Insecta Mundi
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578
Tragosoma soror (Coleoptera: Cerambycidae: Prioninae), a new Nearctic species, is described and compared to its North American congeners. Tragosoma harrisii LeConte, revised status, and Tragosoma spiculum Casey, revised status are both resurrected from synonymy and considered valid species distinct from T. depsarium (Linnaeus). The Mexican T. nigripenne Bates is confi rmed as a valid species. Tragosoma chiricahuae Linsley, new synonym, is placed in synonymy under T. spiculum Casey, and T. sodale Casey, new synonym, T. parvicolle Casey, new synonym, and T. repens Casey new synonym are placed in synonymy under T. harrisii. Tragosoma repens is a junior objective synonym of T. parvicolle. A lectotype is designated for T. sodale. An updated synonymical and distributional checklist of Tragosoma of the world is provided.
577
Two new genera and species of tiger beetles from Baltic amber (Coleoptera: Carabidae: Cicindelinae)
(2017)
Two fossil tiger beetle species (Coleoptera, Carabidae, Cicindelinae) are described from Eocene Baltic amber using light microscopic and X-ray microscopic techniques. Both species are considered representatives of the subtribe Iresina Rivalier, 1971 due to the shared combination of character states: glabrous head, six labral and four suborbital setae, and glabrous pronotum. Palaeopronyssiformia groehni Wiesner, Will, and Schmidt, new genus, new species, is characterized by a glabrous and furrowed head with six labral setae, large eyes, presence of two supraorbital setae on each side, mandibles with two teeth of the incisor region, and a glabrous and furrowed pronotum. Palaeoiresina cassolai Wiesner, Will, and Schmidt, new genus, new species, is characterized by a unicolored, undentated labrum, mandibles with two teeth of the incisor region, glabrous head with six labral setae, two clypeal setae, two supraorbital setae on each side, and a glabrous pronotum, mesepisternum, mesepimeron, and metepisternum. The species described here represent the only known tiger beetle fossils preserved in Baltic amber.
576
575
Redescription of adult and observations of the weevil Revena plaumanni Bondar (Coleoptera: Curculionidae), preying on seeds of the palm tree Butia eriospatha (Mart. ex Drude) Becc. (Arecaceae), as well as oviposition and larval behavior, pupation site, and adult emergence are reported for the fi rst time in Brazil. Photos of host plant, immature forms (egg, larva and pupa) and adult are given. Illustrations of male and female genitalia are also provided.
574
The New World genus Chariessa Forster (Coleoptera: Cleroidea: Cleridae) is revised and includes C. catalina Opitz, new species, C. elegans Horn, C. dichroa (LeConte), C. floridana Schaeffer, C. pilosa (Forster), C. texana Wolcott, C. ramicornis Perty, C. vestita (Chevrolat), and C. duponti (Spinola). Enoplium pilosa var. marginata Say is synonymized with Chariessa pilosa Forster. Lectotypes are designated for C. pilosa (Forster), C. ramicornis Perty, and C. vestita (Chevrolat). Available information indicates that Chariessa adult and immature individuals are predatory on lignicolous insects with a particular affinity for cerambycids and buprestids that infest species of oak. It is postulated that Pleistocene speciation generated the North American components of Chariessa with more ancient southern species generated during the Middle Tertiary; after closures of the Middle American portals and orogeny of the South American Andes. Included in this treatise is a discussion of natural history, key to species, narratives of zoogeography and phylogeny, one diagram of a phylogenetic tree, 35 line drawings, eight SEM micrographs, twelve habitus photographs, nine photographs of male genitalia, and five distributional maps.
573
With an incident in Palo Duro Canyon, Texas, USA, Scolopendra heros Girard (Chilopoda: Scolopendromorpha: Scolopendridae) becomes the third centipede species known to prey on bats; S. gigantea Linnaeus and S. viridicornis Newport have been so documented in Venezuela and Brazil, respectively. The Texas predation was interrupted by the predator/prey pair’s falling around 15–20 m from the canyon wall and, perhaps also, by human presence where they landed. The centipede uncoiled and retreated to shelter under a nearby rock and, after initial immobilization, so did the bat.
572
Abstract. Six new species of Coleoxestia Aurivillius, 1912 (Cerambycidae, Cerambycinae, Cerambycini) are described: C. clarkei from Bolivia; C. chemsaki and C. eyai from Peru; C. fragosoi and C. rafaeli from Brazil; and C. hovorei from Ecuador. Coleoxestia rachelae Eya and Chemsak is also newly recorded from Peru (new country record). Dorsal, ventral and lateral illustrations along with illustrations of various other structures are provided for each of the new species. In some cases, existing key couplets by Eya and Chemsak (2005) or Martins and Monné (2005) are modifi ed to help separate a new species from similar existing species. A reproduction of the general descriptive terminology for the head, prothorax and antennomeres of Coleoxestia, from Fragoso (1993), is also included in the illustrations.
571
A trimaculate male of the diplopod genus Apheloria Chamberlin (Polydesmida: Xystodesmidae/-inae: Apheloriini) from 1.3 km (0.8 mi) west of McKenney, Dinwiddie County (Co.), Virginia, is designated the Neotype of Julus virginiensis Drury 1770, thereby stabilizing the earliest name for a North American milliped and authenticating its prior assignment to this taxon. The existing concept of Apheloria is accepted in the absence of a revisionary treatment, and a modern description of A. v. virginiensis with gonopod drawings and color photos is provided. Drury’s original account and his letter to the Virginian who sent him the original specimens are quoted verbatim to eliminate future library searches. The specific name has been associated with at least three genera, and its confusing history is clarified by summarizing works in each. Authentic localities, mapped to the extent now possible, reveal a distribution south of the James River in piedmont and coastal Virginia that extends southwestward to the Blue Ridge foothills and at least as far south in North Carolina (NC) as Greensboro, the “Triangle” (Raleigh/Durham/Chapel Hill region), and Albemarle Sound in the east. Based on the holotypes, A. aspila and A. tigana, both by Chamberlin, are placed in synonymy under A. v. virginiensis (syns. nov.), and although its status is still under review, A. waccamana Chamberlin, whose type locality is Lake Waccamaw, Columbus Co., in southeastern NC, may be the correct name for today’s A. tigana. All samples so labeled must be reexamined for misidentifications of A. v. virginiensis.
570
A detailed study of the holotype of Sphecomyrma canadensis Wilson, 1985 (Hymenoptera: Formicidae) from Canadian amber has led to the conclusion that the specimen belongs to a new genus, here named Boltonimecia gen.n. Since the taxonomy of stem-group ants is not well understood, in order to find the taxonomic position of this genus, it is necessary to review the classifi cation of stem-group ants in a study of their relation to crown-group ants. In the absence of data for traditional taxonomic approaches, a statistical study was done based on a morphometric analysis of antennae. Scape elongation is believed to play an important role in the evolution of eusociality in ants; however, this hypothesis has never been confirmed statistically. The statistical analysis presented herein lends support to the view that antennal morphology reliably distinguishes stem-group ants from crown-group ants, to determine whether a species belongs to one or the other group. This, in turn, may indicate a relationship exists between eusociality and scape elongation. A review of Cretaceous records of ants is made and the higher classification of Formicidae with definitions of stem and crown groups is proposed. Newly obtained data are discussed focusing particularly on the origin, evolution and diversity of ants.
