var. delicata, Barbella chrysonema, Brachymenium leptophyllum, Brachythecium rutabulum, Cololejeunea longifolia, Cyathodium tuberosum, Dicranella amplexans, Didymodon vinealis, Duthiella wallichi, Fabronia assamica, Haplocladium microphyllum, Himantocladium cyclophyllum, Homalia trichomanoides var. trichomanoides, Isopterygium serrulatum, Leskea perstricta, Lopholejeunea kashyapii, Leptotrichella assamica, Macromitrium turgidum, Rhynchostegium hookeri, Splachnobryum assamicum, Thamnobryum siamense) and another 14 species are new for Kerala State (Atrichum pallidum, Chionostomum rostratum, Claopodium prionophyllum, Cololejeunea lanciloba, Cyathophorum adiantum, Dicranella divaricata, Entodontopsis wightii, Fissidens pellucidus, Glossadelphus glossoides, Isopterygium lignicola, Leucodon secundus, Neckeropsis exserta, Plagiochila bischleriana, Timmiella anomala). The species Jungermannia obliquifolia has also been reported as a taxon new to India (Nair et al. in press (a) from this area.
Three new species of Pselnophorus are described from the Nearctic region. Pselnophorus chihuahuaensis Matthews, Gielis, and Watkins, Pselnophorus hodgesi Matthews, Gielis, and Watkins, and Pselnophorus kutisi Matthews, Gielis, and Watkins, are described and distinguished from the only previously named Nearctic congener Pselnophorus belfragei (Fish). Illustrations of the adults and male and female genitalia are provided along with a key to males.
Besides the two species at present known belonging to the genus Trichonotuloides Balthasar (T. glyptus (Bates) and T. latecrenatus (Bates)), two new Mexican species, T. alfonsinae and T. hansferyi, are herein described (Coleoptera: Scarabaeidae: Aphodiinae). The complete set of fi gures is supplied for all taxa herein dealt with.
Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae)
(2014)
The North American species of the genus Eusphalerum Kraatz (Coleoptera, Staphylinidae, Omaliinae) are revised. The taxonomic history, natural history, geographical distribution of the genus, characters, species groups, diversity, and biogeography of North American species are presented. Two main phylogenetic lineages and 13 species groups are provisionally recognized. The following new synonymies are proposed: Eusphalerum farrarae (Hatch, 1944) = E. lawrencei Hatch, 1957; Eusphalerum californicum (Fauvel, 1878) = E. atriventre (Casey, 1894), = E. nigerrimum (Casey, 1894), = E. gilvipenne (Casey, 1894), = E. dichroum (Fall, 1922), = E. bonnelli (Hatch, 1944), = E. lunae Hatch, 1957; Eusphalerum fraternum (Casey, 1894) = E. minskae (Hatch, 1944); Eusphalerum rugulosum (Mäklin, 1853) = E. grayae (Hatch, 1944); Eusphalerum orientale (Bernhauer, 1912) = E. frosti (Bernhauer, 1928). The following lectotypes are designated: E. subangulatum (Casey), E. californicum (Fauvel), E. gilvipenne (Casey), E. diversicolle (Casey), E. convexum (Fauvel), E. fraternum (Casey), E. horni (Fauvel), E. orientale (Bernhauer), E. pothos (Mannerheim), and E. punctatum (Casey). The following new species are described: Eusphalerum pilosum (California); E. klimaszewskii (British Columbia); E. chatzimanolisi (California); E. carolinensis (Kentucky, Missouri, North Carolina, Ohio, Pennsylvania, Tennessee, Virginia); E. caterinoi (California); E. luteipes (California); E. thayeranum (Alberta, British Columbia, Idaho, Indiana (doubtful record), Oregon, Washington); E. margaretae (Tennessee); E. newtoni (British Columbia, Oregon, California); E. parvispiculum (California, Oregon); E. uncinatum (British Columbia, California, Oregon, Washington). Eusphalerum lapponicum (Mannerheim, 1830) is excluded from the North American fauna. The following new combination is proposed: Xylodromus segmentarius (Mäklin, 1852: 322) (ex Omalium), wrongly attributed to Eusphalerum in the literature. Omalium marginatum Say, 1832 is considered a doubtful species, probably not Eusphalerum. A key to the 27 recognized North American Eusphalerum species and a catalog of the species are provided.