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Highlights
• Protocol for extracting and analyzing pollen grains from fossil insects
• Individual fossil grains can be analyzed using a combined approach
• Simple and fast TEM embedding and sectioning protocol
• Protocol enables a taxonomic assignment of pollen
Summary
This protocol explains how to extract pollen from fossil insects with subsequent descriptions of pollen treatment. We also describe how to document morphological and ultrastructural features with light-microscopy and electron microscopy. It enables a taxonomic assignment of pollen that can be used to interpret flower-insect interactions, foraging and feeding behavior of insects, and the paleoenvironment. The protocol is limited by the state of the fossil, the presence/absence of pollen on fossil specimens, and the availability of extant pollen for comparison.
Fossil dental remains are an archive of unique information for paleobiological studies. Computed microtomography based on X-ray microfocus sources (X-μCT) and Synchrotron Radiation (SR-μCT) allow subtle quantification at the micron and sub-micron scale of the meso- and microstructural signature imprinted in the mineralized tissues, such as enamel and dentine, through high-resolution “virtual histology”. Nonetheless, depending on the degree of alterations undergone during fossilization, X-ray analyses of tooth tissues do not always provide distinct imaging contrasts, thus preventing the extraction of essential morphological and anatomical details. We illustrate here by three examples the successful application of neutron microtomography (n-μCT) in cases where X-rays have previously failed to deliver contrasts between dental tissues of fossilized specimen.
Significance
Identifying the earliest members of the genus Homo is crucial for understanding when and where selective pressures resulted in its emergence from a Plio-Pleistocene hominin taxon. Our revision of a large part of the dental fossil record from southern Africa provides evidence suggesting a paucity of Homo remains and indicates increased levels of dental variation in australopith taxa. Results of the Ba/Ca, Sr/Ca, and elemental mapping of enamel and dentine also indicate that some of the purported Homo specimens show a paleoecological signal similar to that of the australopiths.
Abstract
The origins of Homo, as well as the diversity and biogeographic distribution of early Homo species, remain critical outstanding issues in paleoanthropology. Debates about the recognition of early Homo, first appearance dates, and taxonomic diversity within Homo are particularly important for determining the role that southern African taxa may have played in the origins of the genus. The correct identification of Homo remains also has implications for reconstructing phylogenetic relationships between species of Australopithecus and Paranthropus, and the links between early Homo species and Homo erectus. We use microcomputed tomography and landmark-free deformation-based three-dimensional geometric morphometrics to extract taxonomically informative data from the internal structure of postcanine teeth attributed to Early Pleistocene Homo in the southern African hominin-bearing sites of Sterkfontein, Swartkrans, Drimolen, and Kromdraai B. Our results indicate that, from our sample of 23 specimens, only 4 are unambiguously attributed to Homo, 3 of them coming from Swartkrans member 1 (SK 27, SK 847, and SKX 21204) and 1 from Sterkfontein (Sts 9). Three other specimens from Sterkfontein (StW 80 and 81, SE 1508, and StW 669) approximate the Homo condition in terms of overall enamel–dentine junction shape, but retain Australopithecus-like dental traits, and their generic status remains unclear. The other specimens, including SK 15, present a dominant australopith dental signature. In light of these results, previous dietary and ecological interpretations can be reevaluated, showing that the geochemical signal of one tooth from Kromdraai (KB 5223) and two from Swartkrans (SK 96 and SKX 268) is consistent with that of australopiths.
Neanderthal diet has been on the spotlight of paleoanthropological research for many years. The majority of studies that tried to reconstruct the diet of Neanderthals were based on the analysis of zooarchaeological remains, stable isotopes, dental calculus and dental microwear patterns. In the past few years, there have been a few studies that linked dental macrowear patterns of Neanderthals and modern humans to diet and cultural habits. However, they mostly focused on maxillary molars. Although mandibular molars have been widely used in microwear dietary research, little is known about their usage at the macroscopic scale to detect information about human subsistence strategies. In this study, we compare the macrowear patterns of Neanderthal (NEA), fossil Homo sapiens (FHS), modern hunter-gatherers (MHG), pastoralists, early farmers and Australian Aborigines from Yuendumu mandibular molars in order to assess their utility in collecting any possible information about dietary and cultural habits among diverse human groups. We use the occlusal fingerprint analysis method, a quantitative digital approach that has been successfully employed to reconstruct the diet of living non-human primates and past human populations. Our results show macrowear pattern differences between meat-eater MHG and EF groups. Moreover, while we did not find eco-geographical differences in the macrowear patterns of the fossil sample, we found statistically significant differences between NEA and FHS inhabiting steppe/coniferous forest. This latter result could be associated with the use of distinct technological complexes in these two species, which ultimately could have allowed modern humans to exploit natural resources in a different way compared to NEA.
Palaeosphryon menatensis gen. et sp. nov., first unambiguous representative of the longhorn beetle subfamily Prioninae from the Paleocene of Menat (France), is described and illustrated. The new fossil is placed into the tribe Prionini, showing some similarities with some species of the extant genera Osphryon (Papua New Guinea) and Titanus (Brazil, Colombia, Guianas, Ecuador, Peru), viz. in general body shape, antennomere 3 as long as first and second together but shorter than the length of fourth plus fifth, elongate elytra, and small spines on the lateral margin of the pronotum disposed in a relatively similar way as in Osphryon. Nevertheless, the exact affinities of the new fossil within the Prionini remain uncertain because of the lack of a recent phylogenetic analysis in which it could be integrated. This fossil beetle is exceptional for its very large size, with a body 70 mm long. Some other large longhorn beetles have been found in the same outcrop, and are awaiting description. The positions of the previously described Cerambycidae from Menat are also discussed. This exceptional fauna of Cerambycidae is in accordance with the current palaeoenvironmental reconstruction for the Menat Konservat-Lagerstätte, as a small maar lake surrounded by a warm and humid, probably evergreen forest.
The ammonoids of the family Maenioceratidae from Givetian sedimentary rocks of the Anti-Atlas (Morocco) are investigated. The study is based on new collections stored in the Museum für Naturkunde, Berlin. The genera Maenioceras Schindewolf, 1933 and Afromaenioceras Göddertz, 1987 are revised; the genus Trimaenioceras is newly described. The species Maenioceras afroterebratum sp. nov., Maenioceras mzerrebense sp. nov., Maenioceras oufranense sp. nov., Maenioceras beckeri sp. nov., Afromaenioceras sulcatostriatum (Bensaïd, 1974), Afromaenioceras hiemale sp. nov., Afromaenioceras bensaidi sp. nov., Afromaenioceras brumale sp. nov., Afromaenioceras crassum (Bensaïd, 1974), Trimaenioceras klugi gen. et sp. nov., Trimaenioceras eculeus gen. et sp. nov., Trimaenioceras fuscina gen. et sp. nov. and Trimaenioceras paucum gen. et sp. nov. are described in detail.
The first sawfly from the Oligocene of Céreste (Southern France (Hymenoptera: Tenthredinidae)
(2024)
Luberotenthredo cerestensis gen. et sp. nov. is the first record of the sawfly family Tenthredinidae from the Oligocene of Céreste (Southern France). This taxon is described and illustrated based on a well-preserved specimen. This genus resembles the extant genus Perineura (subfamily Tenthredininae, tribe Perineurini) with which it shares forewing venation similarities and numerous morphological characters. This new taxon is the first fossil representative of the tribe Perineurini and can be used as a calibration point for future investigation of the diversification of the family Tenthredinidae.
Here, we report on 33 molluscan species from Miocene ’Calcari a Lucina’ hydrocarbon-seep deposits in northern Italy. Three new species are described: the chilodontaid gastropod Putzeysia diversii sp. nov., the lucinid bivalve Miltha (sensu lato) romaniae sp. nov., and Sisonia ultimoi sp. nov., a heterodont bivalve of uncertain taxonomic affinity. Fourteen species are described in open nomenclature. The common but enigmatic gastropod species Phasianema taurocrassa is here suggested to belong to the seguenzioid genus Cataegis. Most gastropod species are inhabitants of the deep-sea floor in general, and are not restricted (obligate) to sites of hydrocarbon-seepage. The gastropod Putzeysia diversii sp. nov. and the bivalve Sisonia ultimoi sp. nov. are the geologically oldest members of their genera known to date. While the genus Putzeysia is geographically restricted to the NE Atlantic Ocean and the Mediterranean Sea, Sisonia ultimoi sp. nov. represents another link of the Miocene Mediterranean seep fauna to that of the central Indo-West Pacific Ocean.
Lacewing larvae in the Cretaceous were more diverse in appearance than they are today, best documented by numerous fossils preserved in amber. One morphotype of an unusual larva from about 100 Ma old Kachin amber (Myanmar) was formally recognised as a distinct group called Ankyloleon. The original description erected a single formal species, Ankyloleon caudatus. Yet, it was indicated that among the five original specimens, more species were represented. We here report five new specimens. Among these is the so far largest as well as the so far smallest specimen. Based on this expanded material we can estimate certain aspects of the ontogenetic sequence and are able to recognise a second discrete species, Ankyloleon caroluspetrus sp. nov. We discuss aspects of the biology of Ankyloleon based on newly observed details such as serrations on the mandibles. Long and slender mouthparts, legs and body together with a weakly expressed outer trunk segmentation provide indications for a lifestyle hunting for prey in more confined spaces. Still many aspects of the biology of these larvae must remain unclear due to a lack of a well comparable modern counterpart, emphasising how different the fauna of the Cretaceous was.
The genus Plateosaurus is one of the most challenging aspects of early-diverging sauropodomorph taxonomy, with a total of 29 names and a century of revisions. As a result, nomina dubia have been treated as synonyms of Plateosaurus trossingensis, adding to the confusion about the extent of morphological variability. Here, we provide a thorough revision of the taxonomy proposed by von Huene as well as the subsequent taxonomic interpretations of Plateosaurus through a complete inventory of the specimens stored in the Palaeontological Collection of Tübingen. We reassess the status of the specimens in the collection and propose that the sauropodomorph-bearing layers are not necessarily monospecific. Most of the original fieldwork documentation has been lost, but we use the taxonomy established by von Huene as a historical reference point to reconstruct what was known at the time. This revised taxonomy of Plateosaurus narrows the genus to three species: Pl. trossingensis, Pl. longiceps, and Pl. gracilis (as a metataxon), and restricts the genera Gresslyosaurus and Pachysaurus to large and robust individuals, as pragmatic decisions aimed to test their affinities to other Late Triassic sauropodomorphs. Future studies should consider not only morphological variability, but also stratigraphy, palaeogeography, and environmental data when delineating species within and outside the Plateosaurus plexus.
The Miocene was a key time in the evolution of African ecosystems witnessing the origin of the African apes and the isolation of eastern coastal forests through an expanding arid corridor. Until recently, however, Miocene sites from the southeastern regions of the continent were unknown. Here, we report the first Miocene fossil teeth from the shoulders of the Urema Rift in Gorongosa National Park, Mozambique. We provide the first 1) radiometric ages of the Mazamba Formation, 2) reconstructions of paleovegetation in the region based on pedogenic carbonates and fossil wood, and 3) descriptions of fossil teeth. Gorongosa is unique in the East African Rift in combining marine invertebrates, marine vertebrates, reptiles, terrestrial mammals, and fossil woods in coastal paleoenvironments. The Gorongosa fossil sites offer the first evidence of woodlands and forests on the coastal margins of southeastern Africa during the Miocene, and an exceptional assemblage of fossils including new species.