569
A primitive subfamily of false click beetles (Coleoptera: Eucnemidae: Phlegoninae) distributed primarily in the Neotropical region is revised. Euryphlegon gen. nov. is described from Belize in Central America. New species include: Phlegon chiriquiensis sp. nov. (Panama), Phlegon panamensis sp. nov. (Panama), Euryphlegon jacqueschassaini sp. nov. (Panama) and Euryphlegon parallelus sp. nov. (Belize). Phlegon herculeanus (Lacordaire) stat. res. is resurrected from synonymy with Phlegon buqueti Laporte. One new combination is proposed: Euryphlegon degallieri (Muona) (Phlegon). Based on a number of observed external character traits, Euryphlegon is placed in Orodotini Muona, 1993 within Macraulacinae Fleutiaux, 1922. Identifi cation keys are provided for species of Phlegon and Euryphlegon in the Neotropical region. The relationships among Phlegon, Euryphlegon, Euryptychus LeConte and other groups within Echthrogasterini Cobos, 1964 and Orodotini are discussed.
568
New Bolivian Rhinotragini (Coleoptera, Cerambycidae, Cerambycinae) are described: three species of Phygopoda Thomson, 1864 (P. longiscopifera sp. nov., P. boliviensis sp. nov. and P. chaquensis sp. nov.); and one species of Phygopoides Peñaherrera-Leiva and Tavakilian, 2007 (P. maxwelli sp. nov.). Two Brazilian species of Neophygopoda Melzer, 1933 are transferred to the genus Phygopoda: P. exilis (Melzer, 1933) comb. nov. and P. agdae (Martins, Galileo and Santos-Silva, 2015) comb. nov. All the species are illustrated, and a key to the Bolivian species of Phygopoda and host flower records are provided.
567
The taxonomic history of the rhinotragine genera Phygopoda Thomson, 1864 and Pseudophygopoda Tavakilian and Peñaherrera-Leiva, 2007 (Coleoptera: Cerambycidae: Cerambycinae) are discussed, and evidence is presented to suggest that some recent taxonomic changes made by Carelli and Monné (2015) were unjustified. Consequently, Phygopoda nigritarsis Gounelle, 1911 is moved to the genus Neophygopoda Melzer, 1933, creating the new combination Neophygopoda nigritarsis, the genera Panamapoda Clarke, 2014 and Paraphygopoda Clarke, 2014 are revalidated, and the species Paraphygopoda viridimicans (Fisher, 1952) and Paraphygopoda nappae Clarke, 2014 are also revalidated.
566
Dipropus tequesta Johnson new species (Coleoptera: Elateridae) is described from southern Florida. Dipropus fuscus (LeConte) is a new synonym of D. soleatus (Say), and D. granosus (Fall) is a new synonym of D. asper (LeConte). The fl ightless female of D. asper is described and provides the fi rst report of brachyptery and endogean habits in Dipropus. A key to the species of the eastern United States and a new checklist of species for the country are provided.
564
563
562
The primary types of Onciderini Thomson, 1860 (Coleoptera: Cerambycidae: Lamiinae) of the Museum of Comparative Zoology (MCZ), Harvard University, are catalogued and illustrated. Data on the original combination, current name, gender, and type locality are verifi ed and presented. There are 19 primary types of Onciderini including seven in Oncideres Lacordaire, 1830; two in Hesychotypa Thomson, 1868; and two in Hypsioma Audinet-Serville, 1835. Of the 19 primary types, 18 were described by L. S. Dillon and E. S. Dillon. A brief history of the Coleoptera collection at the MCZ is also presented.
561
A number of nomenclatural issues regarding Neotropical Scarabaeoidea are addressed. Athyreus soesilae Makhan, 2008 is synonymized with Neoathyreus (Neoathyreus) excavatus (Laporte, 1840) (Geotrupidae: Bolboceratinae: Athyreini). Scalmus Zang, 1905 is replaced with its senior synonym Neleuops Kuwert, 1891, nec Neleurops Kuwert, 1891, and the four species it presently contains are thus transferred to Neleuops (Passalidae: Passalinae: Passalini). The spelling of the name Oxysternon oberthueri Olsoufi eff, 1924, a junior synonym of Oxysternon (Oxysternon) conspicillatum (Weber, 1801), is fi xed (Scarabaeidae: Scarabaeinae: Phanaeini). The year of description of Oxysternon (Oxysternon) festivum (Linnaeus, 1758) is corrected, its existing lectotype invalidated, and a new lectotype designated (Scarabaeidae: Scarabaeinae: Phanaeini). The correct spelling of Phanaeus (Phanaeus) prasinus trinidadens Arnaud, 2001 and Phanaeus (Phanaeus) prasinus joliyi Arnaud, 2001 is discussed (Scarabaeidae: Scarabaeinae: Phanaeini). The seniority of Chlorota aulica Burmeister, 1844 over Chlorota metallica Burmeister, 1844 is restored (Scarabaeidae: Rutelinae: Rutelini). The year of description of Rutela lineola (Linnaeus, 1758) is corrected and a lectotype is designated (Scarabaeidae: Rutelinae: Rutelini). Cyclocephala aulustjaorum nomen novum is proposed as a replacement name for the preoccupied name Cyclocephala brevis Höhne, 1923 (Scarabaeidae: Dynastinae: Cyclocephalini). The spelling of the generic name Harposceles Burmeister, 1847 is fi xed (Scarabaeidae: Dynastinae: Cyclocephalini). The incorrect subsequent spelling Stenocrates haackae Ratcliffe, 1977 is preserved (Scarabaeidae: Dynastinae: Cyclocephalini). The name Scarabaeus simson Linnaeus, 1767 is invalidated as a synonym of Megasoma actaeon (Linnaeus, 1758) (Scarabaeidae: Dynastinae: Dynastini). The spelling of the generic name Coelosis Hope, 1837 is found to be correct (Scarabaeidae: Dynastinae: Oryctini). The validity of Steyskal’s emendation of the name Megaceropsis quadridentata Dechambre, 1976 is confi rmed (Scarabaeidae: Dynastinae: Oryctini). The lectotype of Podischnus agenor (Olivier, 1789) is invalidated and a new lectotype is designated (Scarabaeidae: Dynastinae: Oryctini).
560
Chiquitano gen. nov. Chiquitano volcanesensis sp. nov., Compsibidion achiraensis sp. nov. and Compsibidion amboroensis sp. nov. (Coleoptera: Cerambycidae: Cerambycinae: Neoibidionini) are described from Bolivia. Notes on Rhysium Pascoe, 1866 and Rhysium bimaculatum Pascoe, 1866 are provided, and Brechmoidion separatum Martins and Galileo, 2007 is transferred to Rhysium. Keys to species of Compsibidion Thomson, 1864, Brechmoidion Martins, 1969 and Rhysium Pascoe, 1866 are also provided.
559
The Antillean fauna of Mantispidae (Neuroptera) consists of six species in four genera, all in the subfamily Mantispinae. Two new species are here described: Leptomantispa antillesensis Hoffman, n. sp., is described from Puerto Rico, Anguilla, British Virgin Islands (Grand Camanoe, Guana, Tortola), United States Virgin Islands (St. Croix, St. John, St. Thomas), Dominica, and Guadeloupe, and Leptomantispa hispaniolaensis Hoffman, n. sp. is described from Hispaniola (Dominican Republic). Mantispilla zayasi Alayo is synonymized under Leptomantispa pulchella (Banks), new synonomy, and Mantispilla taina Alayo is synonymized under Zeugomantispa minuta (Fabricius), new synonomy. Climaciella cubana Enderlein is recorded for the fi rst time from Hispaniola (Dominican Republic). Dicromantispa sayi (Banks) is newly reported from the Cayman Islands (Cayman Brac and Little Cayman), and Hispaniola (Dominican Republic). Zeugomantispa minuta (Fabricius) is now recorded from the Bahamas (Andros) and Hispaniola (Dominican Republic).
558
Six new species of false click beetle (Coleoptera: Eucnemidae) are described. These new species are: Fornax dixiensis sp. nov. (Florida, USA), Dromaeolus comayaguiensis sp. nov. (Honduras), Asiocnemis bicolor sp. nov. (Ecuador), Miruantennus chinensis sp. nov. (Yunnan prov., China), Miruantennus cuneiformis sp. nov. (Malaysia) and Nematodes africanus sp. nov. (Côte d’Ivoire). Identifi cation keys are provided for species of Fornax in the Nearctic ecozone and Miruantennus in the Palearctic and Indo-Malayan ecozones. A new key to include all the genera within the tribe Nematodini Leiler is also provided.
557
Conognatha iris iris Olivier (Coleoptera: Buprestidae) is recorded for the fi rst time for Venezuela. The record is based on a single female specimen from Caicet hill, Amazonas state. A map of the known distribution of the taxon is provided
555
The monotypic clerid genus Ohanlonella Rifkind (Coleoptera: Cleridae: Clerinae), described from Oaxaca, is reported for the first time from the state of Veracruz. The recently discovered population shows distinctly different coloration from the type series of O. esperanzae Rifkind, but appears to be otherwise identical. The Veracruz morph is briefly characterized and illustrated, and its significance is discussed.
554
A new species of Ctilodes Murray, 1864 (Coleoptera: Nitidulidae) was recovered in material from Vietnam during a large study of Carpophilinae. A description and detailed diagnosis of Ctilodes clinei Powell and Duffy new species is presented here along with a key to identify all currently known species of Ctilodes.
553
Coprophagy and probable saprophagy are reported for larvae of two species of chrysaugine moths (Lepidoptera: Pyralidae). Larvae of Parachma ochracealis Walker are found in rotten pine wood and mulch in North-Central Florida. Larvae of Basacallis tarachodes (Dyar) inhabit feces of an unidentified small mammal in a cave in Central Florida and seem to be troglophilic. These behaviors are compared to similar ones in Neotropical chrysaugines such as Humiphila Becker and Cryptoses Dyar. Saprophagy and coprophagy are predicted to be more general habits among Chrysauginae.
552
Distributional records of the 30 tiger beetle species and subspecies (Coleoptera: Carabidae: Cicindelinae) known for Brunei Darussalam are given together with habitus photos for 21 species. Neocollyris (Neocollyris) labiomaculata (Horn, 1892), Neocollyris (Neocollyris) emarginata (Dejean, 1825), Therates spectabilis fl avissimus Brouerius van Nidek, 1957, Heptodonta analis s. str. (Fabricius, 1801), Cosmodela velata (Bates, 1872), Lophyra (s. str.) fuliginosa (Dejean, 1826), Cylindera (Leptinomera) fi ligera (Bates, 1878), Myriochila (s. str.) specularis brevipennis (Horn, 1897), Abroscelis tenuipes araneipes (Schaum, 1863) and Callytron doriai (Horn, 1897) are reported for the fi rst time for the Sultanate.
551
Scale insects (Hemiptera: Coccoidea) were collected on oak trees (Quercus spp.; Fagaceae) in Korea and identifi ed as Asterodiaspis luteola (Russell) (Asterolecaniidae) and Nidularia japonica (Kuwana) (Kermesidae), which are newly documented in the Korean fauna of scale insects. Characteristics of these species are briefl y given and illustrative photographs and information on their distribution and hosts are provided.
550
Cixidia fusca and Synecdoche impunctata (Hemiptera: Achilidae) are reported from Missouri for the first time, new state records. Ecological and trapping information is also provided.
549
548
We describe a new butterfl y species, Anaeomorpha mirifi ca Simon and Willmott, n. sp. (Lepidoptera, Nymphalidae, Charaxinae), from premontane rain forest of the Chocó region of northwestern Ecuador. This represents the second known species and the fi rst record outside of the Amazon basin for this taxonomically and biologically enigmatic genus. Although the two species are not known to be sympatric, we identifi ed 21 characters in the external color pattern, wing shape and male genitalia that together support distinct species status. Most notably, the new species possesses an ocellus in the ventral hind wing tornus, a character which occurs in the Neotropical Charaxinae only in the genus Prepona Boisduval, 1836. A mean divergence of 6.8% in the COI ‘barcodes’ between the two species underlines their taxonomic distinctness.
547
The eight species in the genus Tomarus Erichson (Coleoptera: Scarabaeidae) in Argentina, Chile, and Uruguay are reviewed. Tomarus roigjunenti new species and Tomarus spinipenis new species are described from Argentina. We include a key to species, representative habitus illustrations for all species, character illustrations, and distribution maps for each, as well as commentary about the natural history and distributions for each species. Diagnostic characters are discussed for each species, and species relationships are hypothesized based on the analysis of internal and external morphological characters. The male of T. bidentulus (Fairmaire) is described for fi rst time. The following taxonomic changes are made: Tomarus guianucai Dechambre and Lumaret, 1985 is a new junior synonym of Tomarus rubripes (Boheman, 1858), which was formerly and incorrectly cited as occurring in Argentina.
545
Descriptions and illustrations of the triungulin, 5th instar, and prepupal larval forms for Nematodes penetrans (LeConte, 1852) (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) are given. The descriptions are based on triungulins collected in a plastic vial as well as 5th instars and prepupal larval stages collected from hard sections of sugar maple (Acer saccharum Marshall; Aceraceae) logs and limbs. Adults were reared from those pieces placed in plastic bags. Biological information is provided, based on literature search and personal observations.
Comparative studies between larvae of N. penetrans and the European species, Nematodes fi lum (Fabricius, 1801) along with other Nearctic larval forms are briefl y discussed. This discovery represents the fi rst larval description for the genus in the Nearctic region.
544
543
Four new species of Chrysina Kirby (Coleoptera: Scarabaeidae: Rutelinae) from Guatemala and Honduras
(2017)
Four new species of the genus Chrysina Kirby (Coleoptera: Scarabaeidae: Rutelinae) are described from cloud forests of Guatemala and Honduras. The Guatemalan species are C. alexae n. sp. from Purulhá, Baja Verapáz in the central mountains and C. woodruffi n. sp. from Bulej, Huehuetenango in Los Cuchumatanes Mountains at elevations from 1500 to 2000 meters. The species from Honduras are C. antonkozlovi n. sp. from Celaque, Lempira and C. maishei n. sp. from El Güisayote, Ocotepeque, at elevations between 2000 and 2600 meters above sea level.
542
Species descriptions, keys to genera and species, and geographical distributions are presented for 43 species of the family Bruchidae (Coleoptera: Chrysomeloidea) for Chile. Of these species, seven are described as new:
Acanthoscelides aricae sp. nov., Lithraeus chillan sp. nov., L. comptus sp. nov., L. elguetai sp. nov., L. limari sp. nov., L. lonquimay sp. nov., and L. penai sp. nov. Eight species are endemic to Chile. A list of true host plants and floral records for those with known host associations is presented. Habitus photographs and drawings of pertinent body parts, including male genitalia, are provided. References pertaining to the previously described species are listed.
541
The genus Paraphloeolaemus Thomas (Coleoptera: Cucujoidea: Laemophloeidae) is described for two new Neotropical species, P. vorticosus Thomas, new species, and P. pterosiagon Thomas, new species. Diagnoses and illustrations are provided.