Thylacocephalans are enigmatic euarthropods, known at least from the Silurian to the Cretaceous. Despite remaining uncertainties concerning their anatomy, key features can be recognised such as a shield enveloping most of the body, hypertrophied compound eyes, three pairs of raptorial appendages and a posterior trunk consisting of eight up to 22 segments bearing appendages and eight pairs of gills. Well-known for its euarthropod diversity, the La Voulte-sur-Rhône Lagerstätte (Callovian, Middle Jurassic, France) has provided many remains of four thylacocephalan species so far: Dollocaris ingens, Kilianicaris lerichei, Paraostenia voultensis and Clausocaris ribeti. In this paper, we study the type material as well as undescribed material. The re-description of La Voulte thylacocephalans reveals an unexpected diversity, with the description of two new species, Austriocaris secretanae sp. nov. and Paraclausocaris harpa gen. et sp. nov., and of specimens of Mayrocaris, a taxon originally described from Solnhofen Lagerstätten. We also reassign Clausocaris ribeti to Ostenocaris. The reappraisal of La Voulte thylacocephalans also provides important insight into the palaeobiology of Thylacocephala. New key anatomical features are described, such as an oval structure or a putative statocyst, which indicate a nektonic or nektobenthic lifestyle. Finally, we document a juvenile stage for Paraostenia voultensis.
Oligoptilomera luberonensis gen. et sp. nov., first fossil representative of the gerrid subfamily Ptilomerinae, is described and figured from the Oligocene of Murs (Vaucluse, Southern France). Extant Ptilomerinae live in streams in warm climates, of the Indo-Malaysian, eastern Palaearctic, and Papouan regions. The discovery of this Oligocene French Ptilomerinae is in accordance with the putative age of the subfamily, at least older than the Eocene, and with the Indo-Malaysian affinities previously recorded for some other insects from the Oligocene of France. The two insect assemblages of Murs and Céreste are compared and the differences discussed. Although of similar ages, that from Murs was possibly corresponding to a more shallow water paleolake than that of Céreste.
This work provides the first revision and illustration of the type material of the ammonite species Parahoplites abichi Anthula [Type species of Protacanthoplites Tovbina] and Parahoplites aschiltaensis Anthula [Type species of Acanthohoplites Sinzow] from the upper Aptian (Lower Cretaceous) of Dagestan, Russia. The close affinities and synonymy between these two species are confirmed, and Acanthohoplites aschiltaensis is here retained as the senior valid name by its long quoting history and its historical use as a zonal index of the upper Aptian. The genus Protacanthoplites should be thus synonymised with Acanthohoplites by priority in the date of publication. Comparison with, and distinction from, closely allied Acanthohoplitidae is provided.
The railway cutting near Oberrödinghausen at the northern margin of the Rhenish Mountains is the cardinal section for the investigation of Early Tournaisian (Early Carboniferous; Mississippian) ammonoids. The ammonoids from the Hangenberg Limestone (= Gattendorfia Limestone) of this and neighbouring outcrops are revised here, using the historical collections as well as undescribed new material. The ammonoid assemblages are composed of a total of 67 species, which occur in four successive ammonoid zones. The assemblages are composed of predominant prionoceratids (Order Goniatitina) with the twenty genera Mimimitoceras (two species), Globimitoceras (one species), Paragattendorfia (two species), Kornia (three species), Stockumites (eleven species), Acutimitoceras (two species), Costimitoceras (one species), Nicimitoceras (four species), Imitoceras (one species), Voehringerites (one species), Gattendorfia (eight species), Zadelsdorfia (two species), Kazakhstania (one species), Gattenpleura (one species), Weyerella (three species), Hasselbachia (three species), Paprothites (five species), Pseudarietites (three species), Rodingites (two species), Paralytoceras (one species) as well as subordinate eocanitids (Order Prolecanitida) with the genera Eocanites (eight species) and Nomismocanites (one species). The new genera Rodingites gen. nov. and Nomismocanites gen. nov. as well as the new species Mimimitoceras perditum sp. nov., Kornia fibula sp. nov., Kornia acia sp. nov., Stockumites parallelus sp. nov., Stockumites voehringeri sp. nov., Acutimitoceras ucatum sp. nov., Acutimitoceras paracutum sp. nov., Imitoceras initium sp. nov., Gattendorfia rhenana sp. nov., Gattendorfia bella sp. nov., Gattendorfia valdevoluta sp. nov., Gattendorfia schmidti sp. nov., Gattendorfia corpulenta sp. nov., Gattendorfia immodica sp. nov., Zadelsdorfia oblita sp. nov., Weyerella lenis sp. nov., Hasselbachia erronea sp. nov., Paprothites beckeri sp. nov., Paprothites kullmanni sp. nov., Eocanites delicatus sp. nov. and Nomismocanites raritas gen. et sp. nov. are described from Oberrödinghausen. Mimimitoceras mina sp. nov., Stockumites marocensis sp. nov., Zadelsdorfia zana sp. nov. and Kazakhstania kana sp. nov. are newly named for material from the Anti-Atlas of Morocco.
The early Tournaisian (Early Carboniferous; Mississippian) ammonoids from the classical abandoned limestone quarry of Gattendorf (Upper Franconia) are revised, using the historical collections as well as so far undescribed material. The ammonoid assemblage is composed of prionoceratid ammonoids of the six genera Mimimitoceras, Paragattendorfia, Stockumites, Acutimitoceras, Gattendorfia and Gattenpleura, which indicate a stratigraphic position near the Devonian–Carboniferous boundary in the earliest Carboniferous. The new species Stockumites hofensis sp. nov. and S. nonaginta sp. nov. are described.
The evolution and interrelationships of carnivorous squamates (mosasaurs, snakes, monitor lizards, Gila Monsters) are a contentious part of reptile systematics and go to the heart of conflict between morphological and molecular data in inferring evolutionary history. One of the best-preserved fossils in this motley grouping is “Saniwa” feisti Stritzke, 1983, represented by complete skeletons from the early-middle Eocene of Messel, Germany. We re-describe it on the basis of superficial examination, stereoradiography, and high-resolution X-ray computed tomography of new and published specimens. The scalation of the lizard is unique, consisting of small, keeled scales on the head (including a row of enlarged medial supraorbitals) and large, rhomboidal, keeled scales (invested by osteoderms) that covered the rest of the body. Two paired longitudinal rows of enlarged scales ran down the neck. The head was laterally compressed and box-shaped due to the presence of a strong canthal-temporal ridge; the limbs and tail were very long. Notable osteological features include: a toothed, strap-like vomer; septomaxilla with a long posterior process; palpebral with a long posterolateral process; a lacrimal boss and a single lacrimal foramen; a well-developed cultriform process of the parabasisphenoid; two hypoglossal (XII) foramina in addition to the vagus; a lack of resorption pits for replacement teeth; and possibly the presence of more than one wave of developing replacement teeth per locus. There are no osteological modifications suggestive of an intramandibular hinge, but postmortem displacement of the angular-prearticular-surangular complex in multiple specimens suggests that there might have been some degree of mobility in the lower jaw based on soft-tissue modifications. Using phylogenetic analyses on a data-set comprising 473 morphological characters and 46 DNA loci, we infer that a monophyletic Palaeovaranidae Georgalis, 2017, including Eosaniwa Haubold, 1977, lies on the stem of Varanidae Merrem, 1820, basal to various Cretaceous Mongolian taxa. We transfer feisti to the new genus Paranecrosaurus n. gen. Analysis of gut contents reveals only the second known specimen of the cryptozoic lizard Cryptolacerta hassiaca Müller, Hipsley, Head, Kardjilov, Hilger, Wuttke & Reisz, 2011, confirming a diet that was at least partly carnivorous; the preservation of the teeth of C. hassiaca suggests that the gastric physiology of Paranecrosaurus feisti (Stritzke, 1983) n. comb. had high acidity but low enzyme activity. Based on the foregoing and linear discriminant function analysis, we reconstruct P. feisti n. comb., as a powerful, widely roaming, faunivorous-carnivorous stem monitor lizard with a sensitive snout. If the molecular phylogeny of anguimorphs is correct, then many of the features shared by Helodermatidae Gray, 1837 and Varanidae must have arisen convergently, partly associated with diet. In that case, a reconciliation of morphological and molecular data would require the discovery of equally primitive fossils on the helodermatid stem.
The present corrigendum corrects errors that occurred in: Zheng Y., Hu H., Chen D., Chen J., Zhang H. & Rasnitsyn A.P. 2021. New fossil records of Xyelidae (Hymenoptera) from the Middle Jurassic of Inner Mongolia, China. European Journal of Taxonomy 733: 146–159. https://doi.org/10.5852/ejt.2021.733.1229
A massive occurrence of microbial carbonates, including abundant sponge remains, within the Devonian Elbingerode Reef Complex was likely deposited in a former cavity of the fore-reef slope during the early Frasnian. It is suggested that the formation of microbial carbonate was to a large part favored by the activity of heterotrophic, i.e., sulfate-reducing bacteria, in analogy to Quaternary coral reef microbialites. The Elbingerode Reef Complex is an example of an oceanic or Darwinian barrier reef system. In modern barrier reef settings, microbialite formation is commonly further facilitated by weathering products from the central volcanic islands. The Devonian microbialites of the Elbingerode Reef Complex occur in the form of reticulate and laminated frameworks. Reticulate framework is rich in hexactinellid glass sponges, the tissue decay of which led to the formation of abundant micrite as well as peloidal and stromatactis textures. Supposed calcimicrobes such as Angusticellularia (formerly Angulocellularia) and Frutexites, also known from cryptic habitats, were part of the microbial association. The microbial degradation of sponge tissue likely also contributed to the laminated framework accretion as evidenced by the occurrence of remains of so-called “keratose” demosponges. Further typical textures in the microbialite of the Elbingerode Reef Complex include zebra limestone, i.e., the more or less regular intercalation of microbial carbonate and cement. Elevated concentrations of magnesium in the microbialite as compared to the surrounding metazoan (stromatoporoid-coral) reef limestone suggests that the microbialite of the Elbingerode Reef Complex was initially rich in high-magnesium calcite, which would be yet another parallel to modern, cryptic coral reef microbial carbonates. Deposition and accretion of the microbialite largely occurred in oxygenated seawater with suboxic episodes as indicated by the trace element (REE + Y) data.
Climatic niches describe the climatic conditions in which species can persist. Shifts in climatic niches have been observed to coincide with major climatic change, suggesting that species adapt to new conditions. We test the relationship between rates of climatic niche evolution and paleoclimatic conditions through time for 65 Old-World flycatcher species (Aves: Muscicapidae). We combine niche quantification for all species with dated phylogenies to infer past changes in the rates of niche evolution for temperature and precipitation niches. Paleoclimatic conditions were inferred independently using two datasets: a paleoelevation reconstruction and the mammal fossil record. We find changes in climatic niches through time, but no or weak support for a relationship between niche evolution rates and rates of paleoclimatic change for both temperature and precipitation niche and for both reconstruction methods. In contrast, the inferred relationship between climatic conditions and niche evolution rates depends on paleoclimatic reconstruction method: rates of temperature niche evolution are significantly negatively related to absolute temperatures inferred using the paleoelevation model but not those reconstructed from the fossil record. We suggest that paleoclimatic change might be a weak driver of climatic niche evolution in birds and highlight the need for greater integration of different paleoclimate reconstructions.