The following 16 species are transferred from Laemophloeus Dejean (s. l.) to Phloeolaemus Casey: Phloeolaemus anticus (Sharp, 1899: 518) [= Laemophloeus anticus Sharp, 1899], new combination; Phloeolaemus boops (Sharp, 1899: 517) [= Laemophloeus boops Sharp, 1899], new combination; Phloeolaemus castaneipennis (Grouvelle, 1876: 494) [= Laemophloeus castaneipennis Grouvelle, 1876: 494], new combination; Phloeolaemus championi (Sharp, 1899: 516) [= Laemophloeus championi Sharp, 1899], new combination; Phloeolaemus curtus (Grouvelle, 1876: xxxiii) [= Laemophloeus curtus Grouvelle, 1876], new combination; Phloeolaemus endomychus (Sharp, 1899: 519) [= Laemophloeus endomychus Sharp, 1899], new combination; Phloeolaemus hoplites (Sharp, 1899: 517) [= Laemophloeus hoplites Sharp, 1899], new combination; Phloeolaemus ignobilis (Sharp, 1899: 518) [= Laemophloeus ignobilis Sharp, 1899], new combination; Phloeolaemus impressus (Grouvelle, 1876: xxxiii) [= Laemophloeus impressus Grouvelle, 1876], new combination; Phloeolaemus lacerdae (Grouvelle, 1877: 211) [= Laemophloeus lacerdae Grouvelle, 1877], new combination; Phloeolaemus macrocephalus (Schaeffer, 1910: 214) [= Laemophloeus macrocephalus Schaeffer, 1910], new combination; Phloeolaemus punctulaticollis (Hetschko, 1929: 94) [= Laemophloeus punctulaticollis Hetschko, 1929], new combination; Phloeolaemus reitteri (Grouvelle, 1877: 210) [= Laemophloeus reitteri Grouvelle, 1877], new combination; Phloeolaemus semiflavus (Grouvelle, 1876: 497) [= Laemophloeus semiflavus Grouvelle, 1876], new combination; Phloeolaemus sharpi (Hetschko, 1929: 41) [= Laemophloeus sharpi Hetschko, 1929], new combination; Phloeolaemus straminipennis (Reitter, 1876: 47) [= Laemophloeus straminipennis Reitter, 1876], new combination; Phloeolaemus teapensis (Grouvelle, 1876: 494) [= Laemophloeus teapensis Grouvelle, 1876], new combination.
540
A new genus and two new species of Argentine Rhinotragini
(Coleoptera, Cerambycidae, Cerambycinae)
(2017)
Rhopalessa irwini sp.nov. and Rhinion parkeri gen. nov., sp. nov. (Coleoptera, Cerambycidae, Cerambycinae, Rhinotragini) are described from Argentina. A key to species of Rhopalessa is provided.
539
Six annotated lists are presented: A, a checklist of the butterflies (Lepidoptera, Papilionoidea) of Tobago (150 species); B, species for which there are no records in the last 80 years (49 species); C, species needing confi rmation from Tobago (5 species); D, species not accepted from Tobago (12 species); E, species which are likely to occur in Tobago, but have not been recorded (6 species); and F, species and subspecies recorded from Tobago, but not from Trinidad (2 species and 2 subspecies). Remarkably, 33% of the 150 recorded species have not been reported
in the last 80 years. While it is possible that some of these are not resident or have become extinct, it seems more likely that most have simply not been found in the last 80 years. The butterfly fauna of Tobago merits further study; year-round collecting in different habitats and areas, using a variety of techniques, will surely fi ll in many of the apparent gaps in our knowledge. Ouleus fridericus sheldoni ssp. nov. (Hesperiidae, Pyrginae) is described from Tobago, with illustrations of adults and male genitalia, and is compared to O. fridericus sinepunctis (Kaye) from Trinidad. Danaus plexippus tobagi A.H. Clark, 1941 is a syn. nov. of D. plexippus nigrippus (Haensch, 1909) (Nymphalidae, Danainae).
538
The female of Callichroma magnifi cum Napp and Martins, 2009 (Coleoptera: Cerambycidae: Cerambycinae: Callichromatini) is described and illustrated for the fi rst time. The geographical distribution of the species is expanded to the Colombian Caribbean and information on the collection site is provided.
537
The Republic of Panama currently has 21 recorded species of stoneflies, all in the genus Anacroneuria (Plecoptera: Perlidae). Herein, we record five species of this genus from the Mount Totumas Cloud Forest and Biological Reserve, in the upper reaches of the Río Chiriquí Viejo watershed. One of these species, A. plutonis (Banks), represents a new country record for Panama. These results are part of an ongoing effort to characterize the aquatic insect fauna of Panama, and to evaluate that country’s major watersheds.
536
A new genus, Dejunaleon Miller and Stange, is described with two new species: Dejunaleon loja Miller and Dejunaleon maculosus Stange (Neuroptera: Myrmeleontidae: Brachynemurini). A key to species is provided as well as distributional data. A description of the larva of Dejunaleon loja is given. A key to genera of the tribe Brachynemurini is provided based on larvae and adults.
535
Aphis elena Lagos-Kutz and Voegtlin, sp. nov. (Hemiptera: Aphididae), is described from specimens collected in Illinois, USA, on the North American native plant, Pycnanthemum virginianum (L.) T. Dur. & B.D. Jacks. ex B.L. Rob. & Fernald (Family: Lamiaceae). Both apterous and alate viviparae are described and illustrated, and a dichotomous key is provided to apterous and alate viviparae of the six species of Aphis that have been recorded on plants in the family Lamiaceae in North America. Neighbor-joining analysis of cytochrome oxidase 1 (Cox1) indicated a close relationship of the new species with Aphis monardae Oestlund, which also feeds on a member of Lamiaceae. The range of pair-wise distances for DNA barcoding of these species is 0.17–0.33%. The newly described Aphis elena is morphologically more similar to Aphis gossypii Glover than it is to Aphis monardae.
534
533
Females of Traumatomutilla André (Hymenoptera: Mutillidae) are organized into 14 species groups; 136 of the 138 Traumatomutilla taxa known from females were studied. Of these species groups, only the inermis species group was adequately established and diagnosed in the literature (Casal 1969). The remaining newly established species groups are as follows: americana, auriculata, bellica, bifurca, diabolica, gemella, indica, integella, juvenilis, quadrinotata, tabapua, trochanterata, and vitelligera. Diagnoses and lists of included species are provided for each species group. Eighty-three new country records are provided for 49 species.
532
Effective island conservation depends on thorough biodiversity surveys and species assessment. The U.S. Fish and Wildlife Service in coordination with Point Blue Conservation Science undertook a two-year insect survey of the Farallon Islands, California, in order to catalog current insect and spider species on the island and to gather information that will be used to examine prey dynamics to aid in the conservation of the endemic salamander Aneides lugubris (Hallowell). The report lists 11 insect orders representing 60 families, 107 genera and 112 insect species on Southeast Farallon Island. Holometabolous orders were the most represented on the island with Coleoptera and Diptera being the most abundant, followed by Lepidoptera and Diptera. One spider order was identified, representing six genera and six species.
531
530
Nine new species of Rhinotragini (Coleoptera: Cerambycidae: Cerambycinae) are described from the Neotropical region: Odontocera galileoae (Mexico); Odontocera mthomasi (Guatemala); Ecliptoides vandenberghei (Nicaragua); Eclipta ricei and Odontocera skelleyi (Costa Rica); Eclipta nearnsi (Panama); Odontocera stangei (Venezuela); Chariergodes lingafelteri and Ischasia martinsi (Costa Rica and Panama). Ischasia rufi na Thomson, 1864 is formally excluded from the fauna of Costa Rica and Panama, and Nicaragua is added as a new country record for Eclipta fi cta Bezark, Martins and Santos-Silva, 2013. Notes on Ecliptoides monostigma (Bates, 1869)
are provided in the “Diagnosis” of E. vandenberghei.
529
The genus Speorthus Chamberlin is revalidated for S. tuganbius Chamberlin and S. reyesi, new species (Diplopoda, Polydesmida, Macrosternodesmidae). The families Polydesmidae and Macrosternodesmidae (as they occur in North America) are briefly discussed, and their component North American genera listed. The validity of Macrosternodesmidae is reaffi rmed; the family is predominantly North American. Nearctodesminae, new status, is formally designated as a subfamily of Macrosternodesmidae. The superfamily Trichopolydesmoidea is synonymized with the superfamily Polydesmoidea, new combination.
528
This revision concerns a small group of Western Palaearctic Copris species (Coleoptera: Scarabaeoidea:
Scarabaeidae) distinguished by having three lateral teeth on the foretibae. According to the literature, this group consists of four taxa: Copris armeniacus Faldermann, 1835, C. felschei Reitter, 1892, C. pueli Mollandin de Boissy, 1905 and C. umbilicatus Abeille de Perrin, 1901. Copris armeniacus is herein deemed a species inquirenda, and C. felschei, for which a neotype is designated, is deemed valid. Copris umbilicatus is recorded for the fi rst time from Turkey. A key to all species of the genus Copris known from the Western Palaearctic is provided. Variability of the cephalic and pronotal armature, and morphology of the parameres are illustrated.
527
The melon fly, Bactrocera cucurbitae (Coquillett) (Diptera: Tephritidae), is a widespread, economically important tephritid fruit fly species. Bactrocera cucurbitae infests fruits and vegetables of a number of different plant species, with many host plants in the plant family Cucurbitaceae, but with additional hosts scattered across many other plant families. Although thought to be native to India, its distribution has spread throughout many countries in Oriental Asia, into a number of Pacifi c Island nations, and into Africa. The documented introductions into countries outside its native distribution show that this species could establish in other countries where it does not presently occur, particularly through the movement of infested fruit. As with other tephritid fruit fly species, establishment of B. cucurbitae can have signifi cant economic consequences, including damage and loss of food production, as well as requirements for implementation of costly quarantine treatments to permit export of commodities susceptible to infestation by B. cucurbitae and inspection of susceptible imported commodities. In order to avoid these adverse economic consequences, one needs to prevent the entry, establishment and spread of B. cucurbitae into a new habitat. To successfully achieve this, an accurate knowledge of the fly’s host plants is essential. Cognizant of this need, we prepared, and present here, a worldwide list of host plants for B. cucurbitae, with annotations on reported laboratory and fi eld infestation data. Overall, 136 plant taxa from 62 plant genera and 30 plant families are identifi ed as hosts of B. cucurbitae, based on reported fi eld infestation data. The predominant family, as expected, is Cucurbitaceae, with 56 plant taxa (41.2% of all host plant taxa) in which fi eld infestation by B. cucurbitae has been documented. The family with the 2nd highest number of documented infested plant taxa is Solanaceae, for which there are published fi eld infestation data for 20 plant taxa (14.7% of plant taxa for which there is documented fi eld infestation). Papers that list plants as hosts of B. cucurbitae based only on laboratory data, those that list plants as a host but do not report any fi eld infestation data, and those that report interception data add an additional 137 host plant taxa, representing a total of 80 genera and 39 plant families, 20 of which are additional plant families beyond those for which there is fi eld infestation data. These additional species must be considered “undetermined” hosts for which additional data are needed to document actual host status. This paper is a comprehensive documentation of host plants of the melon fly based on recorded infestations in laboratory and/or fi eld, interceptions at ports of entry, or “listing only” associations. Host records presented here will be used in vetting and developing the offi cial USDA list of host plants of the melon fly, which will be published by APHIS as a federal order.
526
525
The first myrmecophilous fl ea beetle genus (Myrmeconycha Konstantinov and Tishechkin, new genus) with four new species (M. erwini Konstantinov and Tishechkin, new species – Ecuador, M. gordoni Konstantinov and Tishechkin, new species – Brazil, M. pakaluki Konstantinov and Tishechkin, new species – Panama, and M. pheidole Konstantinov and Tishechkin, new species – Costa Rica) is described and illustrated. It is compared with fl ea beetles of the subtribe Disonychina (Coleoptera: Chrysomelidae: Galerucinae: Alticini) and may be easily differentiated based on the external and internal features, which include the waxy surface of the head and pronotum, reticulated surface of the pronotum, and four longitudinal ridges on each elytron.
524
A new species of Autostichidae, Eraina beechei nov. sp., is described from specimens collected in the Andean mountains of Ñuble, Biobío. Diagnostic characters for the species are presented; photographs of adults, illustrations of male and female genitalia, and a key for identifi cation of Eraina species are provided.
523
The species of Dipropus Germar (Coleoptera: Elateridae) of the west-central region of North America are taxonomically reviewed. Historical records of D. approximatus (Candèze), D. ferreus (LeConte), D. simplex (LeConte) and D. soleatus (Say) in the region are based on misidentifi cations. Dipropus pericu new species is described from Baja California Sur. Dipropus reinae new species, D. sonora new species and D. yaqui new species are described from southern Arizona, New Mexico and Sonora. Dipropus warneri new species is described from Arizona, New Mexico and Texas. Dipropus sus (Candèze) is reported from Morelos, Sonora and Sinaloa as new state records. Keys to the species of the region are provided. The historical value of a specimen of Elater soleatus Say in the Dejean collection is discussed and its lectotype designated. Ischiodontus oblitus Candèze is treated as an objective synonym of Dipropus soleatus.
522
Ptyoiulus Cook 1895, the dominant parajulid diplopod genus in the eastern United States (US), comprises two species – P. impressus (Say 1821), with a slanted, fl ared, circumferentially entire, and marginally serrate apical calyx on the anterior gonopod coxal process, and P. montanus (Cope 1869), n. comb., with a smooth, upright, cupulate calyx that is open caudad and coaxial with the process’ stem. The genus occupies a broad area between the Mississippi River and Atlantic Ocean extending from southern New England, Ontario, and Michigan to the Florida Panhandle and four small disjunct ones – from Montreal, Québec, to northern Vermont, along southwestern Lake Michigan in Wisconsin and Illinois; northeastern/eastcentral Arkansas, primarily in Crowley’s Ridge physiographic feature and beside the “bootheel” of Missouri; and a point locality in northeastern Louisiana just south of the Arkansas line. A male from Chester County (Co.), Pennsylvania, is designated as the neotype of Julus impressus, as is one from Durham Co., North Carolina, for J. montanus. As both species inhabit Montgomery Co., Virginia, the type locality of J. montanus, we exercise the right of first reviser, conserve the latter name, and assign it to the species with the smooth, cupulate, and coaxial calyx. We also exercise first reviser rights and assign Parajulus ectenes Bollman 1887 to this form, thereby relegating it to synonymy under Ptyoiulus montanus. Other new synonymies include Ptyoiulus georgiensis Chamberlin 1943 under P. impressus and P. coveanus Chamberlin 1943 under P. montanus. Both Ptyoiulus and P. impressus are projected for Delaware and Rhode Island and newly reported from Québec, Connecticut, District of Columbia, Maryland, Mississippi, South Carolina, Vermont, West Virginia, and Wisconsin, and the genus and species, respectively, are newly documented from Louisiana and Arkansas; P. montanus is newly cited from Alabama, Arkansas, Georgia, Mississippi, and South Carolina. Ptyoiulus impressus occupies every state except perhaps Louisiana and is the only species in areas that were inundated during the Cretaceous and glaciated during the Pleistocene; by contrast, P. montanus inhabits a relatively narrow east/west transect through the center of the generic range. Their distribution patterns suggest an old species, montanus, being actively displaced by the younger and more successful impressus. The decurvature of the epiproct in uroblaniulinines appears to increase with age and developmental stage. A key is presented to parajulid familygroup taxa in the US and Canada east of the Rocky Mountains.
521
520
519
Cnestus mutilatus (Blandford) (Coleoptera: Curculionidae: Scolytinae) is reported from Pennsylvania for the fi rst time, new state record. Specimens were collected using baited Lindgren funnels as early as 2013. Within Pennsylvania, C. mutilatus is now reported from Berks, Bucks, Lehigh, Montgomery, and York Counties.