Biominerals fossilisation: fish bone diagenesis in plio–pleistocene african hominid sites of Malawi
(2020)
Fish fossilisation is relatively poorly known, and skeletal element modifications resulting from predation, burial and diagenesis need to be better investigated. In this article, we aim to provide new results about surface, structural and chemical changes in modern and fossil fish bone. Fossil samples come from two distinct localities of roughly the same age in the Pliocene–Pleistocene Chiwondo Beds adjacent to Lake Malawi. Optical and scanning electron microscope (SEM) observations, energy dispersive spectroscopy (EDS) analyses and Fourier transform infrared (FTIR) spectrometry were carried out on three categories of fish bones: (i) fresh modern samples collected in the lake, (ii) extracted from modern fish eagle regurgitation pellets, and (iii) fossils from Malema and Mwenirondo localities. A comparison of these data allowed us to detect various modifications of bone surfaces and structure as well as composition changes. Some differences are observed between fresh bones and modern pellets, and between pellets and fossils. Moreover, fossil fish bone surface modifications, crystallinity, and chemical composition from Malema and Mwenirondo differ despite their chronological and spatial proximities (2.5–2.4 Ma, 500 m). In both sites, the post-predation modifications are strong and may hide alterations due to the predation by bird of prey such as the fish eagle. The combination of the used methods is relevant to analyses of diagenetic alterations in fish bones.
Das Cranium eines fossilen Hominiden des Formenkreises Homo sapiens sapiens wurde relativ-geologisch sowie absolut durch Radiokohlenstoff und Aminosäuren auf ungefähr 31 000 Jahre B.P. datiert. Andere absolute sowie relative Daten an Mollusken und Mammutzähnen in überlagernden jüngeren Straten datieren auf 18 000 — 21000 und 16 000 Jahren B.P. Geomorphologische und geophysikalische Datierungen stimmen somit gut überein. Er ist der älteste datierte und früheste Bewohner Zentraleuropas, der dem Homo sapiens sapiens angehört.
Our knowledge of early evolution of snakes is improving, but all that we can infer about the evolution of modern clades of snakes such as boas (Booidea) is still based on isolated bones. Here, we resolve the phylogenetic relationships of Eoconstrictor fischeri comb. nov. and other booids from the early-middle Eocene of Messel (Germany), the best-known fossil snake assemblage yet discovered. Our combined analyses demonstrate an affinity of Eoconstrictor with Neotropical boas, thus entailing a South America-to-Europe dispersal event. Other booid species from Messel are related to different New World clades, reinforcing the cosmopolitan nature of the Messel booid fauna. Our analyses indicate that Eoconstrictor was a terrestrial, medium- to large-bodied snake that bore labial pit organs in the upper jaw, the earliest evidence that the visual system in snakes incorporated the infrared spectrum. Evaluation of the known palaeobiology of Eoconstrictor provides no evidence that pit organs played a role in the predator–prey relations of this stem boid. At the same time, the morphological diversity of Messel booids reflects the occupation of several terrestrial macrohabitats, and even in the earliest booid community the relation between pit organs and body size is similar to that seen in booids today.
The fossil record is widely informative about evolution, but fossils are not systematically used to study the evolution of stem-cell-driven renewal. Here, we examined evolution of the continuous growth (hypselodonty) of rodent molar teeth, which is fuelled by the presence of dental stem cells. We studied occurrences of 3,500 North American rodent fossils, ranging from 50 million years ago (mya) to 2 mya. We examined changes in molar height to determine whether evolution of hypselodonty shows distinct patterns in the fossil record, and we found that hypselodont taxa emerged through intermediate forms of increasing crown height. Next, we designed a Markov simulation model, which replicated molar height increases throughout the Cenozoic and, moreover, evolution of hypselodonty. Thus, by extension, the retention of the adult stem cell niche appears to be a predictable quantitative rather than a stochastic qualitative process. Our analyses predict that hypselodonty will eventually become the dominant phenotype.
In seiner Ordnung der Dinge postuliert Michel Foucault bekanntermaßen die Vorgängigkeit der Geschichte der Dinge vor derjenigen des Menschen: Der Mensch, der am Anfang des 19. Jahrhunderts erscheint, hat ihm zufolge keine 'eigene' Geschichte mehr, er ist 'enthistorisiert', weil ihm die Zeit fortan "von woanders her als von ihm selbst [kommt]". Als Subjekt der Geschichte bildet er sich allein "in der Überlagerung der Geschichte der Lebewesen, Dinge und Wörter" und insofern er selbst als lebendes, arbeitendes und sprechendes Wesen erscheint.
Zum Herzstück seiner Analyse der neu aufkommenden Geschichte der Dinge hat Foucault das Fossil gemacht und dargelegt, wie das zweidimensionale, über Ähnlichkeiten verbundene Wissenstableau der Naturgeschichte aufbrach, als das Fossil bei George Cuvier zum Wissensobjekt der Tiefendimension des Lebens und seiner Funktionen wurde. Während Foucault hier weniger auf geologische als vielmehr auf biologische bzw. generell auf wissensgeschichtliche Raum- und Zeitdimensionen abhebt, möchte ich im Folgenden zum einen nachzeichnen, wie das Fossil um 1800 zum Lot der sogenannten Tiefenzeit wurde und welche Her- und Darstellungsweisen von Zeit sich in und mit ihm ergaben. Zum anderen möchte ich aufzeigen, wie gerade im Narrativ des Fossils als Wissens-, aber eben auch als Nichtwissensobjekt der Zeit die Frage kulminierte, welche Tiefen- bzw. Oberflächendimension dem Mensch und seiner Darstellung noch eigen sein kann. Als Leerstelle im Archiv der Fossilienfunde kam ihm nämlich die Zeit nicht nur "von woanders her als von ihm selbst". In der Überlagerung mit der Geschichte der Dinge, Lebewesen und Wörter erreichte sie ihn vielmehr gar nicht mehr. Um diese gerade literarisch ausgeloteten Abgründe der Tiefenzeit wird es mir in der Auseinandersetzung mit E.T.A. Hoffmanns Erzählung 'Die Bergwerke zu Falun' (1819) gehen. Wie ich darlegen möchte, romantisiert Hoffmann gerade im Zeichen des Fossils die Geschichte des Menschen konsequent zu Ende.
This article was republished on October 14, 2015, to correct the order of Figs 8–13. The publisher apologizes for the error. Please download this article again to view the correct version. The originally published, uncorrected article and the republished, corrected article are provided here for reference.
Diverse epidermal appendages including grouped filaments closely resembling primitive feathers in non-avian theropods, are associated with skeletal elements in the primitive ornithischian dinosaur Kulindadromeus zabaikalicus from the Kulinda locality in south-eastern Siberia. This discovery suggests that “feather-like” structures did not evolve exclusively in theropod dinosaurs, but were instead potentially widespread in the whole dinosaur clade. The dating of the Kulinda locality is therefore particularly important for reconstructing the evolution of “feather-like” structures in dinosaurs within a chronostratigraphic framework. Here we present the first dating of the Kulinda locality, combining U-Pb analyses (LA-ICP-MS) on detrital zircons and monazites from sedimentary rocks of volcaniclastic origin and palynological observations. Concordia ages constrain the maximum age of the volcaniclastic deposits at 172.8 ± 1.6 Ma, corresponding to the Aalenian (Middle Jurassic). The palynological assemblage includes taxa that are correlated to Bathonian palynozones from western Siberia, and therefore constrains the minimum age of the deposits. The new U-Pb ages, together with the palynological data, provide evidence of a Bathonian age—between 168.3 ± 1.3 Ma and 166.1 ± 1.2 Ma—for Kulindadromeus. This is older than the previous Late Jurassic to Early Cretaceous ages tentatively based on local stratigraphic correlations. A Bathonian age is highly consistent with the phylogenetic position of Kulindadromeus at the base of the neornithischian clade and suggests that cerapodan dinosaurs originated in Asia during the Middle Jurassic, from a common ancestor that closely looked like Kulindadromeus. Our results consequently show that Kulindadromeus is the oldest known dinosaur with “feather-like” structures discovered so far.
Prehistoric dental treatments were extremely rare, and the few documented cases are known from the Neolithic, when the adoption of early farming culture caused an increase of carious lesions. Here we report the earliest evidence of dental caries intervention on a Late Upper Palaeolithic modern human specimen (Villabruna) from a burial in Northern Italy. Using Scanning Electron Microscopy we show the presence of striations deriving from the manipulation of a large occlusal carious cavity of the lower right third molar. The striations have a "V"-shaped transverse section and several parallel micro-scratches at their base, as typically displayed by cutmarks on teeth. Based on in vitro experimental replication and a complete functional reconstruction of the Villabruna dental arches, we confirm that the identified striations and the associated extensive enamel chipping on the mesial wall of the cavity were produced ante-mortem by pointed flint tools during scratching and levering activities. The Villabruna specimen is therefore the oldest known evidence of dental caries intervention, suggesting at least some knowledge of disease treatment well before the Neolithic. This study suggests that primitive forms of carious treatment in human evolution entail an adaptation of the well-known toothpicking for levering and scratching rather than drilling practices.
Background: Dental biomechanics based on finite element (FE) analysis is attracting enormous interest in dentistry, biology, anthropology and palaeontology. Nonetheless, several shortcomings in FE modeling exist, mainly due to unrealistic loading conditions. In this contribution we used kinematics information recorded in a virtual environment derived from occlusal contact detection between high resolution models of an upper and lower human first molar pair (M1 and M1, respectively) to run a non-linear dynamic FE crash colliding test.
Methodology: MicroCT image data of a modern human skull were segmented to reconstruct digital models of the antagonistic right M1 and M1 and the dental supporting structures. We used the Occlusal Fingerprint Analyser software to reconstruct the individual occlusal pathway trajectory during the power stroke of the chewing cycle, which was applied in a FE simulation to guide the M1 3D-path for the crash colliding test.
Results: FE analysis results showed that the stress pattern changes considerably during the power stroke, demonstrating that knowledge about chewing kinematics in conjunction with a morphologically detailed FE model is crucial for understanding tooth form and function under physiological conditions.
Conclusions/Significance: Results from such advanced dynamic approaches will be applicable to evaluate and avoid mechanical failure in prosthodontics/endodontic treatments, and to test material behavior for modern tooth restoration in dentistry. This approach will also allow us to improve our knowledge in chewing-related biomechanics for functional diagnosis and therapy, and it will help paleoanthropologists to illuminate dental adaptive processes and morphological modifications in human evolution.
The bug Gyaclavator kohlsi Wappler, Guilbert, Wedmann et Labandeira, gen. et sp. nov., represents a new extinct genus of lace bugs (Insecta: Heteroptera: Tingidae) occurring in latest early Eocene deposits of the Green River Formation, from the southern Piceance Basin of Northwestern Colorado, in North America. Gyaclavator can be placed within the Tingidae with certainty, perhaps it is sistergroup to Cantacaderinae. If it belongs to Cantacaderinae, it is the first fossil record of this group for North America. Gyaclavator has unique, conspicuous antennae bearing a specialized, highly dilated distiflagellomere, likely important for intra- or intersex reproductive competition and attraction. This character parallels similar antennae in leaf-footed bugs (Coreidae), and probably is associated with a behavioral convergence as well.
We describe a new large-sized species of hypercarnivorous hyainailourine–Kerberos langebadreae gen. & sp. nov.–from the Bartonian (MP16) locality of Montespieu (Tarn, France). These specimens consist of a skull, two hemimandibles and several hind limb elements (fibula, astragalus, calcaneum, metatarsals, and phalanges). Size estimates suggest K. langebadreae may have weighed up to 140 kg, revealing this species as the largest carnivorous mammal in Europe at that time. Besides its very large size, K. langebadreae possesses an interesting combination of primitive and derived features. The distinctive skull morphology of K. langebadreae reflects a powerful bite force. The postcranial elements, which are rarely associated with hyainailourine specimens, indicate an animal capable of a plantigrade stance and adapted for terrestrial locomotion. We performed the first phylogenetic analysis of hyainailourines to determine the systematic position of K. langebadreae and to understand the evolution of the group that includes other massive carnivores. The analysis demonstrates that Hemipsalodon, a North American taxon, is a hyainailourine and is closely related to European Paroxyaena. Based on this analysis we hypothesize the biogeographic history of the Hyainailourinae. The group appeared in Africa with a first migration to Europe during the Bartonian that likely included the ancestors of Kerberos, Paroxyaena and Hemipsalodon, which further dispersed into North America at this time. We propose that the hyainailourines dispersed into Europe also during the Priabonian. These migrants have no ecological equivalent in Europe during these intervals and likely did not conflict with the endemic hyaenodont proviverrines. The discovery of K. langebadreae shows that large body size appears early in the evolution of hyainailourines. Surprisingly, the late Miocene Hyainailouros shares a more recent common ancestor with small-bodied hyainailourines (below 15 kg). Finally, our study supports a close relationship between the Hyainailourinae and Apterodontinae and we propose the new clade: Hyainailouridae.
The early Middle Eocene locality of Grube Messel, near Darmstadt (Germany), is famous for its complete vertebrate skeletons. The degree of preservation of soft tissues, such as body silhouettes, internal organs and gut contents, is frequently remarkable. The present specimen was analyzed for remnants of the reproductive system. Classic anatomy and osteology and high-resolution micro-x-ray were applied to describe the fetus of the European Eocene equoid Eurohippus messelensis. Scanning electronic microscopy (SEM) was used for determination of soft tissue remnants. The fetus is the earliest and best-preserved fossil specimen of its kind. The postcranial fetal skeleton is almost complete and largely articulated, allowing the conclusion that the pregnant mare was in late gestation. The apparent intrauterine position of the fetus is normal for the phase of pregnancy. Death of mare and fetus were probably not related to problems associated with parturition. Soft tissue interpreted as the uteroplacenta and a broad uterine ligament are preserved due to bacterial activity and allow considerations on the evolutionary development of the structures.
Nach den Kräften, welche die Oberflächenformen schufen, sondere ich das Diluvium im Westen der Weser nach horizontaler Richtung in ein glaciales, glacialfluviatiles und fluviatiles Diluvium. Von diesen drei Bezirken ist das glacial-fluviatile als eine nur unvollkommen zu begrenzende Zwischenzone zwischen dem nördlichen, glacialen und dem südlichen, fluviatilen Diluvium belegen.
Meine vergleichenden Untersuchungen über das Diluvium im Westen der Weser führten mich betreffs der Gliederung desselben zu Ergebnissen, welche mit den Anschauungen der holländischen Autoren nur teilweise sich decken. So weichen unsere Ansichten über das Alter derjenigen Flussablagerungen voneinander ab, aus denen südlich der Vecht die Mehrzahl der Höhen aufgebaut ist. Auch hinsichtlich der Stellung des "Sanddiluviums" kann ich jenen Forschern nicht in allen Punkten beipflichten. Ferner vermisse ich bei ihnen Angaben über das Vorhandensein einer Innenmoräne, und endlich, glaube ich, ist die Nomenklatur, welche in der niederländischen Diluvialliteratur sich eingebürgert hat, keineswegs einwandfrei. Da ich im Rahmen meiner vorigen Arbeit die Ansichten anderer Autoren nicht in erschöpfender Weise berücksichtigen konnte, so sei es meine Aufgabe, das dort unterlassene hier nachzuholen.
Die Ostrakoden bilden eine der niederen Ordnungen der Krebstiere. Der Panzer ist in eine zweiklappige Schale umgewandelt und erinnert so an die Schale der Muscheltiere. Die Systematik hat sich zwar bei Aufstellung der Gruppen und Gattungen von inneren Merkmalen des Tierkörpers leiten lassen, aber der Bau der Schale geht so sehr mit dem der Weichteile Hand in Hand, dass es im allgemeinen möglich, in den meisten Fällen verhältnismäßig leicht ist, die Tiere nach der Schale allein zu bestimmen, ein Umstand, der für die geologischen Funde von der größten Bedeutung ist. Ostrakoden finden sich nämlich in fast allen geologischen Formationen, welche überhaupt organische Überreste führen. In unseren nordischen Silur Geschieben sind sie beispielsweise von den Herren R. Jones und A. Krause zur Genüge nachgewiesen. Häufiger kommen sie bereits in der Steinkohlenformation vor, wie besonders die Arbeiten des Herrn Professors T. R. Jones dartbun. Am häufigsten treten sie jedoch in der Kreide und besonders im Tertiär auf. Aus dem nordwestdeutschen Tertiär habe ich z. B. 96 Species beschrieben.
Beitrag zur Kenntniss der Foraminiferen-Fauna des Ober-Oligocäns vom Doberg bei Bünde : Theil 2
(1895)
Über Prestwichia rotundata H. Woodw. sp. aus der Steinkohlenformation des Piesberges bei Osnabrück
(1885)
Schon früher ist von mir das Vorkommen der Gattung Prestwichia in der Steinkohlenformation des Piesberges genauer besprochen. Damals suchte ich nachzuweisen, dass die in den hangenden Schieferthonen des Flötz Mittel gefundenen fossilen Krebse wegen ihrer unbeweglich miteinander verwachsenen Rumpf- und Hinterleibs- Segmente zu jener Xiphosuren - Gattung zu stellen seien, und dass von den Species derselben die grösste Verwandtschaft mit den Piesberger Exemplaren Prestwichia rotundata H. Woodw. sp. besässe. Da aber damals von dieser Art nur die Abbildung von Prestwich vorlag und eine genauere Beschreibung erst noch aus der Feder Woodward's zu erwarten war, so liess ich es unentschieden, ob die scheinbar abweichende Ausbildung der Glabella bei dem einen Exemplare die Aufstellung einer besonderen Species erforderte. Nachdem jetzt schon vor längerer Zeit der betreffende Teil der Woodward'schen Monographie über die Merostomata, in welcher die Xiphosuren beschrieben, erschienen ist, glaube ich diese Frage zum Abschluss bringen zu können.
Im Frühling dieses Jahres machte mich der leider seitdem verstorbene Conducteur Peters hier darauf aufmerksam, dass in einem Muschelkalk-Steinbruche, östlich der sogenannten Quellenburg, Versteinerungen ziemlich häufig vorkämen. Da bekanntlich die Muschelkalkschichten der nächsten Umgebung hiesiger Stadt dem mittleren Muschelkalke (dem Dolomit) angehören und sich durch einen gänzlichen Mangel an Versteinerungen auszeichnen; so war mir die erhaltene Mittheilung äußerst wichtig und versäumte ich nicht, den betreffenden Steinbruch einer näheren Untersuchung zu unterziehen.
In dieser Arbeit geht es um die Korallen Micrabacia senoniensis und Cyclabacia Fromenteli. Diese wurden in der sog. Böllerts-Kuhle in der Honschaft Speldorf unweit Mühlheim a. d. Ruhr in Schichten gefunden. Weiterhin werden noch die Korallen Thamnastraea tenuissima, Dimorphastraea cf. parallela und Dimorphastraea Deickei, welche in einem Steinbruche in der Stadt Mühlheim selbst gefunden wurden, beschrieben. Die letzten genannten drei Specien lagen über dem Kohlensandsteine und sind einem an Glaukonit sehr reichen, weissen Pläner eingebacken.
Die betreffenden Schichten gehören ohne Zweifel zur Tourtia. Aus derselben waren bis jetzt aus Westphalen nur drei Species von Korallen bekannt, nämlich Synhclia gibbosa Münst. sp., Micrabacia coronula Goldf. sp. und Anthophyllum conicum Reuss. (s. Bölsche, Korallen des unteren Pläners im sächsischen Elbthale in: Geinitz, Elbthalgebirge, Dd. I, p. 5G).
Als ich in diesem Jahre meine Sammlung einer genaueren Durchsicht unterwarf, fand ich, dass in derselben genügendes Material vorhanden war, um dasselbe für einen kleineren Beitrag zur Paläontologie der Juraformation verwerten zu können. Nicht allein zeigte es sich, dass manche Species, wenn auch schon beschrieben, doch von neuen Fundorten vorlagen, und dass andere noch ganz neu für die jurassischen Schichten des nordwestlichen Deutschland waren, sondern die Untersuchung des vorliegenden Materials gab auch für die Kenntnis und Auffassung mancher bekannter Species neue Gesichtspunkte an die Hand. In dem folgenden ersten Theile, der Versteinerungen des unteren und teilweise des mittleren Jura (bis zu den Coronaten - Schichten inclus.) behandelt, habe ich in Bezug des geognostischen Vorkommen der Species die Einteilung des Jura den Werken von Brauns (der untere Jura im nordwestlichen Deutschland 1871, der mittlere Jura 1869) entnommen. Um eine spätere Kritik zu ermöglichen, ist bei den Species das Werk angegeben, nach dem dieselben bestimmt sind. Von Fundorten sind meistens nur die neuen angeführt.
This study demonstrates that alkaline earth elements in enamel of hippopotamids, in particular Ba and Sr, are tracers for water provenance and hydrochemistry in terrestrial settings. The studied specimens are permanent premolar and molar teeth found in modern and fossil lacustrine sediments of the Western Branch of the East African Rift system (Lake Kikorongo, Lake Albert, and Lake Malawi) and from modern fluvial environments of the Nile River.
Concentrations in enamel vary by two orders of magnitude for Ba (120–9336 μg g−1) as well as for Sr (9–2150 μg g−1). The variations are partially induced during post-mortem alteration and during amelogenesis, but the major contribution originates ultimately from the variable water chemistry in the habitats of the hippopotamids which is controlled by the lithologies and weathering processes in the watershed areas. Amelogenesis causes a distinct distribution of MgO, Ba and Sr in modern and fossil enamel, in that element concentrations increase along profiles from the outer rim towards the enamel–dentin junction by a factor of 1.3–1.9. These elements are well correlated in single specimens, thus suggesting that their distribution is determined by a common, single process, which can be described by closed system Rayleigh crystallization of bioapatite in vivo.
Enamel from most hippopotamid specimens has Sr/Ca and Ba/Ca which are typical for herbivores. However, Ba/Sr ranges from 0.1 to 3 and varies on spatial and temporal scales. Thus, Sr concentrations and Ba/Sr in enamel differentiate between habitats having basaltic mantle rocks or Archean crustal rocks as the ultimate sources of Sr and Ba. This provenance signal is modulated by climate change. In Miocene to Pleistocene enamel from the Lake Albert region, Ba/Sr decreases systematically with time from 2 to 0.5. This trend can be correlated with changes in climate from humid to arid, in vegetation from C3 to C4 biomass as well as with increasing evaporation of the lake water. The most plausible explanation is that Ba mobility decreased with increasing aridification due to preferential deposition with clay and Fe-oxide-hydroxide or barite on the watershed of Lake Albert.