518
Five new species of Bakerius Bondar (Hemiptera: Aleyrodidae: Aleurodicinae) are described and illustrated from the Americas and Vietnam based on the adult, nymph, and pupal stages: Bakerius asiaticus, Bakerius colombianus, Bakerius hondurensis, Bakerius leei and Bakerius peruvianus. The following six species: Bakerius attenuatus Bondar 1923, Bakerius calmoni Bondar 1928, Bakerius marmoratus (Hempel 1923), Bakerius phrygilanthi Bondar 1923, Bakerius sanguineus Bondar 1928, and Bakerius sublatus Bondar 1928 are re-described. An identification key to the New World genera of the subfamily Aleurodicinae, and a key to the adults and the puparia of Bakerius species are provided.
517
516
Himaloaesalus gaoligongshanus Huang and Chen, new species (Coleoptera: Lucanidae: Aesalinae) is described from the Gaoligongshan Mts., Yunnan, China. It is the fi fth species of the genus Himaloaesalus Huang and Chen. This new species is similar to the Himalayan species Himaloaesalus himalayicus Kurosawa and H. saburoi Araya et al., from which it is distinguished. The male and female genitalia of all the known species of Himaloaesalus are illustrated. Dorcus yongreni Huang and Chen, new species (Coleoptera: Lucanidae: Lucaninae) is described from Ruili area, Yunnan, China. It belongs to the elegans group (Huang and Chen 2013). This new species is similar to the Indian species Dorcus apatani (Okuda and Maeda), new combination, originally described in Digonophorus Waterhouse. The male genitalia of both species are compared and illustrated.
514
Four species of the genus Enoclerus (Coleoptera: Cleridae) are reported to prey on chemically protected Coccinellidae in North America. The possibility that aposematically colored Enoclerus also mimic prey ladybird species is discussed, and the further possibility that clerid predators acquire chemical protection through sequestration of ladybird prey’s toxic alkaloids is suggested as an important avenue of investigation. The data presented are primarily based on photographs taken by non-specialists, discovered through Internet search. The crowdsourcing of natural history observations can reveal aspects of animal behavior heretofore unreported and even unsuspected.
513
512
Two new species of Eburiini (Coleoptera, Cerambycidae, Cerambycinae) are described from Mexico (Guerrero): Susuacanga boteroi; and Eburia (Eburia) girouxae. Both are included in previous keys.
511
The Republic of Panama currently has 300 recorded species of Trichoptera distributed among 14 families. Herein we add 42 new country records for Panama, including one new family (Anomalopsychidae) and three new genera (Anomalopsychidae: Contulma; Hydroptilidae: Byrsopteryx and Cerasmatrichia). The newly recorded caddisfly taxa increase Panama’s total known fauna to 342 species, distributed among 15 families and 50 genera. These results are part of an ongoing effort to characterize the caddisfly fauna of Panama, and to evaluate that country’s major watersheds (cuencas).
510
A new species of Neostenoptera (Diptera: Cecidomyiidae: Winnertziinae) from eastern North America
(2016)
A new species of paedogenetic gall midge, Neostenoptera appalachiensis sp. nov., (Diptera: Cecidomyiidae: Winnertziinae: Heteropezini) from the eastern United States is described and illustrated, and pertinent collection and biological data are also provided. It is compared to its congeners, N. kiefferi (Meunier), a subfossil described from African copal, and N. congoensis Gagné, from the Congo. This rare, exciting discovery is the first record of the genus Neostenoptera in the New World.
509
Parajulid milliped studies XI : Initial assessment of the tribe Gosiulini (Diplopoda: Julida)
(2016)
The parajulid milliped tribe Gosiulini (Diplopoda: Julida) comprises two genera – Gosiulus Chamberlin, with three projections on the posterior gonopod and two species in the southcentral/southwestern United States (US) [Arizona, Colorado, New Mexico, and Texas], and monotypic Minutissimiulus Shelley, n. gen., with two projections, in Nuevo León, Mexico. Gosiulus conformatus Chamberlin occupies the plains/fl atlands of Texas, while its congener inhabits high elevations to the west in all four US states. Both are anticipated in Mexico (Coahuila, Chihuahua, and Sonora), and G. conformatus is expected in southeastern Colorado, eastern New Mexico, and the Oklahoma panhandle. The eastern boundary of G. conformatus and the genus/tribe conforms to the western border of the Piney Woods biome in eastern Texas. As shown by the posterior gonopod drawing in the original description, Parajulus timpius Chamberlin, previously considered of “uncertain generic position or validity,” is unquestionably the oldest name for the western species. The anteriormost posterior gonopod projection, absent from Minutissimiulus, is considered the “prefemoral process,” while the “solenomere” and a third branch arise from a common base.
Because of positional homology with “process ‘C’” in Nesoressini, the last projection is accorded this name, which may also apply to the “prefemoral process” in Aniulini. Minutissimiulus biramus Shelley, n. sp., is proposed along with the following new subjective synonymies: Apacheiulus Loomis under Gosiulus; Ziniulus aethes and Z. medicolens, both by Chamberlin, and Z. ambiguus and Z. nati, both by Loomis, under G. conformatus; and A. pinalensis and A. guadelupensis, both by Loomis, under G. timpius, new combination. Ziniulus navajo Chamberlin becomes an objective synonym of P. timpius because its holotype is designated neotype of the latter. Minutissimiulus biramus Shelley is the fi rst Mexican gosiuline and “mainland” Mexican parajulid not in the tribe Parajulini.
508
507
506
A revised annotated checklist for the butterfly family Lycaenidae (Lepidoptera) of Trinidad is presented, updating nomenclature, and indicating synonyms from earlier lists and papers. The checklist includes 131 species of Lycaenidae, comprising 127 species of 49 genera of Eumaeini, Theclinae, and four species of three genera of Polyommatinae.
There are more than 30 new island records. No lycaenid species is endemic to Trinidad, and the fauna consists primarily of widespread species (71%) that occur from Central America to the Amazon Basin. However, the primary biogeographic affi nity is the Amazon Region, where 94% of the Trinidad lycaenid fauna also occurs.
Corrections are made to the literature cited in the first author’s earlier checklists on other Trinidad butterflies.
505
The genus Rhinolaemus Steel is revised. A new island and a new country record are presented for the type species, R. maculatus Steel. A new species, R. niueensis Thomas, new species, is described from Niue, and Rhinolaemus tuberculatus (Grouvelle), new combination, is transferred from Laemophloeus (sens. lat.). The members of the genus are illustrated and a key to their identification is presented.
504
Epimelitta Bates, 1870 (Coleoptera, Cerambycidae) is redescribed with two species: Epimelitta scoparia (Klug, 1825) and Epimelitta rufiventris Bates, 1870; Epimelitta meliponica Bates, 1870 syn. nov. and Epimelitta acutipennis Fisher, 1947 syn. nov. are considered junior synonyms of E. scoparia. Exepimelitta gen. nov. is described with five species: Exepimelitta mimica (Bates, 1873), Ex. nigerrima (Bates, 1892), Ex. consobrina (Melzer, 1931) (=Epimelitta nigerrima var. flavipubescens Fisher, 1947, syn. nov.), Ex. lestradei (Peñaherrera-Leiva and Tavakilian, 2003) and Ex. windsori sp. nov. Charisia Champion, 1892 is revalidated and redescribed with six species: Charisia euphrosyne (Newman, 1840), C. mneme (Newman, 1841), C. melanaria Gounelle, 1911, C. ornaticollis Zajciw, 1973, C. bleuzeni (Peñaherrera-Leiva and Tavakilian, 2003) and C. durantoni (Peñaherrera-Leiva and Tavakilian, 2003).
Erratamelitta gen. nov. is described with two species: Erratamelitta erato (Newman, 1840) (= Epimelitta bicolor (Bates, 1873), syn. nov.) and Er. eliasi sp. nov.. Adepimelitta gen. nov. is described with two species: Adepimelitta debilis (Gounelle, 1911) and Ad. eupheme (Lameere, 1884). A brief synopsis of the genus Acorethra Bates, 1873 is presented, with two species included: Acorethra chrysaspis Bates, 1873, revalidated, and Ac. aureofasciata Gounelle, 1911. All species are illustrated (including genitalia when available); and keys to the genera, and their species, are provided.
503
Seven specimens of Sybra alternans (Wiedemann) (Cerambycidae: Lamiinae: Apomecynini) were captured with a UV light trap and by beating branches of trees in the urban area of Easter Island (Chile) during 2011–2016, representing the fi rst record of the species in this Chilean insular territory, but not continental Chile.
Sybra alternans is native to Southeast Asia and has been introduced accidentally to Hawaii and Florida in the United States of America. Data are presented from the literature on this species’ distribution, host plants, and biological information.
502
Taxonomic supplement (2001 to 2015) to the catalogue of New Zealand Carabidae (Insecta: Coleoptera)
(2016)
A supplement to the “Carabidae (Insecta: Coleoptera): catalogue” published by Larochelle and Larivière (2001; Fauna of New Zealand 43), is provided. A total of 99 genera and 547 species and subspecies belonging to 21 tribes and 8 subfamilies of Carabidae (including Cicindelini and Rhysodini), is recorded for New Zealand. Endemism is high, with 93% of the species and subspecies as well as 62% of the genera recorded from this country only. Changes to the 2001 catalogue are documented. The synonymy and type locality of taxa described between 2001 and 2015 are also provided.
500
Andaspis recurrens Takagi and Kawai, Hypaspidiotus jordani (Kuwana), Lepidosaphes kamakurensis (Kuwana) and Selenomphalus distylii Takagi from warm temperate forests are newly documented in the Korean fauna of armored scale insects (Hemiptera: Diaspididae). Characteristics of these species are briefl y given and illustrative photographs and information on their distribution and hosts are provided.
499
Nine new species of Hydroptilidae (Trichoptera) are described and illustrated from the western United States, Canada, Mexico, and Belize: Seven new species in the genus Neotrichia Morton, N. buenoi, N. chihuaha, N. contrerasi, N. maya, N. palitla, N. pinnacles, and N. sandersoni, one new species in the genus Oxyethira Eaton, O. pembertonensis, and one new species in the genus Metrichia Ross, M. mastelleri. As well, new records for N. mobilensis and Mayatrichia tuscaloosa and new illustrations of the male genitalia of Neotrichia jarochita Bueno-Soria and Mayatrichia tuscaloosa Harris and Sykora are provided.
498
The Iranian fauna of Cucujidae, Laemophloeidae, and Silvanidae (Coleoptera: Cucujoidea) are summarized in this paper. In total 2 species of Cucujidae (1 genus: Pediacus Shuckard), 6 species of Laemophloeidae (3 genera: Cryptolestes Ganglbauer, Laemophloeus Dejean, and Placonotus MacLeay) and 7 species of Silvanidae (6 genera: Uleiota Latreille, Psammoecus Latreille, Ahasverus Gozis, Nausibius Lentz, Oryzaephilus Ganglbauer, Psammoecus Latreille, and Silvanus Latreille) are listed in this paper. Synonymies and distribution data are given.
497
The previously published list of Coleoptera holotypes in the Museo de Historia Natural, Noel Kempff Mercado (MNKM), Santa Cruz, Bolivia is updated to include those deposited in the Museo since then though the end of 2015. Literature citations for the original descriptions of each listed holotype are also provided along with summary comments regarding family composition and authorship of included species. Photographs of collector, authors, habitats, and major Bolivian type repositories are provided.
496
495
Descriptions or diagnoses are given for 36 species of New World Eremoleon including 12 new species:
Eremoleon attenuatus, E. durangoensis, E. jacumba, E. jamaica, E. inca, E. monagas, E. morazani, E. pygmaeus, E. samne, E. tanya, E. tepuyiensis and E. venezolanus. A neotype is designated for Hesperoleon atomarius Navás 1933.
The larvae of 25 species are described and keyed. Keys to the adults and larvae are given, and biological notes are provided. Twenty-five species were reared from larvae found in cave mouths, rock overhangs, or other less common habitats; for example, E. nigribasis were found in deep recesses of mammal burrows, E. gracile were reared from reptile holes in the ground, E. punctipennis were found in beetle frass beneath logs, and E. femoralis in small twig holes or abandoned termite galleries in termite frass. Many new bombyliid and chalcidid parasites are listed.
494
The primary types of Onciderini Thomson, 1860 of the Field Museum of Natural History (FMNH) are catalogued and illustrated. Data on the original combination, current name, gender, and type locality are verifi ed and presented. There are 14 primary types of Onciderini including fi ve in Oncideres Lacordaire, 1830; three in Charoides Dillon and Dillon, 1945; and two in Jamesia Jekel, 1861. Of the 14 primary types, 13 were described by L.S. Dillon and E.S. Dillon. One lectotype is designated. A brief history of the Coleoptera collection at the FMNH is also presented.
493
Fifteen species of longhorn beetles (Coleoptera, Cerambycidae) are reported for the fi rst time for Venezuela:
Ambonus electus (Gaham, 1903), Eusapia guyanensis Huedepohl, 1988, Acyphoderes abdominalis (Olivier, 1795), Isthmiade ichneumoniformis Bates, 1870, Drychateres bilineatus (Olivier, 1795), Polyschisis rufi tarsalis Waterhouse, 1880, Carphina petulans (Kirsch, 1875), Lepturges zonula Monné, 1976, Oreodera albata Villiers, 1971, Psapharochrus chrysopus (Bates, 1861), Estola fratercula Galileo y Martins, 1999, Oncideres cephalotes Bates, 1865, Polyrhaphis spinosa (Drury, 1773), Nicias alurnoides (Thomson, 1857) and Ialyssus tuberculatus (Olivier, 1795). The known prior distributions for each species, collecting methods and additional information on the collection sites of each specimen are also provided.
492
491
Four new species of the genus Polyphylla Harris (Coleoptera: Scarabaeidae: Melolonthinae) from the southwestern United States and Baja California, Mexico, are described and illustrated: Polyphylla anivallis, P. koso, P. morroensis, and P. socorriana. Two nomenclatural changes are proposed: Polyphylla ratcliffei Young is placed into synonymy with P. avittata Hardy and Andrews, new synonymy; P. uteana Tanner is removed from synonymy and reinstated as a valid species, reinstated status. The females of P. monahansensis Hardy and Andrews and P. stellata Young are described and illustrated. New distributional records and ecological associations are amended for P. avittata Hardy and Andrews, P. cavifrons LeConte, P. concurrens Casey, P. crinita LeConte, P. diffracta Casey, P. mescalerensis Young, P. petitii (Guérin-Méneville), and P. rugosipennis Casey. A checklist of the “Western Clade” species of Polyphylla, north of Mexico, is included.
490
Recent shifts in US policies towards Cuba suggest a relaxation or lifting of the embargo may occur in the near future. With the prospects of open travel and trade with Cuba come concerns over the introduction of agricultural pests. In an effort to assess these concerns the distribution-based introduction risk of pests listed in the 2015 Cooperative Agricultural Pest Survey’s (CAPS) list of priority pests of economic and environmental importance is reviewed. Of the 59 pests on the CAPS priority pest list, 20 have been recorded in the literature as being present in the Caribbean Basin, South America and Central America. For these 20 New World pests a commodity and distribution-based risk rating was assigned to describe their potential for introduction through the Cuba-Florida pest pathway. The highest rating was given to the six listed pests currently reported as being present in Cuba, and potential for introduction and subsequent impact of these six pests on Florida agriculture is discussed. In addition to the pests found on the 2015 CAPS priority pest list, information regarding pests of concern in the family Tephritidae and the Old World bollworm Helicoverpa armigera (Hübner), is also included, as is a description of the Cuban plant health and regulatory structure.