Zwischen dem westlichen Teile des Teutoburger Waldes und der Weserkette findet sich eine Reihe tertiärer Hügel, die als einzelne Schollen von dem Ganzen der tertiären Ablagerungen dieser Gegend übrig geblieben sind. Hierzu gehört das Ober-Oligocän in der Bauerschaft Krevinghausen bei SchIedehausen und der bei weitem wichtigste dieser Hügel, der Doberg. Nach einleitenden Bemerkungen und einem Verzeichnis der Schriften, welche auf den Doberg Bezug nehmen, wird die oligozäne Fauna des Gebietes mit folgenden Kapiteln dargestellt: I. Vertebrata, Wirbeltiere. II. Cephalopoda, Kopffüsser. III. Glossophora, Schnecken. IV. Lamellibranchiata, Muscheln. V. Brachiopoda, Armfüsser. VI. Crustacea, Krustentiere. VII. Annelida, Ringelwürmer. VIII. Echinoidea, Seeigel. IX. Bryozoa, Moostierchen. X. Anthozoa, Korallen. XI. Foraminiferen. XII. Algen (1 Art).
Aus einer verlassenen Mergelgrube am Rubbenbrucher See westlich von Osnabrück werden 61 Foraminiferen-, 10 Bryozoen- und 22 Ostrakoden-Arten namhaft gemacht. Diese Funde, zusammen mit den Resten anderer Organismen, ergeben als Biotop ein ufernahes Litoral des Oberoligozän-Meeres, vergleichbar mit den klassischen Vorkommen von Astrup und Bünde.
It has been forty years since the Jurassic site of Veillon in Talmont-Saint-Hilaire (Vendée, France), where footprints of vertebrates were found, was plundered. This outstanding site belonging to Vendée's geological patrimony needs to be preserved and developed. Some footprints from Veillon are displayed in the collections of about thirty public institutions and six museums.
Using the fossil dinosaur footprints of the lower Lias of Le Veillon at Talmont-Saint-Hilaire, (Vendée) as a starting point, the reconstitution by pupils of a unique fossil-bearing site is under consideration. The paleobiology and paleoecology of reptiles in this geological site are dealt with.
Palichnites et muséologie
(2003)
After they had intrigued the inquiring or observant mind for decades, it was realised that certain "natural curiosities" were tracks of extinct creatures, related to gigantic birds or other antediluvian animals. At the beginning of the XIXth century a new science, ichnology, came into being and developed, thanks to the collection, preservation and study of fossil prints. The increasing quantity of evidence thus unearthed reveals the diversity of vertebrate faunas which have trod the earth’s surface and left behind valuable information about their characteristics and behaviour. Like other institutions labouring on behalf of conservation, knowledge of nature and the preservation of our natural heritage, the Nantes Natural History Museum has, for nearly two centuries, collected, conserved, displayed and brought to the attention of the public those valuable pieces of evidence, fossil footprints. Displays of fossil prints are regularly presented during different exhibitions. Examples of them permit one to follow the evolution of our knowledge of these prints and illustrate how the information yielded by these prints has been transmitted.
The Ichnospace is a specific exhibition dedicated to traces and tracks left by recent and fossil animals on ground. The Ichnospace is located at Luzech, a little town with a long historic past, near Cahors (about 17 km W) in the valley of the river Lot. Every living being leaves traces of its activities (habitat, locomotion, diet, etc.) in its environment. These traces are the signing of the individual and they may be preserved and fossilised in geological time. They provide information about biodiversity, the habits of individuals and their distribution in time and space. In the Causses of Quercy area very near of Luzech and well known by palaeontologists and geologists, many dinosaur and pterosaur tracks have been discovered. Some of them are exposed with various fossil invertebrates, turtles, crocodiles and so on, from upper Jurassic when the sea Tethys ran along shores of the large Central island. Movie and an audio-visual show complete the exhibited specimens with French and English comments.
Hettangian stratotype shows an example of preservation and valorisation of a scientifical historical place. Earth Story is understood thanks to geological heritage; this heritage is very vulnerable. Judicial framework and mentality evolution allow us to make a conservation of particularly interesting places.
The Liassic Talmont-Saint-Hilaire (Vendée, France) outcrop has been studied here according to the abundance of fossil plants of the Cheirolepidiaceous family (Gymnosperms, Coniferales). A new species of Brachyphyllum has been studied in light microscopy, even in scanning electron microscopy and transmission electron microscopy. It is, at that time, one of the most complete study realized on a fossil taxon. Epidermal cells, subsidiary cells and guard cells have been described on both adaxial and abaxial surfaces, in transversal and longitudinal sections. Cuticle ultrastructural variations have been observed between epidermal and stomatal cells. To conclude, we propose a discussion about the cuticle structure variations related to environmental conditions and the functional morphology of the different cells.
Analyse d'un exemple de fossilisation d'une trace de pas de Dinosaure (Lias inférieur des Causses)
(2003)
A calcareous block made of algo-laminated (stromatolitic) material exhibits at its upper surface a foot print of a Dinosaurian. A vertical section (sawing) and a thin section allow to make detailed observations. The early diagenesis permits the preservation of the deformations caused by the foot print.
In the inter/supratidal algal mats, the early diagenesis products a progressive induration. It is enough to preserve all the deformations affecting the mats, including footprints of vertebrates, tracks and burrows of small invertebrates, and physical actions (desiccation cracks).
Comparaison des ichnopopulations des Grands Causses (Sud de la France) et du Veillon (Vendée)
(2003)
From descriptive statistic method concerning length (L), width (W) and L/W, it is suggested that Grallator variabilis samples from Vendée (Le Veillon) and Grands Causses belong to the same ichnospecies. Also, by this way, G. maximus of Hettangian of Vendée could be synonym of G. minusculus from USA Hettangian.
Numerous footprints areas has been found, since 1962, in Permian, Triassic and Hettango-Sinemurian. All show precised oriented trackways. So that they were undestood like walking ways but their precise meaning is still enigmatic.
Footprints are numerous in France where they found from Carboniferous to Quaternary levels. Those of Middle Triassic (Anisian-Ladinian) are especialy interesting because they are tridactyl footprints made probably by first dinosaurs. First skelettons are known from Upper Ladinian / lower Carnian.
The Veillon ichnofauna, early Liassic in age, includes various Reptile taxa: quadruped Pseudosuchian, Coelurosaurians, Theropods, primitive Iguanodon – like Ornithopods and some unspecified forms. This ichnofauna is very comparable to the early Liassic footprint assemblage of the Connecticut.
Clastic deposits related to alluvial and estuarine environments sedimented during the early Liassic in the Veillon area (south of Talmont-Saint-Hilaire, Vendée, France). A reptile fauna including various taxa, just known by innumerable footprints, lived in that environment, in rather hot and dry climatic conditions.
On the intertidal zone of Le Veillon at Talmond-Saint-Hilaire (Vendée, France), in 1963 Gilbert Bessonnat discovered traces of vertebrate footprints in a Hettangian formation. On March 28th & 29th, at this site a study session was held on the theme: "sites with vertebrate footprints on the Triassic-Jurassic limit". Palæontologists, palæobotanists, sedimentologists, hydrologists, scientific historians and naturalists compared their results and projected further research. This exceptional Vendée heritage site is to be protected and developed.
Anhand mikrolazieller, paläontologischer und sedimentologischer Untersuchungen wird die tempestitische Natur des oberen Teils der Zwischenbank (Mittelkimmeridge VII, Oberer Jura) am Kalkrieser Berg (SW-Niedersachsen) dargelegt. An der Oberkante der Kalkbank ist ein hardground ausgebildet, der erosiv Ireigelegt und anschließend besiedelt wurde. Das Milieu stellt sich als Ilachmarin und küsten nah dar. Die Arbeit ist die erste Beschreibung der Genese eines deutschen jurassischen hardgrounds s. s. und der erste Nachweis eines kalkigen oberjurassischen Tempestits.
Unter den Geschieben des Kies-Sand-Rückens ,Laer-Heide' (Landkreis Osnabrück) fand sich ein Geschiebe des ,grünlich-grauen Graptolithengesteins' mit Calymene orthomarginata SCHRANK 1970 und Leonaspis mutica (EMMRICH 1844) aus dem baltischen Silur. Es stammt vermutlich aus dem Gebiet der "North Mid-Sea Bank" SSW von Gotland und ESE der Südspitze Ölands. Stratigraphisch ist es dem Bereich des Unterwenlock bis unteren Oberwen lock zuzuordnen. Nach einem kurzen Überblick über den Fossilinhalt des Geschiebes und der systematischen Einordnung der gefundenen Trilobitentaxa werden folgende Larvalstadien von Trilobiten beschrieben: 1 Paraprotaspis-Stadium eines Odontopleuriden, das wahrscheinlich Leonaspis mutica zuzuordnen ist, 1 nahezu vollständiges frühes Meraspis-Stadium und eine Reihe frühmeraspider und spätmeraspider Cranidien und Pygidien, die Calymene orthomarginata zugeordnet werden. Abschließend werden einige Beobachtungen zur Ontogenie von Calymene orthomarginata während der meraspiden Phase dargelegt.
Der erste Insektenrest (linker Vorder- und Hinterflügel) aus Gesteinen des Westfalium D (Ober-Karbon = Silesium) vom Piesberg N'Osnabrück wird als Erasipterella piebergensis n. g. n. sp. beschrieben und abgebildet. Er gehört in die nächste Verwandtschaft der ältesten bekannten Libellen-Gattung Erasipteron (Ordnung Odonata; Unterordnung Meganisoptera), unterscheidet sich von dieser aber vornehmlich durch die wesentlich kleineren Ausmaße, den weniger schlanken Flügel-Umriß und die stärkere Auflösung der hinteren Anal- Adern in Zellen-Reihen. In dem letztgenannten Merkmal ist Erasipterella n. g. weniger ursprünglich als ihre vermutliche Ahnenform Erasipteron und weist schon in Richtung moderner Typen der Odonata. Das Exemplar vom Pies berg ist der zweite Nachweis dieser Unterordnung aus dem westdeutschen Ober-Karbon.
Xiphosuriden (Schwertschwänze) sind in einigen Fazies-Bereichen des Ober- Karbons (Silesium) recht weit verbreitet und an einigen wenigen FundsteIlen sogar häufig. Im paralisehen Ober-Karbon West-Deutschlands sind sie jedoch sehr selten. Vor kurzem ist nun ein neues Opisthosoma der Xiphosuren-Gattung Euproops in Gesteinen des Westfalium 0 vom Piesberg nördlich Osnabrücks gefunden worden, das wahrscheinlich konspezifisch ist mit den 4 schon früher an derselben Lokalität aufgesammelten Stücken (BÖLSCHE 1875; BOEKSCHOTEN 1968). Die Erhaltung aller Stücke vom Piesberg erlaubt jedoch keine genauere Bestimmung als Euproops sp. In diesem Zusammenhang wird auch darauf hingewiesen, daß noch erhebliche Schwierigkeiten hinsichtlich der infragenerischen Systematik von Euproops bestehen, da noch nicht genügend bekannt ist über die spezifische Variabilität und Ontogenie dieser merkmalsarmen Gattung. Nach den neuen Untersuchungen über die Lebensweise von Euproops danae durch FISHER (1979) dürfte auch die Euproops-Form vom Piesberg weitgehend subaerisch gelebt haben.
An der Basis einer Brekzie aus zechsteinzeitlichen Karbonatgesteinen konnten auf dem Hüggel-Horst Sedimente mit mariner Fauna (Ober-Miozän: Langenfelde-Gram) beobachtet werden. Ihre Fossilführung und Petrographie wird beschrieben. Sie sind in Höhlen, die bei Subrosion von Zechstein-Sulfaten entstanden, im küstennahen Bereich des Miozän-Meeres eingespült und abgelagert worden. Sie belegen, daß der Hüggel-Horst schon im Miozän gehoben und bis auf die Zechstein-Gesteine abgetragen worden war.
Eine Lagerstätte kreidezeitlicher und paläogener Chondrichthyes-Reste bei Fürstenau (Niedersachsen)
(1979)
Aus einer durch Glaziotektonik verstellten Kiesbank innerhalb einer Folge mariner Sedimente bei Fürstenau wird eine 49 Arten umfassende Liste von Chondrichthyes genannt. Diese Fauna enthält Arten des Campans, Oberpaläozäns und Unter- bis Mitteleozäns. Die Entstehung dieser Kiesbank könnte einerseits einer Transgressionsphase oder andererseits einer Kondensation verbunden mit der Erosion von Folgen des oben genannten Alters während des Lediums zugeschrieben werden. Diese Kiesbank besitzt in Oldenburg und im Emsland eine weite Verbreitung.
Die Knochenansammlung im grauen vulkanischen Tuff der Südserengeti gibt als ökologisch unmögliches
Gemisch ein gutes Abbild des Gesamtbestandes und des Lebensraumes der altquartärcn ost- und innerafrikanischen Fauna. Diese lebte formenreich in Urwald, Savanne und offener Steppe. Das Fehlen wasserlebender Tiere ist hier auf örtliche Umstände zurückzuführen: die vulkanischen Aschen gingen auf Steppenboden nieder, Die benachbarten, ungefähr gleichalten Knochenlager enthalten solche Tiere. Neu ist an der Serengetifauna der bereits beträchtliche Anteil von Kleinsäugern (Nager; Insektivoren fehlen noch). Diese wird weitere Forschung vermehren. Die klimatische Entsprechung der Fauna wird in tropischen, feuchtwarmen Bedingungen erblickt. Obwohl viele tertiäre Formen enthaltend, wird die Fauna nicht als jungtertiär angesehen, sondern wegen des Auftretens moderner· Formen als eine Tiergesellschaft, welche das Quartär eröffnet. Als Leitfossil für dessen Beginn wird der Gattung Archidiskodon, aus welcher die echten Elefanten, darunter auch der afrikanische (Loxodonta africana ) entstanden sind, vor den Equiden der Vorzug gegeben.
Ostafrika, das ja als Tierparadies schlechthin gilt, lebte bis vor kurzem noch im Quartär. Das Schrifttum über seine Tierwelt scheint zwar fast unermeßlich groß, aufs Ganze gesehen ist das Wissen weder tief noch auch nur oberflächlich vollständig. Die Hauptleistung des "weißen Mannes" bestand in der Störung und Vernichtung der Fauna. Aber die Natur ist groß; sie hält noch einen Schatz im Inneren ihrer Gebirge bereit, damit der Mensch seine Stellung zu ihr und den Sinn seines Lebens ergründe, einen von vielen: dle quartäre Lebewelt selbst. Möge die hohe Aufgabe, ihn zu heben, uns Deutschen vergönnt sein! Eine Probe hat Dr. KOHL-LARSEN gesichert.
Fossile Libellen sind eine echte paläontoloqische Rarität. Wenn man außerdem noch von ihnen erwartet, daß sie besonders gut erhalten sind, dann werden sie zu wahren wissenschaftlichen Kostbarkeiten. Sowohl der Paläontologe als auch der Zoologe sind deshalb gleichermaßen angetan von dem einzigartigen Fossilmaterial (Abb. 1), das sich durch viele Seitenblicke auf die Formenvielfalt rezenter Libellen, ihre Morphologie, ihre Lebensweise, kurzum ihre gesamte Biologie, wieder "zu neuem Leben erwecken" läßt.
Aus dem Obercampan von Münster und Umgebung werden neben 5 biostratigraphisch leitenden Foraminiferen noch 9 Arten beschrieben und abgebildet, die bisher in N-Deutschland nur aus der höchsten Oberkreide und dem Dan publiziert worden sind. Viele der letztgenannten Foraminiferen sind in den Ablagerungen der Tethys häufig, sogar in der Oberkreide unterhalb des Maastricht.
An Hand der im einzelnen dargelegten Anschauungen der pflanzlichen Gesellschaftslehre der Neobotanik wird das Vorhaben verfolgt, gleichartige Wege für die Beantwortung soziologischer Fragen bei fossilen Floren auf paläobotanischer Grundlage zu finden. Geologisch-historisch ergeben sich dabei zwei Probleme, welche dem Neobotaniker als Zeitgenossen seiner Objekte unbekannt sind und für ihn aus der heute vorliegenden Situation gar nicht entstehen können. Um paläobotanische Soziologie überhaupt betreiben zu können, sind für die Realität einer fossilen Pflanzengemeinschaft erst zwei Voraussetzungen zu gewinnen: die Synchronie, die regionale Gleichaltrigkeit der Gesteinsschicht, in oder auf der die fossile Flora bis heute konserviert wurde, der Individuums beg riff, der erst auf Grund der jetzigen Fundumstände geprägt werden kann. Unter Auswertung von Untertage-Vorkommen von fossilreichen Pflanzenschichten im gefalteten Westfal A und B des mittleren und nördlichen Ruhrgebietes wurde von zwei Mitarbeitern des Autors eine größere Anzahl von Synchronen ermittelt und unter Formung und Präzisierung des Individuumsbegriffes die Möglichkeit qeschaffen. die Paläosoziologie für einen Teil der oberkarbonischen Pflanzenwelt der euramerischen Floraprovinz zu erarbeiten.
Zum ersten Male wird berichtet über nicht-marine Muschelgesellschaften mit mehr als 170 Exemplaren. Die Mehrzahl stammt aus dem Hangenden des Flözes Dreibänke, einige Funde kommen aus dem Hangenden des Flözes Bänkchen und Zweibänke (Unteres Westfal D) des Piesberges. Die Muscheln gehören alle der Gattung Anthraconaia an und wurden unter fluvio-Iakustrinen Bedingungen abgelagert.
Ein "Taqesaufschluß", aufgeschürft bei Baggerarbeiten an der Straßenböschung E Niedermoschel/Pfalz, erbrachte den ersten Nachweis von Limuliden im saarpfälzischen Unter-Rotliegenden. Die Funde werden als neue Art beschrieben und einer neuen Gattung zugeordnet: Palatinaspis beimbaueri n.g. et n.sp. Eine aus dem Stefanium der Halleschen Mulde als Pringlia fritschi W. & R. REMY 1959 beschriebene Art wird in die neue Gattung einbezogen.
Während die klassische Darstellungsweise der Paläontologie stets vom Niederen zum Höheren und vom Alteren zum Jüngeren führt, entspricht die Fragestellung der paläontologischen Forschung einem vergleichenden Hin und Her zwischen den Lebensformen der Gegenwart und der Vergangenheit. Die folgende Darstellung versucht dem dadurch Ausdruck zu verleihen, daß die Gegenwart in ihr nicht nur als Ziel des Ausblicks, sondern auch als Ausgangspunkt der Betrachtung fungiert.
The first fossil cyphophthalmid harvestman (Opiliones: Cyphophthalmi) from Palaeogene (Eocene) Baltic amber is described. This is only the third fossil example of this basal harvestman lineage; the others being from the probably slightly younger Bitterfeld amber and the much older, early Cretaceous, Myanmar (Burmese) amber. Although incomplete and lacking most of the appendages, the new Baltic amber fossil can be identified as a female. The somatic characters preserved, especially spiracle morphology and the coxo-genital region, allow it to be assigned with some confidence to the extant genus Siro Latreille, 1796 (Sironidae). This fossil is formally described here as Siro balticus sp. nov. It resembles modern North American Siro species more than modern European ones, and can be distinguished principally on its relatively large size and the outline form of the body.
Ein neuer isolierter Insekten-Flügel aus dem Westfalium D vom Piesberg N' Osnabrück gehört zu einer neuen Art: Aspidothorax aestatis n. sp. (Megasecoptera: Aspidothoracidae). Diese ist nahe verwandt mit Aspidothorax triangularis BRONGNIART 1894 aus dem Stephanium B von Commentry (Zentral-Frankreich), unterscheidet sich aber vor allem durch (1) den weniger deutlich markierten Apex, (2) die breitere Praeradial-Area, (3) die ungeteilten Adern MP- und CuP- sowie (4) die unregelmäßiger angeordneten Queradern.
Die Faunenhorizonte der "Subfurcaten-Schichten" (Bajocium, Niortense-Zone) in Nordwestdeutschland
(1992)
Aufgrund neuer Aufschlüsse der "Subfurcaten-Schichten" (Bajocium, Niortense- Zone) in Nordwestdeutschland ist es möglich, die Schichtenfolge in Ammonitenfaunen-Horizonte zu untergliedern. Eine paläontologische Revision der Ammonitengattung Strenoceras HYATTmit den Untergattungen Strenoceras HYATT(m) und Garantiana HYATT(M) wird durchgeführt.
Aus ordovizischen Geschieben des Kies-Sand-Rückens ,Laer-Heide' (Landkreis Osnabrück; NW-Deutschland) werden 2 mißgebildete Pygidien von IlIaeniden (Trilobita) beschrieben: Ein vergleichsweise dünnschaliges Pygidium von lIIaenus jevensis HOLM 1886 weist eine Verletzung in Form eines dellenartigen Eindrucks auf. An einem Pygidium von Bumastus cf. nudus (ANGELIN 1854) läßt sich eine beulenförmige Verwachsung beobachten. Mögliche Entstehungsursachen beider Anomalien werden diskutiert.
Neue Insekten-Funde (Palaeodictyoptera: Breyeriidae) aus dem Ober-Karbon von Osnabrück (Deutschland)
(1995)
Zwei neue, zu den Palaeodictyoptera: Breyeriidae gehörende Insekten-Flügel aus Schichten des Westfalium 0 (Ober-Karbon) vom Piesberg nördlich Osnabrück werden beschrieben. Breyeria bistrata n.sp. ähnelt den beiden Arten B. rappi Carpenter, 1967 und B. barborae Kukalova, 1959, unterscheidet sich aber von diesen vor allem durch den fast geraden Flügel-Vorderrand und die größere Anzahl von Zweigen. Hasa/a inferiorsaxonica n.g. n.sp. ist gekennzeichnet durch die sehr lange, in den Vorderrand mündende ScP-, die Anwesenheit einer Stützader rp-ma und die deutlich ausgebildete sekundäre Korrugation.
We describe and analyze a Neandertal postcranial skeleton and dentition, which together show unambiguous signs of right-handedness. Asymmetries between the left and right upper arm in Regourdou 1 were identified nearly 20 years ago, then confirmed by more detailed analyses of the inner bone structure for the clavicle, humerus, radius and ulna. The total pattern of all bones in the shoulder and arm reveals that Regourdou 1 was a right-hander. Confirmatory evidence comes from the mandibular incisors, which display a distinct pattern of right oblique scratches, typical of right-handed manipulations performed at the front of the mouth. Regourdou's right handedness is consistent with the strong pattern of manual lateralization in Neandertals and further confirms a modern pattern of left brain dominance, presumably signally linguistic competence. These observations along with cultural, genetic and morphological evidence indicate language competence in Neandertals and their European precursors.