The significance of plant-pest introductions between Cuba and Florida is discussed, with an emphasis on proactive engagement in research and collaboration to address these issues.
489
Chionaspis acer (Takagi and Kawai) and Chionaspis wistariae Cooley, occurring respectively on Acer and Wisteria plants, are newly reported in the Korean fauna of armored scales (Hemiptera: Diaspididae). The characters of these species are briefl y redescribed with illustrative photographs and information on distribution and hosts. A dichotomous key to species of Chionaspis Signoret in Korea is provided for correct species identifi cation.
488
Two Neoibidionini from Bolivia are described as new: Neocompsa schneppi sp. nov. and Tropidion nancyae sp. nov. The new species are compared to closely related species and previous keys by Martins and Galileo (2007, 2009) are modifi ed to include them. Illustrations of dorsal, ventral and lateral habitus, as well as head structures are included for both species. Comments are also included on color and pattern variation of each species.
487
New records of Myrmeleontidae from Paraguay are presented: Dimares elegans (Perty, 1833), Glenurus croesus Banks, 1922, Glenurus penningtoni (Navás, 1918a) and Vella fallax (Rambur, 1842). The total number of species now recorded from the country is fourteen among 11 genera. Specimens of Dimares elegans from east (Cerrado) and west (Chaco) of the Paraguay River are phenotypically distinct; those west of the Paraguay River are assigned to the form “lepida” Navás, 1912. More research is required to understand the taxonomic limits in this genus.
486
Cyrea Gordon and Canepari (121 species) and Tiphysa Mulsant (2 species) are discussed, species are described, illustrations are provided, and a key to all recognized species is included. New synonyms recognized are:
Hyperaspis arrowi var. darwini Brèthes = Cyrea arrowi (Brèthes); Hyperaspis trivittata Weise, Hyperaspis mundula Weise = Cyrea emiliae (Mulsant); Cleothera scapulata Mulsant, Cleothera mercabilis Mulsant, Hyperaspis iheringi Weise = Cyrea flavoguttata (Mulsant); Cleothera gracilis Mulsant = Cyrea hexastigma (Mulsant); Cleothera triacantha Mulsant = Cyrea novemsignata (Herbst); Cleothera distinguenda Mulsant = Cyrea ormanceayi (Mulsant); Cleothera sexnotata Brèthes = Cyrea quinquenotata (Mulsant); Cleothera schaufussi Vogel and Hyperaspis adelaida Gorham = Cyrea tessulata (Mulsant). A total of 76 new species of Cyrea are described: Cyrea agnes, C. allison, C.alma, C. annette, C. arlene, C. audrey, C. beatrice, C. bernice, C. bessie, C. brittany, C. carla, C. charlene, C. claudia, C. colleen, C. constance, C. courtney, C. dana, C. dolores, C. dora, C. eileen, C. ella, elsie, erica, georgia, gertrude, gina, glenda, heidi, holly, ida, jackie, jeanne, jeannette, jessie, jo, C. joy, C. june, C. katie, C. kristen, C. laurie, C. lillie, C. lucille, C. lucy, C. lydia, C. marcia, C. marian, C. marion, C. marlene, C. mattie, C. maureen, C. maxine, C. melanie, C. melinda, C. minnie, C. natalia, C. nellie, C. pearl, C. pseudospinalis, C. renee, C. roberta, C. rosemary, C. samantha, C. stacy, C. stella, C. sue, C. tamara, C. tanya, C. tara, C. terry, C. vanessa, C. vera, C. vicki, C. viola, C. vivian, C. willie, C. wilma, C. yolanda, C.
yvonne. Lectotypes here designated for C. collaris, C. compta, C. distinguenda, C. exclamationis, C. fasciata, C. ferruginiceps, C. flavoguttata, C. iheringi, C. languida, C. maculosa, C. melaneura, C. mundula, C. noticollis, C. novemsignata, C. ormanceayi, C. ornaticolis, C. quinquenotata, C. renifera, C. schaufussi, C. sexguttata, C. spinalis, C. tessulata, C. trepida, C. triacantha
485
The Bermuda grass scale Odonaspis ruthae Kotinsky, 1915 (Hemiptera: Coccomorpha: Diaspididae) is reported for the first time in Colombia. The scale insect was collected in two localities, in northwestern and southwestern Colombia. This is the first record of the tribe Odonaspidini in Colombia. Aspects of the distribution and biology of the species are discussed. A key to separate the species of Odonaspis recorded in the Neotropical region is provided.
0484
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0480
Crypticerya brasiliensis (Hempel) and Crypticerya genistae (Hempel) (Hemiptera: Monophlebidae:
Iceryini) are herein reported for the fi rst time in Colombia. The above two species and Crypticerya multicicatrices Kondo and Unruh and Crypticerya zeteki (Cockerell) are briefl y diagnosed based on the adult females. The presence of C. zeteki in Colombia is confi rmed and a key to the adult females of species of the tribe Iceryini reported in Colombia is provided.
0479
0478
Cedar Point Biological Station (CPBS) is located in the mixed grass prairie of the central Great Plains, at the transition between the subregions known as the “tall grass” and “short grass” prairies. Adding to the habitat diversity, there are wetlands and riparian habitats associated with the North Platte River and the edge of the Sandhills region of north central Nebraska. This concurrence of habitats supports a diverse small mammal community. The purpose of this paper is to assemble all published information on ectoparasites associated with small mammals (Insectivora, Rodentia) of southwestern Nebraska, and to report the results of an intensive survey carried out by students of the Parasitology fi eld course during two summers at CPBS. In 2012 and 2013, 27 species of mammal-associated arthropods were collected, including fi ve species of sucking lice (Anoplura), a chewing louse (Ischnocera), six species of fl eas (Siphonaptera), thirteen species of mesostigmatic mites (Laelapidae, Macronyssidae, Macrochelidae), and two species of metastigmatic ticks (Ixodidae). These specimens were brushed from the pelage of 11 species of small mammals that were captured in a variety of habitats around CPBS. The arthropod
list includes 17 new records for the State of Nebraska. This collection is housed in the Harold W. Manter Laboratory of Parasitology (HWML), University of Nebraska State Museum, at the University of Nebraska-Lincoln, and serves as a taxonomic base for our continued efforts to establish a long-term catalog of parasites associated with small mammals in southwestern Nebraska.
0477
Two new species of the mexicanus group of Vaejovis C.L. Koch are described from the Madrean pine-oak forests of the Sierra Madre Occidental in the state of Durango, Mexico. These species, Vaejovis sierrae sp. nov. and Vaejovis mcwesti sp. nov., are distinguished from each other and the only other species of the mexicanus group known from this mountain range, Vaejovis montanus Graham and Bryson, by morphometrics, carinal development of the pedipalps, granulation of the metasoma, and body size. A key to the species of the mexicanus group from
the Sierra Madre Occidental is provided.
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0474
Descriptions and illustrations of the fi fth instar and prepupal larval stages of Stethon pectorosus LeConte, 1866 (Coleoptera: Eucnemidae: Eucneminae: Mesogenini) are provided. These are the fi rst larval descriptions for the genus Stethon LeConte within the subfamily Eucneminae in the Nearctic region. Biological information is provided and comparison with larval morphology of other Eucnemidae is briefl y discussed.