Yuanmou Basin of Yunnan, SW China, is a famous locality with hominids, hominoids, mammals and plant fossils. Based on the published megaflora and palynoflora data from Yuanmou Basin, the climate of Late Pliocene is reconstructed using the Coexistence Approach. The results indicate a warm and humid subtropical climate with a mean annual temperature of ca. 16–17°C and a mean annual precipitation of ca. 1500–1600 mm in the Late Pliocene rather than a dry, hot climate today, which may be due to the local tectonic change and gradual intensification of India monsoon. The comparison of Late Pliocene climate in Eryuan, Yangyi, Longling, and Yuanmou Basin of Yunnan Province suggests that the mean annual temperatures generally show a latitudinal gradient and fit well with their geographic position, while the mean annual precipitations seem to be related to the different geometries of the valleys under the same monsoon system.
Despite being internal organs, digestive structures are frequently preserved in Cambrian Lagerstätten. However, the reasons for their fossilisation and their biological implications remain to be thoroughly explored. This is particularly true with arthropods--typically the most diverse fossilised organisms in Cambrian ecosystems--where digestive structures represent an as-yet underexploited alternative to appendage morphology for inferences on their biology. Here we describe the phosphatised digestive structures of three trilobite species from the Cambrian Weeks Formation Lagerstätte (Utah). Their exquisite, three-dimensional preservation reveals unique details on trilobite internal anatomy, such as the position of the mouth and the absence of a differentiated crop. In addition, the presence of paired pygidial organs of an unknown function is reported for the first time. This exceptional material enables exploration of the relationships between gut phosphatisation and the biology of organisms. Indeed, soft-tissue preservation is unusual in these fossils as it is restricted to the digestive structures, which indicates that the gut played a central role in its own phosphatisation. We hypothesize that the gut provided a microenvironment where special conditions could develop and harboured a source of phosphorus. The fact that gut phosphatization has almost exclusively been observed in arthropods could be explained by their uncommon ability to store ions (including phosphorous) in their digestive tissues. However, in some specimens from the Weeks Formation, the phosphatisation extends to the entire digestive system, suggesting that trilobites might have had some biological particularities not observed in modern arthropods. We speculate that one of them might have been an increased capacity for ion storage in the gut tissues, related to the moulting of their heavily-mineralised carapace.
For reconstructing environmental change in terrestrial realms the geochemistry of fossil bioapatite in bones and teeth is among the most promising applications. This study demonstrates that alkaline earth elements in enamel of Hippopotamids, in particular Ba and Sr are tracers for water provenance and hydrochemistry. The studied specimens are molar teeth from Hippopotamids found in modern and fossil lacustrine settings of the Western Branch of the East African Rift system (Lake Kikorongo, Lake Albert, and Lake Malawi) and from modern fluvial environments of the Nile River.
Concentrations in enamel vary by ca. two orders of magnitude for Ba (120–9336 μg g−1) as well as for Sr (9–2150 μg g−1). Concentration variations in enamel are partly induced during post-mortem alteration and during amelogenesis, but the major contribution originates from the variable water chemistry in the habitats of the Hippopotamids which is dominated by the lithologies and weathering processes in the watershed areas. Amelogenesis causes a distinct distribution of Ba and Sr in modern and fossil enamel, in that element concentrations increase along profiles from the outer rim towards the enamel-dentin junction by a factor of 1.3–1.5. These elements are well correlated with MgO and Na2O in single specimens, thus suggesting that their distribution is determined by a common, single process. Presuming that the shape of the tooth is established at the end of the secretion process and apatite composition is in equilibrium with the enamel fluid, the maturation process can be modeled by closed system Rayleigh crystallization.
Enamel from many Hippopotamid specimens has Sr/Ca and Ba/Ca which are typical for herbivores, but the compositions extend well into the levels of plants and carnivores. Within enamel from single specimens these element ratios covary and provide a specific fingerprint of the Hippopotamid habitat. All specimens together, however, define subparallel trends with different Ba/Sr ranging from 0.1 to 3. This ratio varies on spatial and temporal scales and traces provenance signals as well as the fractionation of the elements in the hydrological cycle. Thus, Sr concentrations and Ba/Sr in enamel differentiate between habitats having basaltic or Archean crustal rocks as the ultimate sources of Sr and Ba. The provenance signal is modulated by climate change. In Miocene to Pleistocene enamel from the Lake Albert region, Ba/Sr decreases systematically with time from about 2 to 0.5. This trend can be correlated with changes in climate from humid to arid in vegetation from C3 to C4 biomass as well as with increasing evaporation of the lake water. The most plausible explanation is that with time, Ba mobility decreased relative to that of Sr. This can arise if preferential adsorption of Ba to clay and Fe-oxide-hydroxide is related to increasing aridification. Additionally, weathering solutions and lake water can become increasingly alkaline and barite becomes stable. In this case, Ba will be preferentially deposited on the watershed of Lake Albert and rivers with low Ba/Sr will feed the habitats of the Hippopotamids.
Orangutans (Pongo) are the only great ape genus with a substantial Pleistocene and Holocene fossil record, demonstrating a much larger geographic range than extant populations. In addition to having an extensive fossil record, Pongo shows several convergent morphological similarities with Homo, including a trend of dental reduction during the past million years. While studies have documented variation in dental tissue proportions among species of Homo, little is known about variation in enamel thickness within fossil orangutans. Here we assess dental tissue proportions, including conventional enamel thickness indices, in a large sample of fossil orangutan postcanine teeth from mainland Asia and Indonesia. We find few differences between regions, except for significantly lower average enamel thickness (AET) values in Indonesian mandibular first molars. Differences between fossil and extant orangutans are more marked, with fossil Pongo showing higher AET in most postcanine teeth. These differences are significant for maxillary and mandibular first molars. Fossil orangutans show higher AET than extant Pongo due to greater enamel cap areas, which exceed increases in enamel-dentine junction length (due to geometric scaling of areas and lengths for the AET index calculation). We also find greater dentine areas in fossil orangutans, but relative enamel thickness indices do not differ between fossil and extant taxa. When changes in dental tissue proportions between fossil and extant orangutans are compared with fossil and recent Homo sapiens, Pongo appears to show isometric reduction in enamel and dentine, while crown reduction in H. sapiens appears to be due to preferential loss of dentine. Disparate selective pressures or developmental constraints may underlie these patterns. Finally, the finding of moderately thick molar enamel in fossil orangutans may represent an additional convergent dental similarity with Homo erectus, complicating attempts to distinguish these taxa in mixed Asian faunas.
Background: Mammalian fossils from the Eppelsheim Formation (Dinotheriensande) have been a benchmark for Neogene vertebrate palaeontology since 200 years. Worldwide famous sites like Eppelsheim serve as key localities for biochronologic, palaeobiologic, environmental, and mammal community studies. So far the formation is considered to be of early Late Miocene age (~9.5 Ma, Vallesian), representing the oldest sediments of the Rhine River. The stratigraphic unity of the formation and of its fossil content was disputed at times, but persists unresolved.
Principal Findings: Here we investigate a new fossil sample from Sprendlingen, composed by over 300 mammalian specimens and silicified wood. The mammals comprise entirely Middle Miocene species, like cervids Dicrocerus elegans, Paradicrocerus elegantulus, and deinotheres Deinotherium bavaricum and D. levius. A stratigraphic evaluation of Miocene Central European deer and deinothere species proof the stratigraphic inhomogenity of the sample, and suggest late Middle Miocene (~12.5 Ma) reworking of early Middle Miocene (~15 Ma) sediments. This results agree with taxonomic and palaeoclimatic analysis of plant fossils from above and within the mammalian assemblage. Based on the new fossil sample and published data three biochronologic levels within the Dinotheriensand fauna can be differentiated, corresponding to early Middle Miocene (late Orleanian to early Astaracian), late Middle Miocene (late Astaracian), and early Late Miocene (Vallesian) ages.
Conclusions/Significance: This study documents complex faunal mixing of classical Dinotheriensand fauna, covering at least six million years, during a time of low subsidence in the Mainz Basin and shifts back the origination of the Rhine River by some five million years. Our results have severe implications for biostratigraphy and palaeobiology of the Middle to Late Miocene. They suggest that turnover events may be obliterated and challenge the proposed ‘supersaturated’ biodiversity, caused by Middle Miocene superstites, of Vallesian ecosystems in Central Europe.
Bioapatite in mammalian teeth is readily preserved in continental sediments and represents a very important archive for reconstructions of environment and climate evolution. This project provides a comprehensive data base of major, minor and trace element and isotope tracers for tooth apatite using a variety of microanalytical techniques. The aim is to identify specific sedimentary environments and to improve our understanding on the interaction between internal metabolic processes during tooth formation and external nutritional control and secondary alteration effects. Here, we use the electron microprobe to determine the major and minor element contents of fossil and modern molar enamel, cement and dentin from Hippopotamids. Most of the studied specimens are from different ecosystems in Eastern Africa, representing modern and fossil lacustrine (Lake Kikorongo, Lake Albert, and Lake Malawi) and modern fluvial environments of the Nile River system. Secondary alteration effects - in particular FeO, MnO, SO3 and F concentrations – are 2 to 10 times higher in fossil than in modern enamel; the secondary enrichment of these components in fossil dentin and cement is even higher. In modern and fossil enamel, along sections perpendicular to the enamel-dentin junction (EDJ) or along cervix-apex profiles, P2O5 and CaO contents and the CaO/P2O5 ratios are very constant (StdDev ∼1%). Linear regression analysis reveals tight control of the MgO (R2∼0.6), Na2O and Cl variation (for both R2>0.84) along EDJ-outer enamel rim profiles, despite large concentration variations (40% to 300%) across the enamel. These minor elements show well defined distribution patterns in enamel, similar in all specimens regardless of their age and origin, as the concentration of MgO and Na2O decrease from the enamel-dentin junction (EDJ) towards the outer rim, whereas Cl displays the opposite trend. Fossil enamel from Hippopotamids which lived in the saline Lake Kikorongo have a much higher MgO/Na2O ratio (∼1.11) than those from the Neogene fossils of Lake Albert (MgO/Na2O∼0.4), which was a large fresh water lake like those in the western Branch of the East African Rift System today. Similarly, the MgO/Na2O ratio in modern enamel from the White Nile River (∼0.36), which has a Precambrian catchment of dominantly granites and gneisses and passes through several saline zones, is higher than that from the Blue Nile River, whose catchment is the Neogene volcanic Ethiopian Highland (MgO/Na2O∼0.22). Thus, particularly MgO/Na2O might be a sensitive fingerprint for environments where river and lake water have suffered strong evaporation. Enamel formation in mammals takes place at successive mineralization fronts within a confined chamber where ion and molecule transport is controlled by the surrounding enamel organ. During the secretion and maturation phases the epithelium generates different fluid composition, which in principle, should determine the final composition of enamel apatite. This is supported by co-linear relationships between MgO, Cl and Na2O which can be interpreted as binary mixing lines. However, if maturation starts after secretion is completed, the observed element distribution can only be explained by equilibration of existing and addition of new apatite during maturation. It appears the initial enamel crystallites precipitating during secretion and the newly formed bioapatite crystals during maturation equilibrate with a continuously evolving fluid. During crystallization of bioapatite the enamel fluid becomes continuously depleted in MgO and Na2O, but enriched in Cl which results in the formation of MgO, and Na2O-rich, but Cl-poor bioapatite near the EDJ and MgO- and Na2O-poor, but Cl-rich bioapatite at the outer enamel rim. The linkage between lake and river water compositions, bioavailability of elements for plants, animal nutrition and tooth formation is complex and multifaceted. The quality and limits of the MgO/Na2O and other proxies have to be established with systematic investigations relating chemical distribution patterns to sedimentary environment and to growth structures developing as secretion and maturation proceed during tooth formation.
Bioapatite in mammalian teeth is readily preserved in continental sediments and represents a very important archive for reconstructions of environment and climate evolution. This project intends to provide a detailed data base of major, minor and trace element and isotope tracers for tooth apatite using a variety of microanalytical techniques. The aim is to identify specific sedimentary environments and to improve our understanding on the interaction between internal metabolic processes during tooth formation and external nutritional control and secondary alteration effects. Here, we use the electron microprobe, to determine the major and minor element contents of fossil and modern molar enamel, cement and dentin from hippopotamids. Most of the studied specimens are from different ecosystems in Eastern Africa, representing modern and fossil lakustrine (Lake Kikorongo, Lake Albert, and Lake Malawi) and modern fluvial environments of the Nile River system.
Secondary alteration effects in particular FeO, MnO, SO3 and F concentrations, which are 2 to 10 times higher in fossil than in modern enamel; secondary enrichments in fossil dentin and cement are even higher. In modern and fossil enamel, along sections perpendicular to the enamel-dentin junction (EDJ) or along cervix-apex profiles, P2O5 and CaO contents and the CaO/P2O5 ratios are very constant (StdDev ~1 %). Linear regression analysis reveals very tight control of the MgO (R2∼0.6), Na2O and Cl variation (for both R2>0.84) along EDJ-outer enamel rim profiles, despite large concentration variations (40 % to 300 %) across the enamel. These minor elements show well defined distribution patterns in enamel, similar in all specimens regardless of their age and origin, as the concentration of MgO and Na2O decrease from the enamel-dentin junction (EDJ) towards the outer rim, whereas Cl displays the opposite variation.
Fossil enamel from hippopotamids which lived in the saline Lake Kikorongo have a much higher MgO/Na2O ratio (∼1.11) than those from the Neogene fossils of Lake Albert (MgO/Na2O∼0.4), which was a large fresh water lake like those in the western Branch of the East African Rift System today. Similarly, the MgO/Na2O ratio in modern enamel from the White Nile River (∼0.36), which has a Precambrian catchment of dominantly granite and gneisses and passes through several saline zones, is higher than that from the Blue Nile River, whose catchment is the Neogene volcanic Ethiopian Highland (MgO/Na2O∼0.22). Thus, particularly MgO/Na2O might be a sensitive fingerprint for environments where river and lake water have suffered strong evaporation.
Enamel formation in mammals takes place at successive mineralization fronts within a confined chamber where ion and molecule transport is controlled by the surrounding enamel organ. During the secretion and maturation phases the epithelium generates different fluid composition, which in principle, should determine the final composition of enamel apatite. This is supported by co-linear relationships between MgO, Cl and Na2O which can be interpreted as binary mixing lines. However, if maturation starts after secretion is completed the observed element distribution can only be explained by recrystallization of existing and addition of new apatite during maturation. Perhaps the initial enamel crystallites precipitating during secretion and the newly formed bioapatite crystals during maturation equilibrate with a continuously evolving fluid. During crystallization of bioapatite the enamel fluid becomes continuously depleted in MgO and Na2O, but enriched in Cl which results in the formation of MgO, and Na2O-rich, but Cl-poor bioapatite near the EDJ and MgO- and Na2O-poor, but Cl-rich bioapatite at the outer enamel rim.
The linkage between lake and river water composition, bioavailability of elements for plants, animal nutrition and tooth formation is complex and multifaceted. The quality and limits of the MgO/Na2O and other proxies have to be established with systematic investigations relating chemical distribution patterns to sedimentary environment and to growth structures developing as secretion and maturation proceed during tooth formation.
Tubular carbonate concretions of up to 1 m in length and perpendicular to bedding, occur abundantly in the Upper Pliensbachian (upper Amaltheus margaritatus Zone, Gibbosus Subzone) in outcrops (Fontaneilles section) in the vicinity of Rivière-sûr-Tarn, southern France. Stable isotope analyses of these concretions show negative δ13C values that decrease from the rim to the center from −18.8‰ to −25.7‰ (V-PDB), but normal marine δ18O values (−1.8‰). Carbon isotope analyses of Late Pliensbachian bulk carbonate (matrix) samples from the Fontaneilles section show clearly decreasing C-isotope values across the A. margaritatus Zone, from +1‰ to −3‰ (V-PDB). Isotope analyses of coeval belemnite rostra do not document such a negative C-isotope trend with values remaining stable around +2‰ (V-PDB). Computer tomographic (CT) scanning of the tubular concretions show multiple canals that are lined or filled entirely with pyrite. Previously, the formation of these concretions with one, two, or more central tubes, has been ascribed to the activity of an enigmatic organism, possibly with annelid or arthropod affinities, known as Tisoa siphonalis. Our results suggest tisoan structures are abiogenic. Based on our geochemical analyses and sedimentological observations we suggest that these concretions formed as a combination of the anaerobic oxidation of methane (AOM) and sulfate reduction within the sediment. Fluids rich in methane and/or hydrocarbons likely altered local bulk rock carbon isotope records, but did not affect the global carbon cycle. Interestingly, Tisoa siphonalis has been described from many locations in the Grands Causses Basin in southern France, and from northern France and Luxemburg, always occurring at the same stratigraphic level. Upper Pliensbachian authigenic carbonates thus possibly cover an area of many thousand square kilometers. Greatly reduced sedimentation rates are needed to explain the stabilization of the sulfate-methane transition zone in the sedimentary column in order for the tubular concretions to form. Late Pliensbachian cooling, reducing run-off, and/or the influx of colder water and more vigorous circulation could be responsible for a halt in sedimentation. At the same time (thermogenic) methane may have destabilized during a major phase of Late Pliensbachian sea level fall. As such Tisoa siphonalis is more than a geological curiosity, and its further study could prove pivotal in understanding Early Jurassic paleoenvironmental change.
Die lithologische Abfolge des höheren Malm in Nordwestdeutschland wird mit Hilfe von biostratigraphisch aussagekräftigen Ammonitenfunden datiert und mit der westfranzösischen Gliederung parallelisiert. Die Grenze zwischen der Mutabilis- und der Eudoxus-Zone liegt innerhalb der "Stollenbank". Der "Knollenkalk" mit dem "Yo-Lager" des "Mittel-Kimmeridqe" entspricht basalen Abschnitten der Caletanum-Subzone der Eudoxus-Zone. Die "Ballersteinbank" enthält den "caletanum-Faunenhorizont", eine jüngere Kalkbank innerhalb der "Virgula-Bänke" den "quercynum-Faunenhorizont". Höchstes gesichertes marines Ober-Kimmeridgium (Autissiodorensis-Zone, Irius-Subzone) liegt in Gestalt der sogenannten "Gigas-Schichten" von Holzen/lth vor. An der Porta Westfalica kann hingegen eine Schichtlücke an der Basis der dortigen "Gigas-Schichten" , dem untertithonischen Gravesienkalk, weiter untermauert werden. Die jüngsten mit Ammoniten datierbaren Jura-Schichten lassen sich in den "Gigas-Schichten" am Südrand des Deister nachweisen.
In dieser Arbeit wird die Begleitflora zu den nicht-marinen Muschelfunden des Piesberges bei Osnabrück, worüber in Band 24 dieser Mitteilungen berichtet wurde, näher untersucht und dargestellt. Die nicht-marinen Muscheln wurden z.T. direkt aus dem Hangenden des Flözes Dreibänke, zum anderen und grössten Teil aus einer nahegelegenen Halde geborgen. Aufgrund des Vorkommens von Neuropteris dussartii Laveine, Sphenopteris coemansii Andrae und Alethopteris davreuxii (Brongniart) Goeppert kann festgestellt werden, dass das Haldenmaterial aus dem untersten Teil des Westfal D stammt.
Climate change research is increasingly focusing on the dynamics among species, ecosystems and climates. Better data about the historical behaviours of these dynamics are urgently needed. Such data are already available from ecology, archaeology, palaeontology and geology, but their integration into climate change research is hampered by differences in their temporal and geographical scales. One productive way to unite data across scales is the study of functional morphological traits, which can form a common denominator for studying interactions between species and climate across taxa, across ecosystems, across space and through time—an approach we call ‘ecometrics’. The sampling methods that have become established in palaeontology to standardize over different scales can be synthesized with tools from community ecology and climate change biology to improve our understanding of the dynamics among species, ecosystems, climates and earth systems over time. Developing these approaches into an integrative climate change biology will help enrich our understanding of the changes our modern world is undergoing.
The influence of dispersal limitation on species ranges remains controversial. Considering the dramatic impacts of the last glaciation in Europe, species might not have tracked climate changes through time and, as a consequence, their present-day ranges might be in disequilibrium with current climate. For 1016 European plant species, we assessed the relative importance of current climate and limited postglacial migration in determining species ranges using regression modelling and explanatory variables representing climate, and a novel species-specific hind-casting-based measure of accessibility to postglacial colonization. Climate was important for all species, while postglacial colonization also constrained the ranges of more than 50 per cent of the species. On average, climate explained five times more variation in species ranges than accessibility, but accessibility was the strongest determinant for one-sixth of the species. Accessibility was particularly important for species with limited long-distance dispersal ability, with southern glacial ranges, seed plants compared with ferns, and small-range species in southern Europe. In addition, accessibility explained one-third of the variation in species' disequilibrium with climate as measured by the realized/potential range size ratio computed with niche modelling. In conclusion, we show that although climate is the dominant broad-scale determinant of European plant species ranges, constrained dispersal plays an important supplementary role.
In diesem ersten Nachtrag zur Monographie der Insekten (ausschließlich der Odonatoptera und Blattodea) aus Schichten des Westfalium D vom Piesberg bei Osnabrück (Niedersachsen, Deutschland) werden weitere neue Angehörige der Palaeodictyoptera beschrieben: Homaloneura kiliani n. sp. (Spilapteridae) und Lithomantis meyeri n. sp. (Lithomanteidae). Neue Flügel-Funde zu bekannten Arten aus der Familie der Breyeriidae (Palaeodictyoptera) und Aspidothoracidae (Megasecoptera) ergänzen den bisherigen Fossilbericht von dieser Lokalität.
Neanderthal diets are reported to be based mainly on the consumption of large and medium sized herbivores, while the exploitation of other food types including plants has also been demonstrated. Though some studies conclude that early Homo sapiens were active hunters, the analyses of faunal assemblages, stone tool technologies and stable isotopic studies indicate that they exploited broader dietary resources than Neanderthals. Whereas previous studies assume taxon-specific dietary specializations, we suggest here that the diet of both Neanderthals and early Homo sapiens is determined by ecological conditions. We analyzed molar wear patterns using occlusal fingerprint analysis derived from optical 3D topometry. Molar macrowear accumulates during the lifespan of an individual and thus reflects diet over long periods. Neanderthal and early Homo sapiens maxillary molar macrowear indicates strong eco-geographic dietary variation independent of taxonomic affinities. Based on comparisons with modern hunter-gatherer populations with known diets, Neanderthals as well as early Homo sapiens show high dietary variability in Mediterranean evergreen habitats but a more restricted diet in upper latitude steppe/coniferous forest environments, suggesting a significant consumption of high protein meat resources.